Landouria schepmani ( Möllendorff, 1897 ) Nurinsiyah & Neiber & Hausdorf, 2019

Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard, 2019, Revision of the land snail genus Landouria Godwin-Austen, 1918 (Gastropoda, Camaenidae) from Java, European Journal of Taxonomy 526, pp. 1-73 : 54-56

Landouria schepmani ( Möllendorff, 1897) comb. nov.

Figs 54 , 76 ; Tables 3–4

Plectotropis schepmani Möllendorff, 1897: 68 (“ Java ”, more exactly ( van Benthem Jutting 1950; Zilch 1966): “Bungbulan, 1500 feet ”).

Landouria winteriana – van Benthem Jutting 1950: 463 (in part). — Zilch 1966: 294, pl. 7, fig. 7 (in part). (not Pfeiffer, 1842)

Diagnosis

Landouria schepmani is characterized by a depressed conical, sharply keeled shell with distinct incised spiral lines but no discernible scales, and a narrow umbilicus.

Material examined

Lectotype

INDONESIA • lectotype (designated by Zilch 1966); West Java, Sukapura, Gunung Bungbulang ; 7°27′ S, 107°36′ E?; ca 450 m a.s.l.; SMF 8877 .

Paralectotype

INDONESIA • 1 spec.; same data as for lectotype; SMF 8878 .

Description

SHELL ( Fig. 76 ; Tables 3–4 ). Depressed conical, with 5–5.25 slightly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles and distinct incised spiral lines; no scales discernible; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 21% of shell diameter, hardly obscured by columellar edge.

GENITALIA. Unknown.

Remarks

Van Benthem Jutting (1950) and Zilch (1966) synonymized L. schepmani with L. winteriana . However, L. schepmani differs from L. winteriana in having a sharply keeled, more conical shell with distinct incised spiral lines and a narrower umbilicus (U/D 0.21 vs 0.29–0.51 in L. winteriana ).

According to the original label, the type locality is Sukapura, Gunung Bungbulan. Van Benthem Jutting (1950: 465) listed a Bungbulan south of Gunung Papandayan (7°27′ S, 107°36′ E). It is not clear whether this is the Bungbulan of Fruhstorfer, because there are also several other localities called Bungbulan in the former Sukapura Regency. For example, there is a Gunung Bungbulang, now called Gunung Bangkuang, in West Java at 7°12′ S, 108°05′ E. Thus, the distribution area of L. schepmani remains dubious.

Distribution

Landouria schepmani is known only from “Gunung Bungbulan” in West Java (Fig. 54; see Remarks).

Figures

Figs 52–55 (opposite page). Distribution of species of Landouria in Java (10′ grid). 52. L. winteriana (Pfeiffer, 1842), L. naggsi sp. nov., L. nusakambangensis sp. nov., L. petrukensis sp. nov., L. tholiformis sp. nov., L. tonywhitteni sp. nov., L. smimensis (Mousson, 1848). 53. L. conoidea (Leschke, 1914), L. abdidalem sp. nov., L. sewuensis sp. nov., L. sukoliloensis sp. nov., L. pacitanensis sp. nov., L. intumescens (Martens, 1867), L. madurensis sp. nov. 54. L. rotatoria (Pfeiffer, 1842), L. nodifera sp. nov., L. parahyangensis sp. nov., L. schepmani (Möllendorff, 1897), L. zonifera sp. nov., L. ciliocincta (Möllendorff, 1897), L. pakidulan sp. nov. 55. L. epiplatia (Möllendorff, 1897), L. monticola van Benthem Jutting, 1950, L. davini Dharma, 2015, L. menorehensis sp. nov., L. dharmai sp. nov., L. moussoniana (Martens, 1867).

Figs 76–80. Shells of species of Landouria from Java. 76. L. schepmani (Möllendorff, 1897), West Java, Sukapura, Gunung Bungbulang (lectotype, SMF 8877). 77. L. monticola van Benthem Jutting, 1950, West Java, Gunung Cikuray (paratype, SMF 8880). 78. L. smimensis (Mousson, 1848), East Java, Gunung Semeru, ca 1200 m a.s.l., 8°06′ S, 112°55′ E (syntype of Helix smimensis, ZMZ 508598). 79. L. smimensis, East Java, Nongkojajar (holotype of Plectotropis tenggerica Schepman, 1912, ZMA 174453). 80. L. smimensis, East Java, Tosari (ZMA 390881). Scale bar: 5 mm.