Anotylus pseudopsinus ( Fauvel, 1895 )

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 232-240

Anotylus pseudopsinus ( Fauvel, 1895)

( Figs 94 , 202–208 )

Oxytelopsis pseudopsina Fauvel, 1895: 201 ; CAmeRoN 1930: 206; HeRmAN 1970: 413; HAm- moNd 1975: 169, 170; HAmmoNd 1984: 200; CoiffAit 1984: 151; HeRmAN 2001: 1414.

Oxytelopsisrufotestacea Cameron, 1925: 176, syn. nov.; HeRmAN 1970: 413; HeRmAN 2001: 1415.

Oxytelopsisbrevipennis Bernhauer, 1926: 313, syn. nov. (preoccupied); HeRmAN 1970: 413; HeRmAN 2001: 1413. (seeremark)

Oxytelopsischapmani Cameron, 1934: 32, syn. nov.; HeRmAN 1970: 413; HeRmAN 2001: 1413.

Oxytelopsisnigripennis Cameron, 1934: 32, syn. nov.; HeRmAN 1970: 413; HeRmAN 2001: 1414.

Oxytelopsis cassagnaui Coiffait, 1982: 163 , syn. nov.; CoiffAit 1984: 50; HeRmAN 2001: 1413.

Examinedtypematerial – Oxytelopsispseudopsina Fauvel – Lectotype (m, heredesignat- ed): “Lecto-; type [lilacmargineddisc, curatorlabel] \ [Mt.] Carin; AsciuiiGhecù [19°17’N, 96°42.5’E]; 1400–1500 m.; L. Fea. III-IV.[18]88. \ pseudopsina; Fvl. \ R.I.Sc.N.B. 17.479; Oxytelopsis ; Coll. et det. A. Fauvel \ Syntype \ Oxytelopsis ; pseudopsina Fvl. ; P.M. Hammond; det. 1972; Lectotype \ Anotylus ; pseudopsinus (Fauvel) ; det. Makranczy, 2012” (ISNB); Paralectotypes (2): “Para-; lecto-; type [lightbluemargineddisc] \ Carin; Asciuii Ghecù; 1400–1500 m.; L. Fea. III-IV.[18]88. \ Coll. et det. A. Fauvel; Oxytelopsis ; pseudopsina; Fauv.; R.I.Sc.N.B. 17.479 \ Oxytelopsis ; pseudopsinaFvl.; P.M. Hammond; det. 1972; Paralectotype \ Anotylus ; pseudopsinus (Fauvel) ; det. Makranczy, 2015” (2 f, ISNB); Oxytelopsisrufotestacea Cameron – Syntype (f): “Para-; lecto-; type [lightbluemargineddisc] \ Para-; type [yellowmargineddisc] \ PrinsenEil.[and = PulauPanaitan]; 21.I.1922; Damm. [erman] \ Oxytelopsis ; rufotestacea; Cam. \ M. Cameron.; Bequest.; B.M. 1955-147. \ Oxy- telopsis; rufotestaceaCam.; P.M. Hammond; det. 1970; Paralectotype ” (BMNH); Syntype (f, teneral): “PrinsenEil.[and]; 21.I.1922; Damm.[erman] \ Oxytelopsis ; rufotestacea; Cam; Type. \ Type [redcard]” (RMNH); Oxytelopsischapmani Cameron – Syntype (f): “Lecto-; type [lilacmargineddisc, curatorlabel] \ Type [redmargineddisc] \ Hoa- Binh.; Tonkin. 193 \ O.; chapmani; Type [underlined] Cam. \ M. Cameron.; Bequest.; B.M. 1955-147. \ Oxytelopsis ; chapmaniCam.; P.M. Hammond; det. 1970; Lectotype ” (BMNH); Oxytelopsis nigripennis Cameron – Syntype (f): “Lecto-; type [lilacmargineddisc, curatorlabel] \ Type [redmargineddisc] \ Hoa- Binh.; Tonkin. 193 \ O.; nigripennisType [underlined] Cam. \ M. Cameron.; Bequest.; B.M. 1955-147. \ Oxytelopsis ; nigripennisCam.; P.M. Hammond; det. 1970; Lectotype ” (BMNH), Syntype (f): “Hoa- Binh; Tonkin \ Cotype \ Oxytelopsis ; nigripennis; Cam. \ Oxytelopsis ; nigripennis; Cameron \ Coll. W. Chapman; inColl. P. Griveau; MHNG-2007 \ nigripennis; Cam.” (MHNG); Oxytelopsisbrevipennis Bernhauer – Holotype (m): “[leg.] L.J. Toxopeus; Buru, Station 9; 12. VI.[19]’21 \ Virginjungle;; decay- ingleaves \ Oxytelopsis ; brevipennis; Brnh. Typusunic[us] \ brevipennis; Bernh. Treubia; 1926, p. 313 \ ChicagoNHMus; M. Bernhauer; Collection \ Holo-; type [redmargineddisc, curatorlabel] \ Oxytelopsis ; brevipennisBnh; P.M. Hammond; det. 1974; Holotypem \ FMNH-INS; 0000 063 472 \ Photographed; D. Le 2011; EMu catalog” (FMNH).

