Anotylus tanator, Makranczy, 2017

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 255-257

Anotylus tanator sp. n.

( Figs 97 , 251–257 , 277 )

Typematerial – Holotype (m): INDONESIA / SULAWESI: S-, TanahToraja, [Baruppu,] Pulu Pulu [02°48.770’S, 119°47.616’E], 1700 m, 13-16.VIII.1990, leg. A. Riedel, sifted in for- est ( SMNS) ; Paratypes (25): samedataasholotype (2 m, 1 f, SMNS, 1 m, HNHM); Tanah Toraja, [Baruppu,] Ponding - Pulu Pulu [02°49.550’S, 119°47.335’E], 1600–1800 m, 13-17. VIII.1990, leg. A. Riedel, sifted in forest (1 m, 2 f, SMNS, 1 m, MHNG, 1 m, NHMW); W-, Mamasa, Penanang, WofMamasa, 1600 m [02°54‘38“S, 119°18‘30“E], 9.IV.1991, leg. D. Agosti ( F91724 ) (16), montanerainforest, leaflitter (1, FMNH) , samebut ( F91722 ) (2, MHNG) ; W-, Mamasa, Ulumambi, WofMamasa, 1000 m [02°55‘57“S, 119°15‘05“E], 11.IV.1991, leg. D. Agosti ( F91741 ) (17), remnantsofrainforest / coffeeplantation, leaflitter (3, MHNG, 1, AMNH); W-, Mamasa, Ulumambi, WofMamasa, 1300 m [02°55‘09“S, 119°16‘22“E], 12.IV.1991, leg. D. Agosti (18), montane forest, leaf litter (6, MHNG, 1, ISNB, 1, ZMUC, 1, MNHP) .

Description – Measurements (in mm, n=10): HW = 0.70 (0.64–0.74); TW = 0.69 (0.62– 0.72); PW = 1.02 (0.93–1.08); EW = 1.10 (1.05–1.16); AW = 1.18 (1.08–1.26); HL = 0.55 (0.50– 0.57); EL = 0.14 (0.13–0.15); TL = 0.21 (0.19–0.23); PL = 0.60 (0.54–0.65); SC = 0.65 (0.61–0.69); FB = 1.88 (1.75–1.95); BL = 3.62 (3.35–3.90). Habitus as in Fig. 97 . Forebody weakly shining becauseofsculpture; abdomenwithsuchextremelydensecoriaceousmicrosculpturethat noteventracesofpunctationobservable, opaque. Forebody, legsandabdomenreddish mediumtodarkbrown, exceptexplanatepartofpronotumandclypealarea (bothfor transparency), these, mouthpartsandantennaereddishmediumbrown. Headwithathin rimonanteriormargin, truncateinmiddle, laterallyevenlyarchedtowardseyes, supraan- tennalprominencesasmoderatelyelevatedobliqueridges, discslightlyimpressednear eyes. Epistomalsutureformingsubcircularimpression (almostimpunctateinside), upper surfacewithouttactilesetae, eyesmoderatelysmall, veryslightlybulgingfromsidelineof head; indorsalviewtemples (1.5× lengthofeye) alatiform, laterallyimperfectlyrounded (withoutbeingangled) andposteriorlyconstrictedtoformdistinctneck, notseparated dorsallybyoccipitalfurrow, onlybyneckbeingmicrosculpturedratherthanpunctured. Antennomere 1 club-shaped, finelysculptured, segment 2 elongateoval, segment 3 clubshaped, segment 4 smallandisodiametrical, segments 5–11 withbasaldishes, articles 6–10 transverse (about 1.5× broaderthanlong), segments 9–10 bulkierthanpreceding. Head andpronotumwithmostlylacunosesculpture. Pronotumwithlateralportionexplanate andslightlyup-turned, edgealatiformwithsparseshortsetae; lateralmarginmoreorless evenlyarchedwithanumberofsmall, irregularlydistributeddenticles, posteriormargin broadlyincisedbeforecorners; discmediallywithtwoparallellongitudinalridges, almost confluentposteriorly, impressedalongsidethem, laterallyimpressedaroundmiddleof sides. Elytrawithmembranousposteriormarginextendingfromsuturalcornerto 3/4 of hindmargin. Elytralepipleuralridgepresentwithup-turnedlateralalatiformportions, disclongitudinallyimpressedalongside; shoulderswelldeveloped, surfacefoveolate, in betweenpuncturestorulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesrath- erarcuate, secondsegmentwithparatergitesstronglybroadeningposteriorly (abdomen appearingconstrictedatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad, posterioredgeoftergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasin Fig. 251 , sterniteVIII ( Fig. 252 ), tergitesIX ( Fig. 253 ), X ( Fig. 254 ) andaedeagusasinFigs 255–257 , spermathecaasinFig. 277 .

0.11 mm for Figs 253–254 , 0.13 mm for Fig. 252

Etymology – The specific epithet is composed of the beginnings of the two words of thetypelocalityarea, TanahToraja (nouninapposition).

Distribution – Accordingtoadministrativeareasthespeciesisknown fromanareapartlybelongingtoWSulawesi (Mamasaregency), partlyS Sulawesi (TanaTorajaregency).

Remarks – Itisapparentlyrelatedto A. rurukan (alsofromSulawesi, but fromthenorthernpart).

Figures

Figs 96–97. Habitus of Anotylus species. 96 = A. rurukan sp. n., 97 = A. tanator sp. n.

Figs 251–257. Anotylus tanator sp. n., male. 251 = median part of sternite VII, 252 = sternite VIII, 253 = tergite IX, 254 = tergite X, 255 = aedeagus, frontal view, 256 = median lobe, lateral view, 257 = paramere, lateral view. Scales: 0.09 mm for Fig. 251, 0.10 mm for Figs 255–257,

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali