Dolichoiulus Verhoeff, 1900

Enghoff, Henrik, 2012, Three new species of Dolichoiulus millipedes from the underground of Gran Canaria, with notes on the circumscription of the genus (Diplopoda, Julida, Julidae), European Journal of Taxonomy 15, pp. 1-12 : 2-3

Genus Dolichoiulus Verhoeff, 1900

Dolichoiulus Verhoeff, 1900: 85 .

Nesopachyiulus Attems, 1902: 599 , 607.

Trichopachyiulus Verhoeff, 1910: 194 .

Anagaiulus Enghoff, 1992: 144 syn. nov.

See Enghoff (1992) for an in-depth discussion of Dolichoiulus . Anagaiulus was erected by Enghoff (1992) for a species which differs from the Dolichoiulus species in having a whorl of metazonital setae. Enghoff (1992) realized that this is a poor diagnostic character but preferred not to ‘dilute’ the already rather vague concept of Dolichoiulus by including a species with metazonital setae. Since then, two lines of evidence have emerged that contradict separate status for Anagaiulus :

One of the new species described in the present paper has metazonital setae; the other doesn’t, but 1. apart from this, and a size difference, the two species are as good as identical.

A preliminary molecular phylogenetic analysis by I. Sanmartín (unpublished) suggests that 2. Anagaiulus is nested deeply inside Dolichoiulus .

Diagnosis

(from Enghoff 1992, emended): Pachyiulini . Eyes present or absent. No frontal setae. Mostly without metazonital setae (exceptions: D. blancatypa ( Enghoff, 1992) comb. nov. and D. oromii sp. nov.). Metazonital striae mostly distinct also dorsally. No grooves in suture. No (or almost no) preanal projection. Preanal pilosity marginal or extensive. Anal valve pilosity extensive. Male mandibles mostly without lobes. First and second male legs typically julid. Male legs mostly with ventral pads. Anterior gonopod spoon-shaped, with a mesal ridge, a lateral prominence, and two apical denticles (which may be fused or reduced). Posterior gonopod without a sperm fovea, with a mesomerital process and with two opisthomerital processes, posterior process varying from short-triangular to flagelliform; one or both opisthomerital processes sometimes more or less reduced.

Included species

Dolichoiulus blancatypa ( Enghoff, 1992) , comb. nov. ex Anagaiulus , D. typhlocanaria sp. nov., D. oromii sp. nov., D. longunguis sp. nov., plus the 56 species included in Enghoff (1992).