Other material – VIETNAM: Tuyen Quang prov., NaHang Reserve , 300 m, 16- 20. V.1997, leg. S. Peck (97-8), rainforest, FIT (1, MHNG) ; Bien- ho, ПЛАНТАЦИЯ КИНКИНЬ, 17.X.1979, probe 0–10cm (1 m, 2, coll. Ryvkin); N-, NinhBinhPr., 90 kmSWHanoi, Cuc PhuongNP, primatesresc. centr., 190 m, 20°14’24”N, 105°42’53”E, 25.IV.2012, leg. A. Weigel & Li-Fa (2, NKME) ; NinhBinhProv., Res. Cuc- Phüöng , 200 m, 14.X.1963, leg. T. Pócs, dansforêttropic. pluv., àlalumière (1, HNHM) ; Hanoi, 20-30.IV.1991, leg. E. Jendek, at light (3, NHMW) ; Hanoi (1, ISNB) ; S-, 40 km NW An Khe, Buon Luoi , 14°10’N, 108°30’E, 620–750 m, 28.III-12.IV.1995, leg. P. Pacholátko & L. Dembický (4, NHMW) ; S-, Nam Cat TienNat. Park , 1-15. V.1994, leg. P. Pacholátko & L. Dembický (2, NHMW) ; Muongson , 8. XII.1971, leg. Gy. Topál (268), sifted litter (1, HNHM) ; Ha Son Binh , Hoa Binh, 30.I.1986, leg. S. Mahunka & J. Oláh (BerlesateNo. 23) (1, HNHM) , LamDongprov., DucMe ( Maria stream), 15 kmSBaoLoc, 700 m, 23.X.1988, leg. S. Mahunka & T. Vásárhelyi (No. 372), at light (7, HNHM) ; CHINA: Yunnan, Ninglang env., 27°19’N, 100°55’E, 6-10.VII.1992, leg. S. Bečvář (1, MHNG) ; INDIA: Kerala, CardamomHills, Mundakayam , 100 m, 9.XI.1972, leg. C. Besuchet, I. Löbl & R. Mussard (16), tamisagesdansplantationd’hévéas (2, MHNG) ; Kerala, CardamomHills, ValaraFall , à 46 kmSWMunnar, 450–500 m, 25.XI.1972, leg. C. Besuchet, I. Löbl & R. Mussard (49), tamisagesdanslaforêtprèsdelarivière (3, MHNG) ; Kerala, NelliampathiHills (NWAnaimalaiHills), Kaikatty , 900 m, 30.XI.1972, leg. C. Besuchet, I. Löbl & R. Mussard (58), tamisagesdenforêtprèsd’unruisseau (33, MHNG) ; MadhyaPradesh, MahadeoHills, 4.5 kmSWPachmarhi, VanshreeVihar 22°26’34”N, 78°23’25”E, 950 m, 16.X.2000, leg. G. Cuccodoro (1a), siftingflooddebrisatstream (32, MHNG) ; MadhyaPradesh, MahadeoHills, 4.5 kmSWPachmarhi, VanshreeVihar 22°26’34”N, 78°23’25”E, 950 m, 16.X.2000, leg. G. Cuccodoro (1b), siftingleaflitternearstream (15, MHNG) ; MadhyaPradesh, MahadeoHills, 5.5 kmSWPachmarhi, Tridhara 22°26’09”N, 78°23’01”E, 900 m, 18.X.2000, leg. G. Cuccodoro (3b), siftingmoistleaflitteronrocksingalleryforest (10, MHNG) ; Goa, Western Ghats , 1.5 km E Tambdi Suria Temple, 15°26’27”N, 74°15’49”E, 350 m, 31.X.2000, leg. G. Cuccodoro (3b), siftingvegetabledebrisinclearing (1, MHNG) ; Orissa [Odisha], Daitari , 24.XII.1966, leg. Gy. Topál (54), extractedfrommosses (1, HNHM) , same but (105), 2.I.1967 (1, HNHM) ; Orissa, Jajpur, Jajpur- KendujharDistr., Daitari , 23.XI.1967, leg. Gy. Topál (925), siftedlitter (2, HNHM) ; Orissa, Jajpur, Jajpur- KendujharDistr. , Dai- tari, 25.XI.-4.XII.1967, leg. Gy. Topál (941), siftedbarkoftree (2, HNHM) ; Orissa, Jajpur, Jajpur- KendujharDistr., Daitari , 4.XII.1967, leg. Gy. Topál (1012), fromunderbark (1, HNHM) ; UttarPradesh, Rishikesh, Laxman- Bridge , 3-4.VII.1989, leg. A. Riedel (1, SMNS) ; N-, Uttar Pradesh, 15-22 km N Rishikesh , 8.VIII.1989, leg. Hiermeier (47) (1, NHMW) ; Kumaon (UttarPradesh), BhimTal , ca. 1500 m, 4.X.1979, leg. I. Löbl (1), lisièred‘uneforêt secondaire, versantest, assezsec, tamisage (3, MHNG) ; Kumaon (UttarPradesh), BhimTal , 1550 m, 5.X.1979, leg. I. Löbl (2b) sous des pierres au bord d‘un ruisseau (1, MHNG) ; Kumaon (UttarPradesh), KathgodamprèsHaldwani , 600 m, 6.X.1979, leg. I. Löbl (3), ravin boisé, tamisagedefeuillesmortesprèsd‘unruisseau (97, MHNG) ; Kumaon (Uttar Pradesh), entreBhimTaletSatTal, ca. 1500 m, 7.X.1979, leg. I. Löbl (4), forêtsecondaire, versantnord, tamisagedefeuillesmortes (1, MHNG) ; Kumaon (UttarPradesh), Garijaà 10 kmdeRamnagar (CorbettNationalPark), 450 m, 15.X.1979, leg. I. Löbl (12b) tamisagede feuillesmortesdansunravinprofond (1, MHNG) ; Garhwal (UttarPradesh), routepour Mussoorie , 1300 m, 18.X.1979, leg. I. Löbl (14), versant sud, ravin boisé, tamisage de feuilles etbranchesmortes, humide (3, MHNG) ; Garhwal ( UttarPradesh ), 20 kmausuddeChamba, 1150 m, 20.X.1979, leg. I. Löbl (16), ravinavecunesource, gravieraveclesracinesde l‘herbeetaccumulationduboispourri (9, MHNG) ; Garhwal ( UttarPradesh ), 4 kmausud deBhatwari, 1400 m, 23.X.1979, leg. I. Löbl (22), lisièred‘uneforettrèsdégradée, tamisage demoussesetfougères (2, MHNG) ; Garhwal ( UttarPradesh ), entreTehrietSrinagar, à 20 kmdeSrinagar, ravin, 900 m, 25.X.1979, leg. I. Löbl (24 and 24c), tamisage, terreauaudessusdebuissons (14, MHNG) ; Garhwal ( UttarPradesh ), entreSrinagaretDevaprayag, à 16 kmdeSrinagar, 550 m, 29.X.1979, leg. I. Löbl (29), aubordd‘unerivière, tamisagede terreautrèshumide (2, MHNG) ; Garhwal ( UttarPradesh ), 22 kmNdeRishikesh, 450 m, 30.X.1979, leg. I. Löbl (31), versantsud, ravinboisé, tamisagedefeuillesetbranchesmortes, humide (5, MHNG) ; HimachalPradesh, ShivalikHills , 6kmSENahan, 800–900 m, 6.X.1996, leg. Schulz & Vock (1, coll. Assing) ; [Uttarakhand] Siwaliks, Nakraunda [30°14‘N, 78°08‘E], 8.X.1922, leg. M. Cameron (1, NMPC) ; [Uttarakhand] KolhuKhetGad. Mussooriedistrict , 1.XI.1921, leg. M. Cameron (1, MHNG) ; Uttarakhand, left side of Kosi River , 5 km N Ramnagar, 29.432°N, 79.140°E, 19-23. VI.2011, leg. A. Shavrin (1, coll. Shavrin) ; NE-, Assam / ArunachalPr. border, Bhalukpong , 27°00‘48“N, 92°39‘08“E, 150 m, 1-8. V.2012, leg. L. Dembický, FlightInterceptionTrap (1, ZFMK) , samebutsifting + Winklerapp. extraction (1, ZFMK) ; Assam, Umrongso env. 700 m, 25°27‘N, 92°43‘E, 3-8. VI.2002, leg. M. Trýzna & P. Benda (9, coll. Schülke) ; Assam, Gauhati , 200 m, 20.X.1978, leg. C. Besuchet, I. Löbl (22), tamisageaupiedd‘uneparoideterre, dansunecollineplusoumoinsboisée (3, MHNG) ; Assam, Manas ( Manaswildlifesanctuary ), 200 m, 21.X.1978, leg. C. Besuchet, I. Löbl (23), tamisagedanslaforêtprèsdesbungalows, solsablonneux (13, MHNG) ; Assam, Manas , 200 m, 22.X.1978, leg. C. Besuchet, I. Löbl (24a), sous des écorces (1, MHNG) , same but (24c), tamisageenforêt (4, MHNG); Meghalaya, GaroHills, Songsak , 400 m, 2.XI.1978, leg. C. Besuchet, I. Löbl (38c), tamisageenforêt, principalementsousdebambous (2, MHNG) ; Meghalaya, GaroHills, Rongrengiri , 400 m, 3.XI.1978, leg. C. Besuchet, I. Löbl (39b), tamisageenforêt, aupieddegrandsarbres (3, MHNG) ; Meghalaya, KhasiHills, Nongpoh , forêt au-dessusduvillage, 700 m, 5.XI.1978, leg. C. Besuchet, I. Löbl (42b), tamisagedanslaforêt, versantnord, aupiedderochersetdegrandsarbres (7, MHNG) ; Meghalaya, 1 kmEof Tura , 300–600 m, 25°30‘N, 90°14‘E, 2-5. V.2002, leg. M. Trýzna & P. Benda (13, coll. Schülke) ; WestBengal, Darjeelingdistr., Teesta , 3 kmenamontduvillage, 250 m, 10.X.1978, leg. C. Besuchet, I. Löbl (7 and 7b), tamisageenlisièredeforêt, aupieddegrandsarbresavecli- anes (13, MHNG) ; WestBengal, Darjeelingdistr., Sukna , 200 m, 7.X.1978, leg. C. Besuchet, I. Löbl (2b), tamisageenforêt (8, MHNG) ; WestBengal, Darjeelingdistr., entreTeestaet Rangpo , à 11 km de Teesta, 350 m, 12.X.1978 leg. C. Besuchet & I. Löbl (11b), sous des écorces (3, MHNG) ; WestBengal, Darjeelingdistr., Tonglu , versantnord, 2700 m, 16.X.1978 leg. C. Besuchet & I. Löbl (17), tamisagesdenforêt (2, MHNG) ; WestBengal, Darjeeling distr., Singla , 300 m, 18.X.1978, leg. C. Besuchet, I. Löbl (18b), tamisagedansuneforêtsec- ondairedetecks (1, MHNG) ; SRILANKA: Lindula , 10 kmWSWNuwaraEliya, leg. G. Benick, 29.III.1973, Quelle (1, coll. Schülke) ; Dickoya , 20 kmWSWNuwaraEliya, leg. G. Benick, 28.III.1973, QuelleimFels, gesiebt (1, coll. Schülke) ; Centralprov., Kandy , env. 600 m, forêt près du “Chalet Guesthouse”, 15.I.1970, leg. C. Besuchet, I. Löbl & R. Mussard (3c), tamisages (1, MHNG) ; Centralprov., Kandy, env. 600 m, UdawattekeleSanctuary , 19.I.1970, leg. C. Besuchet, I. Löbl & R. Mussard (11), tamisages en forêt vierge (3, MHNG) ; Centralprov. , au-dessusdeTalatuoya, entre 850 et 1000 m, 27.I.1970, leg. C. Besuchet, I. Löbl & R. Mussard (27a), tamisagesdansdesrestesdeforêt (4, MHNG) ; Centralprov., Hanguranketa , 750 m, 27.I.1970, leg. C. Besuchet, I. Löbl & R. Mussard (28), écorcesetPoly- poresd’unevieillesouche (3, MHNG) ; Centralprov., MahaweliGanga, à 7 mienavalde Kandy , env. 450 m, 30.I.1970, leg. C. Besuchet, I. Löbl & R. Mussard (34), au bord du fleuve, surlesablehumideaupieddesgraminées (1, MHNG) ; Centralprov., Ginigathena , env. 650 m, au bord du Mahaweli Ganga, 9.II.1970, leg. C. Besuchet, I. Löbl & R. Mussard (56), surlesablehumide (1, MHNG) ; Centralprov., Kandy , env. 700 m, collinesboiséesausud du lac, 14.II.1970, leg. C. Besuchet, I. Löbl & R. Mussard (67b), tamisage de feuilles mortes accumuléesdansungrandtrou (24, MHNG) ; THAILAND: S-, Yalaprov., Betongdistr. , Gunung Cang dun vill. [5°52‘N, 101°07‘E, 400 m], 26.III-22.IV.1993, leg. J. Horák & J. Strnad (91, NHMW) ; Phang- ngaprov., Takuapadistr., KhaoLak , 08°37.623‘N, 98°15.091‘E, 50 m, 23.VIII.-2.IX.2010, leg. A. Skale ( NHMW) ; Prov. Phitsanulok, ThungSa- LaengLuangN.P., 31. V.2001, leg. E. Horváth & Gy. Sziráki (21), atlight (1, HNHM) ; KhaoYaiNP [several smallstreams, veryshaded, withsandybottom, approx. 