Anotylus franzi ( Coiffait, 1982 )

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 187-193

Anotylus franzi ( Coiffait, 1982)

( Figs 86 , 104–110 )

Oxytelopsis franzi Coiffait, 1982: 162 ; CoiffAit 1984: 150; HeRmAN 2001: 1413.

Examined type material – Oxytelopsis franzi Coiffait – Neotype (m, here designated): “ NEPAL: (prov. Bagmati); below Thare Pati ; 3300 m, 10. V.[19]81; [leg. I.] Löbl & [A.] Smetana; [on the backside:] 18b [tamisage de mousses, de fougères et de feuilles mortes d’érables sur une pente raide près d’un ruisseau] \ Neotypus; Oxytelopsis ; franzi Coiffait ; des. Makranczy, 2015 \ Anotylus ; franzi (Coiffait) ; det. Makranczy, 2015” ( NHMW).

Other material – NEPAL: prov. Bagmati, Burlang Bhanjyang, 2600 m, 5.IV.1981, leg. I. Löbl & A. Smetana (7b), tamisage de mousses et de feuilles mortes près d’une source (2, MHNG, 1, ISNB); prov. Bagmati, au-dessus de Gul Bhanjyang, 2600 m, 6.IV.1981, leg. I. Löbl & A. Smetana (9), vieille chênaie sur le versant nord, pente raide, tamisage de feuilles mortes, mousses et bois pourri, au pied des arbres (13, MHNG) ; prov. Bagmati, Malemchi , 2900 m, 14. V.1981, leg. I. Löbl & A. Smetana (23), tamisage de branches pourries, feuilles mortes, herbe et mousses au pied d‘une muraille près du village (1, MHNG) ; prov. Bagma- ti, Malemchi, 2800 m, 14. V.1981, leg. I. Löbl & A. Smetana (24), tamisage de branches pour- ries, feuilles mortes, herbe et mousses au pied d‘une muraille près du village (9, MHNG, 1, CNCI, 1, NIBR, 1, ZMHB, 1, HNHM); prov. Bagmati, Malemchi , 2900 m, 18. V.1981, leg. I. Löbl & A. Smetana (30), mousses sur les tsuga, tamisage; tamisage de bois pourri dans un chêne creux (9, MHNG) ; prov. Bagmati, Tarke Ghyang [28°00.0‘N, 85°33.5‘E], 2650 m, 19.IV.1981, leg. I. Löbl & A. Smetana (32a), tamisage de feuilles mortes et de mousses dans un ravin près d‘un ruisseau (4, MHNG, 1, NKME, 1, SDEI); prov. Bagmati, above Sher- mathang, 2900 m, 26.IV.1981, leg. I. Löbl & A. Smetana (47c), tamisage de feuilles mortes et fougères entre les roches dans une forêt de chênes-rhododendrons (4, MHNG, 1, BMNH, 1, MNHP, 1, ZMUC, 1, FMNH, 1, NMPC, 1, SMNS, 1, AMNH, 1 m, 1, HNHM) .

Redescription – Measurements (in mm, n=10): HW = 0.60 (0.58–0.61); TW = 0.58 (0.56–0.60); PW = 0.79 (0.75–0.84); EW = 0.81 (0.78–0.84); AW = 0.86 (0.82–0.91); HL = 0.63 (0.46–0.49); EL = 0.105 (0.10–0.11); TL = 0.19 (0.18–0.20); PL = 0.52 (0.50–0.54); SC = 0.38 (0.36–0.39); FB = 1.42 (1.38–1.45); BL = 2.90 (2.75–3.06). Habitus as in Fig. 86 . Forebody moderatelyshiningdespitesculpture, abdomenwithcoriaceousmicrosculpture, moderately finelyandshallowlypunctured, lesslustrous. Bodyreddishmediumtodarkbrownexcept anteriormarginofhead, lateralmarginofpronotum, antennomeres 1–4 andmouthparts mediumbrown, apexoflastantennomereoccasionallyalsolighter. Headwithanterior margintruncateinmiddle, obliquetosupraantennaltubercles (asmoderatelyelevated obliqueridges) wheregentlyangledagain, possessingslightrim (obsoleteatfront), disc slightlyimpressedneareyes. Epistomalsutureformingsubcircularimpression (almostun- puncturedinside), uppersurfacewithouttactilesetae, eyessmallandextremelybulging (likehalfsphere); indorsalviewtemples (1.8× lengthofeye) alatiform, laterallyslightly broadeningandangled, posteriorlyconstrictedtoformdistinctneck, however, dorsalside notseparatedbyoccipitalfurrow, ratherbydorsumofneckbearingmicrosculpturein- steadofpunctation. Antennomere 1 flattenedclub-shapedandsmooth, segment 2 basally broadovoid, segment 3 club-shaped, segment 4 smallandisodiametrical, segments 5–11 withbasaldishes, articles 6–10 slightlytransverse (about 1.3× broaderthanlong). Head andpronotumwithmostlylacunosesculpture. Pronotumwithlateralportionexplanate andslightlyup-turned, edgewithafewshortsetae, lateralborderalmostevenlyarched, withoutposteriorcorners, withanumberofsmall, irregularlydistributedtubercles; disc mediallyratherconvex, withtwofaint, parallellongitudinalridges, laterallyimpressed aroundmiddleofsides. Elytrawithverythinmembranousposteriormarginextending fromsuturalcornertolateral 3/4 ofhindmargin. Elytralepipleuralridgepresentwithupturnedlateralalatiformportions, discdepressed; shouldersmoderatelydeveloped, surface foveolate, inbetweenpuncturestorulose. Legsshort, pro-andmesotibiawithseveralspi- nuloserows, metatibiawithlongitudinalctenidiumofspinulesindistalhalfindistalhalf. Abdomenwithsidesweaklyarcuate, secondsegmentwithparatergitesstronglybroaden- ingposteriorly (abdomenappearingconstrictedatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad. PosterioredgeoftergiteVIIwiththinpalisadefringe, median part of sternite VII as in Fig. 104 , sternite VIII ( Fig. 105 ), tergites IX ( Fig. 106 ), X ( Fig. 107 ) andaedeagusasinFigs 108–110 .

Distribution – Onlyknownfromarathernarrowareaaroundtheoriginal

typelocalityincentralNepal.

Remarks – Theoriginalmaterialof Oxytelopsisfranzi ismissing. Neither theholotypefrom “ Népal (central), zw.[ischen] Mulk[h]arkaundTarePati [TharePati, 28°00’16”N, 85°29’40”E]” inH. Franz’ scollection ( NHMW), nor thesupposedlyfemaleparatypeinthecollectionofH. Coiffait ( MNHP) are foundinthedepositoriesindicatedinthedescription. Thelistoftheauthor’ s typespecimens ( CoiffAit 1986) doesnoteveninclude Oxytelopsisfranzi. Thereforeaneotypemustbedesignatedtoclarifythetaxonomicstatusofthe taxon – luckily, asuitablespecimenexistsfromalmostexactlytheoriginal typelocalityandthereforethereisnodoubtaboutthecorrectinterpretation of the name.

Figures

Figs 86–87. Habitus of Anotylus species. 86 = A. franzi (Coiffait, 1982), 87 = A. jaljulensis (Coiffait, 1984)

Figs 104–110. Anotylus franzi (Coiffait, 1982), male. 104 = median part of sternite VII, 105 = sternite VIII, 106 = tergite IX, 107 = tergite X, 108 = aedeagus, frontal view, 109 = median lobe, lateral view, 110 = paramere, lateral view. Scales: 0.10 mm for Figs 108–110, 0.12 mm