14°22‘26“N, 101°21‘42“E], 14. XI.1988, leg. M. Jäch (3) (1, NHMW) ; MaePing , 5-10.IX.1991, leg. H. Malicky, licht (2, NHMW) ; W-, SaiYokYaiNP, NWKanchanaburi, 3.XII.1999, leg. M. Jäch (8) (1, NHMW) ; Soppong , 19°29‘N, 98°18‘E, 750 m, 13. V.1993, leg. L. Bocák (3, SMNS) ; Soppong , 19°29‘N, 98°18‘E, 750 m, 13. V.1993, leg. V. Kubáň (4, SMNS) ; Chiang Mai, Doi Suthep , versant nord, 1050 m, 5.XI.1985, leg. D. Burckhardt & I. Löbl (10), ravin très humide, tamisage de bois pourri, desécorcesetchampignons (1, MHNG) ; ChiangMai, DoiInthanon , prèsdesmaisonsdu „ForestryDepartment“, 1250 m, 6.XI.1985, leg. D. Burckhardt & I. Löbl (13), tamisagededébrisvégétauxaubordd‘unruisseau (1, MHNG) ; Phetchaburi, KaengKrachan NationalPark , 200 m, auxenvironsde „Headquarters“, 16.XI.1985, leg. D. Burckhardt & I. Löbl (23), tamisagededébrisvégétauxàlalisièredelaforêt (57, MHNG) ; Phetchaburi, KaengKrachanNationalPark, 300–400 m, 25 à 30 kmde „ Headquarters “, 17.XI.1985, leg. D. Burckhardt & I. Löbl (24), tamisageaupieddegrosarbres (24, MHNG) ; Phetchaburi prov., KaengKrachanNationalPark, 450 m, à 30 kmde „ Headquarters “, 18.XI.1985, leg. D. Burckhardt & I. Löbl (25), tamisageaupieddegrosarbres (1, MHNG) ; Phetchaburi, Kaeng KrachanNationalPark, 450 m, à 35 kmde „Headquarters“, 19.XI.1985, leg. D. Burckhardt & I. Löbl (26), tamisagededébrisvégétauxdanslaforêt (36, MHNG) ; Chanthaburi, Khao SabapNationalPark, environsdePhliuWaterfalls, 150–300 m, 23-24.XI.1985, leg. D. Burck- hardt & I. Löbl (27a), tamisagededébrisvégétauxetdemoussesàproximitédecoursd’eau (3, MHNG) ; NEBangkok, KhaoYaiNationalPark, auxenvironsde “ Headquarters ”, 750– 850 m, 26.XI.-3.XII.1985, leg. D. Burckhardt & I. Löbl (28), sous les écorces (3, MHNG) ; NE Bangkok, KhaoYaiNationalPark, auxenvironsde “ Headquarters ”, 750–850 m, 26.XI.-3. XII.1985, leg. D. Burckhardt & I. Löbl (28b), tamisagededébrisvégétaux (26, MHNG) ; NE Bangkok, KhaoYaiNationalPark, CollinesEHeoSuwatWaterfalls, 800 m, 1.XII.1985, leg. D. Burckhardt & I. Löbl (30a), tamisagededébrisvégétauxàlalisièredelaforêt, prèsd’un merécageetprèsd’unruisselet (3, MHNG) ; ChiangMaiprov., ChiangDaodistr. , DoiChiangDaoWildlifeSanctuary, 510 m, 22.IX.-25.X.1990, leg. P. Schwendinger, pitfalltraps (2, MHNG) ; Chiang Mai prov., Doi Angkhang , 10 km W Fang, 1500 m, 30.X.1987, leg. P. Schwendinger, (2, MHNG) ; ChiangMaiprov., DoiSuthep, 1180 m, I-II.1986, leg. P. Schwendinger, pitfalltrap (1, MHNG) ; ChiangMaiprov., DoiInthanon , 1020 m, 17.II.1987, leg. P. Schwendinger (1, MHNG) ; ChiangRaiprov., 10 kmWWiangPaPao, BanHuayYa Sai , 780 m, 28.I.1988, leg. P. Schwendinger (6, MHNG) ; ChiangRaiprov., ThamPla , 500 m, 31.X.1991, leg. P. Schwendinger (3, MHNG) ; MaeHongSonprov., PongDüat , 750 m, 10. IV.1987, leg. P. Schwendinger (1, MHNG) ; Chanthaburiprov., NamtokPhliu , 50–100 m, 20.XI.1991, leg. P. Schwendinger (1, MHNG) ; Trangprov. , KhaoChongBotanicalGarden, 26.IX.2003, leg. A. Orosz & Gy. Sziráki (no. 30), atlightatstaffcenter (1, HNHM) ; Trang prov., KhaoChongBotanicalGardenatKhaoChongriver, 29.IX.2003, leg. A. Orosz & Gy. Sziráki (no. 39), atblacklight (1, HNHM) ; LAOS: Khammouanprov. , BanKhounNgeun, 300 m, 16. V.-6. VI.2007, leg. M. Štrba (5, SMNS) ; S-, Prov. Champasak, ca. 50 kmSPakse, Umg. BanPhatoumphone , burntclearings, 50–100 m, 23-24. V.1996, leg. H. Schillhammer (1a), at light (2, NHMW) ;; CAMBODIA: Silhanoukville , 5 m, 1.-13.I.2009, leg. S. & M. Mur- zin (1 m, coll. Schülke) ; Paifin , 200 m, 11-16. V.2009, leg. S. Murzin (1 m, coll. Schülke) ; MY- ANMAR: SagaingDivision , AlaungdawKatthapaNP, ca. 1.5 kmSWLogCabinCamp, alongPagodaRoad, 22°18.902’N 94°28.060’E, ca. 400 m, 6. V.2003, leg. H. Schillhammeret al. (108), semi-evergreenforestclosetoPagodaStream ; siftingofmoistleaflitterinsmall ravines (10, NHMW); SagaingDivision , AlaungdawKatthapaNP, aroundKuMaramon- astery, 22°18.560’N, 94°27.679’E, ca. 400 m, 8. V.2003, leg. H. Schillhammer et al. (117), siftingofmoistleaflitteramongstonesindried-upstream-bed, shadedbylargebushes (11, NHMW) ; SagaingDivision , AlaungdawKatthapaNP, KhaungDinStream, between 22°18.360’N 94°25.937’E and 22°19.650’N 94°25.768’E, ca. 450 m, 11.5.2003, leg. Schillhammeretal. (120), siftingofdryandmoistleaflitteralongstreamandinnearbyforest (1, NHMW) ; env. Maymyo , 700–900 m, 12-13.II.1996, leg. S. Kurbatov (2, MHNG) , samebut 4-5.III.1996 (1, MHNG) ; YangonDivision, Yangon, Insein , HighlandLodge ( PyayRoad , 7.5 miles), 11-12. V.1999, leg. H. Schillhammer & R. Schuh (38), at light (4, NHMW) ; Pegu , 60 kmNWYangoon, 17°19’N, 96°28’E, 22.XI.2003, leg. M. Hornburg, lighttrap (1, coll. Schülke) ; NEPAL: Seti (province), Bajhang (district), 12 kmNEChainpur, TalkotGadSofTalkot, 29°36’17”N, 81°17’54”E, 1400 m, 28. VI.2009, leg. A. Weigel (37), riverside, at light (1, NKME) ; Mahakali, Darchula, Latinathbachufer , 29°44’01”N, 80°46’00”E, 1030 m, 31.V-1. VI.2005, leg. A. Weigel (3, NKME) ; prov. Bagmati, DobateRidgeNEBarahbise, 2800 m, 2. V.1981, leg. I. Löbl & A. Smetana (55d), tamisage de bois pourri, de feuilles mortes et de moussesdansunechênaieavecrhododendrons (1, MHNG) ; prov. Bagmati, 4 kmSTarang Marang , 900 m, 28.IV.1981, leg. I. Löbl & A. Smetana (50), tamisage d’herbe et de feuilles mortesprèsd’unesource (2, MHNG) ; prov. Bagmati, GokanaForestnr. Kathmandu , 1400 m, 31.III.1981, leg. I. Löbl & A. Smetana (1b), tamisage dans la forêt (2, MHNG) ; prov. Bag- mati, Gokana Forest nr. Kathmandu , 1400 m, 1.IV.1981, leg. I. Löbl & A. Smetana (2b), tamisage, dansunravinboisé, demoussesetdebranchesmortes (2, MHNG) ; prov. Bag- mati, NagarjunForestWKathmandu, 1650 m, 2.IV.1981, leg. I. Löbl & A. Smetana (3), forêt secondairejeune, surleversantouest, tamisagedemoussesetfeuillesmortes (1, MHNG) ; distr. Kathmandu , GokarnaForestReserve, 1350 m, 20.X.1983, leg. I. Löbl & A. Smetana (40), tamisagedefeuillesmortesdanslaforêtassezsèche (1, MHNG) ; distr. Kathmandu, Godawari , 1600 m, forêtau-dessusdujardinbotanique, 31.III.1984, leg. I. Löbl (1), tamisage de feuilles mortes près d‘un ruisseau, humide (6, MHNG) ; MALAYSIA: W Ipoh, 5 km of TanjongRambutan , 13-15.IV.2000, leg. M. Snižek (1, coll. Schülke) ; MALAYSIA /BORNEO: Sabah, SepilokForestRes. (nearSandakan), 8. VI.1968, leg. R.W. Taylor, exhumus (1, MHNG) ; Sabah, KinabaluPark , PoringHotSprings, ca. 500 m, forêtàDipterocarpaceae relativementsèche, 6. V.1987, leg. D. Burckhardt & I. Löbl (14a), tamisagedeboispourri avecchampignonsalalisièredelaforêt, etdefeuillesmortesaupieddesarbreslaforêt (4, MHNG) ; Sabah, Mt. KinabaluNationalPark, PoringHotSprings 480 m, 10. V.1987, leg. A. Smetana (B39), siftingofleaflitter, debrisandfermentingfruitsonforestfloor (2, FMNH) ; Sabah, Mt. KinabaluNationalPark, PoringHotSprings, 485 m, 25.VIII.1988, leg. A. Smetana (B147), siftingofaccumulatedoldfruitsof Areca palms, includingdebrisandhumus underneath, openarea (1, FMNH) , samebut 29.VIII.1988 (B160) (2, FMNH) ; Sabah, Bing- korNKeningau, 400–500 m, 19-20.XI.1996, leg. W. Schawaller (1, SMNS) ; Sabah, Poring HotSprings , 450–600 m, 9-11.III.2007, leg. W. Schawaller (3, SMNS) ; Sarawak, Gn. Gading N. P., 150–300 m, 9-12.III.2008, leg. W. Schawaller (1, SMNS) ; PHILIPPINES /MINDANAO: Davao prov., 25 km NW New Bataan, 1200 m, 20-22. V.1996, leg. Bolm (1, SMNS) ; PHILIP- PINES / LEYTE: ViscaNBaybay, 200–500 m, 22.II.1991, leg. W. Schawalleretal., primary forest (3, SMNS) , same but 22.II.-10.III.1991 (4, SMNS) ; Visca N Baybay , 3.III.1991, leg. W. Schawalleretal., cultivatedland (1, SMNS) ; PHILIPPINES / LUZON: Lagunas (Calabar- zon) ., Mt. Makiling , summit road (S of Los Banos), 500 m, 20.XI.1995, leg. I. Löbl (2), degradedrainforest, bambooleaflitterandotherdebrisnearroad (14, MHNG) ; Lagunas (Calabarzon)., Mt. Makiling , aboveMadSprings, 400–700 m, 19-22.XI.1995, leg. J. Kodada, degradedrainforest (5, MHNG) ; Lagunas (Calabarzon), Mt. Makiling , summitroad ( Sof LosBanos ), 600 m, 21-22.XI.1995, leg. I. Löbl (3), degradedrainforest, moss, epiphytesand rottenwoodonlargefallentrees (1, MHNG) ; Lagunas (Calabarzon), Mt. Banahawabove Kinabuhayan , 600–700 m, trailtoCrystalino, 24.XI.1995, leg. I. Löbl (5a), degradedrainfor- est, forestfloorlitter (1, MHNG) , samebutleg. J. Kodada & B. Rigová (1, MHNG) ; Lagunas (Calabarzon), Mt. Banahawnearschoolabout 1 kmKinabuhayan, 500 m, 26.XI.1995, leg. I. Löbl (7a), degradedrainforest, forestfloorlitter (5, MHNG) , samebutleg. J. Kodada (4, MHNG) ; CentralLuzon, Mt. BanahawaboveKinabuhayan , summittrail, ca. 800 m, 25. XI.1995, leg. I. Löbl (6), degraded rainforest, on slope (3, MHNG) ; PHILIPPINES / PALA- WAN: Central , aboveSanRafael, ca. 300 m [10°04’34”N, 118°54’14”E], 4.XII.1995, leg. J, Kodada, degradedrainforest, onslope (4, MHNG) , samebutleg. I. Löbl (13), leaflitter (9, MHNG) ; Central , alongTarabananriverNESanRafael, ca. 30 m, 7.XII.1995, leg. J. Kodada & B. Rigová (1, MHNG) ; Central, Concepcion, largelogsacrossConcepcionriverNESan Rafael , ca. 20 m [10°01’17”N, 119°04’35”E], 8.XII.1995, leg. J. Kodada & B. Rigová (3, MHNG) ; INDONESIA / SUMATRA: Jambi, Mt. Kerinci, 1750 –1850 m, 11-12.XI.1989, leg. I. Löbl, D. Agosti, D. Burckhardt (11), siftingofvegetationaldebrisinmontaneLithocarpus- Castanopsisforest (1, MHNG) ; INDONESIA / SULAWESI: Maros , KaraentaNatureReserve, 500 m, 18.IV.1991, leg. D. Agosti (22) ( F91791 ), limestone, rainforest, clay, thinleaf litterlayerandhandcollecting (3, MHNG) ; Pattanetegang , 1600 m (?, ca. 5°25’S, 119°59’E), 9.IV.1991, leg. D. Agosti (15), secondaryforest, leaflitterlayerthin, soilclay, fewmycor- rhiza (11, MHNG) ; Ulumambi (W of Mamasa), 1000 m, 11.IV.1991, leg. D. Agosti ( F91741 ), remnantsofrainforest / coffeeplantation, leaflitter (3, MHNG) ; Larompang- Temboe (Sof Palopo), 50 m, Naturereserve , secondarylowlandrainforest, 20.IV.1991, leg. D. Agosti (24), leaflitterlayer, thickandheavilyinfestedwithmycorrhiza, leaflittersampleandhand collecting (1, MHNG) ; SE-, nearKolaka, 300 m, 20. V.1997, leg. S. Kurbatov (1, MHNG) ; INDONESIA / BALI: Penelokan , 25.XI.1978, leg. J. T. Huber (2, MHNG) ; INDONESIA / JAVA: WestJava, Bogor, Parung , Churuk , 17. VI.-15.VII.2005, leg. S. K. Alekseyev (5, coll. Ryvkin) ; INDONESIA /SUMBAWA: NE-, Calabai (TambonaN. P.), 11-13.II.1994, leg. Bolm (16, SMNS, 2, HNHM) ; INDONESIA /LOMBOK: Senaro, N slope of Rinjani , 1100 m, 2-5. II.1994, leg. Bolm (2, SMNS) ; Suranadi , park with secondary forest, 190 m, 9.III.1991, leg. D. Agosti (1) ( F91507 ), soilwithclay, distinctleaflitterlayer (4, MHNG) ; INDONESIA / TI- MOR: East Nusa Tengarra prov., between Soe and Kapan , 16 km N Soe, 1000 m, 30.III.1991, leg. D. Agosti (13) ( F91654 ), evergreenforestonlimestone, talltreeswithbutressesand epiphytes, onslopeverythinleaflitterlayer, afterrain, winkler (18, MHNG) , samebut 29.III.1991 ( F91643 ) (3, MHNG) ; INDONESIA /MOLUCCAS: Tanimbar, Yamdena isl., Bomaki (NWofSaumlaki), 50 m, 18.IX.1991, leg. D. Agosti (F911146), limestone, Kanary nutforest, evergreen, leaflitter (8, MHNG) ; INDONESIA /IRIANJAYA: Sorongprov., Sala- watti Isl., Solol , 200–300 m, 6-7.XI.1996, leg. A. Riedel, sifted (6, NHMW) ; Sorong, Sala- watti Isl., Kalobo , 10–30 m, 19-22.X.1996, leg. A. Riedel, sifted (1, NHMW) ; Sorong prov., Batanta Isl., Waylebet , 500 m, 29.X.1996, leg. A. Riedel, sifted (2, NHMW) , same but 800 m (1, NHMW) ; Sorong prov., Batanta Isl., Waylebet , 10 m, 30.X.1996, leg. A. Riedel, sifted (8, NHMW) ; Sorongprov., BatantaIsl., Waylebet , 150 m, 2.XI.1996, leg. A. Riedel, sifted (17, NHMW) ; Sorongprov., WaigeoIsl., KabuiBay , Wawiay , 50 m, 14.XI.1996, leg. A. Riedel (4, NHMW) ; Sorong, Intimpuraloggingcamp , dusunMeibo, 100–150 m [00°50’45”S, 131°23’03”E], 19.I.2001, leg. A. Riedel, siftedinforest (14, SMNS) ; WandammenBay, Won- diwoi Mts., Wasior , 500–700 m [02°43’14”S, 134°32’41”E], 14.I.2001, leg. A. Riedel, sifted in forest (1, SMNS) ; Japen Isl., E Serui, Wawuti , 300 m [01°50’47”S, 136°24’50”E], 17.XII.2000, leg. A. Riedel, siftedinforest (7, SMNS) ; JapenIsl., onroadtoAmbaidiru, Kontiunai , 600– 700 m [01°47.5’S, 136°18.0’E], 26.XII.2000, leg. A. Riedel, sifted in forest (4, SMNS) ; Jayapuraprov., RouffaerMts., Mamberamo , Noiadi , 150–200 m, 17.III.1999, leg. A. Riedel, sifted (3, NHMW) ; PAPUA NEW GUINEA: nr. Lae , ca. 50 m, 11. VI.1972, leg. R.W. Taylor ( ANIC Berlesate 391), rainforest (1, ANIC) ; Passam , nearWewak, ca. 270 m, 5.VII.1972, leg. R.W. Taylor (1, ANIC) ; near Kokoda , ca. 500 m, 1. VI.1972, leg. R.W. Taylor (4, ANIC) ; 8 km S of Kokoda , 800 m, 1. VI.1972, leg. R.W. Taylor (3, ANIC) ; Sandaunprov. , NMianmin, 1100 m [4°53‘9.68“S 141°36‘50.42“E], 20. V.1998, leg. A. Riedel, sifted (9, NHMW, 2 m, HNHM) ; West Sepik prov., Vanimo, Denake-range , km 12, 500 m [2°44’58”S, 141°14’54”E], 28- 29.X.1992, leg. A. Riedel (25B), roadsideforest, sifting (6, SMNS) ; Madangprov., centreof Baiteta forest (05°01’S, 145°45’E), 1. V.1996, leg. O. Missa (AR41), at light (7, ISNB) , same but 21. V.1996 (AR10) (2, ISNB) , same but 24. VI.1996 ( T2 ) (2, ISNB) , same but 20. VI.1996 ( T2 ) (3, ISNB) , samebut 15. V.1996 ( T11 ) (2, ISNB) ; [Madangprov.,] env. Madang, Nagada [Mission] [5°09.5’S, 145°47.3’E], VI.1979, leg. W.G. Ullrich (3, MHNG) ; Morobeprov., Mindik- Pindiu, near Kuat river , 500–580 m, 28.IV.1998, leg. A. Riedel, sifted (5, NHMW) ; [Morobe prov.,] Finschhafen, Heldsbach [6°29.5’S, 147°50.0’E], XI.1979, leg. W.G. Ullrich (3, MHNG) ; Morobeprov., Wauregion, WauEcologyInstitute ( WEI) area, 1400 m, 16. V.1992, leg. G. Cuccodoro (1a), lowermontanerainforest, vegetationaldebris (34, MHNG) ; Morobeprov., Wauregion , WEIarea, 1150 m, 17. V.1992, leg. G. Cuccodoro (2b), verydegradedlower montanerainforest, vegetationaldebrisatfootofisolatedKlinkiipines- 80% itsleaves (10, MHNG) , samebut (2c), vegetationaldebrisonrocksinsmallgalleryforest (22, MHNG); Morobeprov., Wauregion , WEIarea, 1450 m, 21. V.1992, leg. G. Cuccodoro (5b), lower montanerainforest, rottingtrunkandvegetationaldebris (20, MHNG) ; Morobeprov., Wauregion, Mt. Kaindiroad , 1350 m, 24. V.1992, leg. G. Cuccodoro (7), gardenbordering lowermontanerainforest, vegetationaldebrisandrottingtrunks (106, MHNG) ; Morobe prov., Wauregion, Mt. Kaindiroad , 1700 m, 25. V.1992, leg. G. Cuccodoro (8a), degraded lowermontanerainforest, vegetationaldebrisandrottingtrunks (1, MHNG) ; Morobe prov., Wauregion , WEIarea, 1200 m, 26. V.1992, leg. G. Cuccodoro (9a), coffeeplantation, rottingwood (2, MHNG) , samebut (9b), vegetationaldebris (1, MHNG), samebut (9c), barkandbasidiomycetes (2, MHNG); Morobeprov., Wauregion , WEIarea, 1150 m, 27. V.1992, leg. G. Cuccodoro (10b), verydegradedlowermontanerainforest, rottingtrunk (4, MHNG) , samebut (10c), basidiomycetes (1, MHNG); Morobeprov., Wauregion, Bieru road ( Kaisenikschool ), 1050 m, 28. V.1992, leg. G. Cuccodoro (11c), galleryforest, mossand vegetationaldebris (2, MHNG) , samebut (11d), vegetationaldebris (7, MHNG).

Redescription – Measurements (in mm, n=10): HW = 0.55 (0.52–0.58); TW = 0.51 (0.47– 0.54); PW = 0.72 (0.67–0.78); EW = 0.82 (0.73–0.92); AW = 0.81 (0.74–0.92); HL = 0.43 (0.39– 0.46); EL = 0.15 (0.14–0.16); TL = 0.13 (0.12–0.14); PL = 0.44 (0.42–0.47); SC = 0.53 (0.47–0.60); FB = 1.44 (1.34–1.55); BL = 2.62 (2.40–2.88). Habitus as in Fig. 94 . Forebody moderately shin- ingdespitesculpture, abdomenveryfinelymicrosculpturedandwithlargebutextremlyshallow, obscuredpunctures, lesslustrous. Elytrareddishdarkbrowntosometimes blackish, head, pronotum, abdomenandlegsmediumtodarkbrown, exceptexplanate partofpronotumandtransparentclypealarealighter, mouthpartsandantennomeres 1–4 mm for Figs 202, 204–205 , 0.125 mm for Fig. 203

mediumbrown, otherantennomeresdarkbrown. Headwithanextremelythinrimon anteriormargin, latterrathertruncateinmiddle, slightlyangledalsoatsupraantennal prominences, theseasmoderatelyelevatedobliqueridges, discslightlyimpressednear eyes. Epistomalsutureformingsubcircularimpression (lesspuncturedinside), uppersur- facewithouttactilesetae, eyesmoderatelylargeandsomewhatbulgingfromsidelineof head; indorsalviewtemples (lessthanlengthofeyeorslightlymoreincaseofbrachypter morph) alatiform, laterallyroundedandposteriorlyconstrictedtoformdistinctneck, however, dorsallynotseparatedbyoccipitalfurrow, onlybydorsumofneckbeingmicrosculpturedratherthanpunctured. Antennomere 1 club-shapedandverylightlymicrosculptured, segment 2 basallybroadoval, segment 3 stoutclub-shaped, segment 4 slightly transversediscoid, segments 5–11 withbasaldishes, segments 6–10 transverse (about 1.5× broaderthanlong). Headandpronotumwithmostlyalveolatesculpture. Pronotumwith lateralportionexplanateandslightlyup-turned, edgealatiformwithsparseshortsetae; lateralmarginunevenlyarchedwithanumberofsmall, irregularlydistributeddenticles, posteriormarginbroadlyandrathershallowlyincisedbeforecorners; discmediallywith twonearlyparallellongitudinalridges (posteriorlyconfluentoralmost), impressedalong- sidethem, laterallyimpressedaroundmiddleofsides. Elytrawithverythinmembranous posteriormarginextendingfromsuturalcornerto 4/5 ofhindmargin. Elytralepipleural ridgepresentwithslightlyup-turnedlateralalatiformportions, disclongitudinallyweakly impressedalongside; shoulderswelldeveloped, surfacefoveolate, inbetweenpunctures torulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlon- gitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesratherarcuate, second segmentwithparatergitesstronglybroadeningposteriorly (abdomenappearingconstrict- edatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad; posterioredgeof tergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasinFig. 202 , sterniteVIII ( Fig. 203 ), tergitesIX ( Fig. 204 ), X ( Fig. 205 ) andaedeagusasinFigs 206–208 .

Distribution – The species is widely distributed from the whole range of theHimalayatosouthernChinaandVietnamtothePhilippines, theGreater SundaIslandsandNewGuinea (seealsobelow).

Remarks – The holotype of Oxytelopsis cassagnaui Coiffait, 1982 is lost (similarly to that of O. franzi described in the same article). While most of the synonymsofthisnameseemtorepresentformswellwithinthe (great) range ofvariationknownforthisspecies, someremarksmustbemadeasregards ‘ brevipennis Bernhauer’. Apartfromdifferentbodyproportions, examination ofmalegenitaliaandterminaliaonlyrevealedafewsubtledifferencesfrom themainlandSEAsianforms; thesedeviationsaretoosmalltoallowforsafe recognitionofadistincttaxon, soitisherekeptwithin A. pseudopsinus . This formwithsmallerelytraandcomparativelysmallereyesandlongertemples existsmostlyeastofWallace’sLine. Thereisstillagreatrangeofvariation existingwithinthosesoutheasternpopulations, withratherdifferentexternal appearanceswithgenitaliaquiteuniform. Toconfusemattersevenmore, a similar population with smaller elytra exists on the island of Ceylon (Sri Lan- ka). Thedescriptionandgenitaliaillustrationsprovidedherearebasedonthe mainlandSEAsianform (cf. thetypeof‘ pseudopsinus ’), butthehabituspho- tographisofaspecimenwithsmallerelytra (cf. thetypeof‘ brevipennis ’). The name‘ brevipennis ’ isayoungersecondaryhomonymandmustbereplacedif usedasvalid.

Figures

Figs 94–95. Habitus of Anotylus species. 94 = A. pseudopsinus (Fauvel, 1895), 95 = A. malaisei (Scheerpeltz, 1965)

Figs 202–208. Anotylus pseudopsinus (Fauvel, 1895), male. 202 = median part of sternite VII, 203 = sternite VIII, 204 = tergite IX, 205 = tergite X, 206 = aedeagus, frontal view, 207 = median lobe, lateral view, 208 = paramere, lateral view. Scales: 0.10 mm for Figs 206–208, 0.115