Archiphthiracarella baloghorum Mahunka, 1997

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 43-59

South African, endemic brevisetus Plonaphacarus (Mahunka, 1984)

Hoplophthiracarus brevisetus Mahunka, 1984

Hoplophthiracarus brevisetus: Niedbała 1992 .

Hoplophthiracarus (Plonaphacarus) brevisetus: Mahunka 1992 .

Plonaphacarus brevisetus: Niedbała 2001a , Liu et al. 2011b, Niedbała & Liu 2018.

Rhacaplacarus (Rhacaplacarus) brevisetus: Subías 2004 .

Hoplophorella (Rhacaplacarus) breviseta: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1984a, 1992, Niedbała 1992, 2001a, 2017c, Subías 2022).

Afrotropical, Tanzania, endemic brevisetus Notophthiracarus Niedbała, 2021

Hoplophthiracarus (Notophthiracarus) brevisetus: Subías 2022 .

South Africa: Eastern Cape (Niedbała et al. 2021).

South African, endemic brevispinus Hoplophthiracarus Niedbała, 2019

Hoplophthiracarus (Hoplophthiracarus) brevispinus: Subías 2022 .

Afrotropic: Ivory Coast ( Niedbała & Ermilov 2019a).

Afrotropical, Ivory Coast, endemic bruneiensis Notophthiracarus (Calyptophthiracarus) Subías, 2010 synonym of Plonaphacarus aculeatus (Mahunka, 1995)

bryani Austrotritia ( Jacot, 1929)

Euphthiracarus (Indotritia) bryani Jacot, 1929

Indotritia bryani: Jacot 1934c .

Austrotritia bryani: Niedbała 1994c , Liu et al. 2009b, Liu & Zhang 2014 a, Niedbała & Liu 2018, Subías 2022.

Australasia: Hawaii Islands ( Jacot 1929, 1934c, Niedbała 1994c, 2012a, Subías 2022).

Australasian, Hawaii Islands, endemic bryobius Phthiracarus Jacot, 1930

Phthiracarus setosellum bryobium Jacot, 1930

Phthiracarus bryobius: Jacot 1938 , Niedbała 1986b, 1992, 1994a, 2000, 2002a, 2008e, f, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Mahunka & Mahunka-Papp 2004, Ni & Liu 2014, Niedbała & Liu 2018, Liu 2021b.

Phthiracarus (Archiphthiracarus) bryobius: Mahunka 1990b , Niedbała et al. 2021, Subías 2022, Niedbała & Błoszyk 2022.

Phthiracarus crinitossimilis Willmann, 1939 syn. by Niedbała 1986b.

Archiphthiracarus crinitossimilis: Balogh & Mahunka 1980 a, 1983 .

Archiphthiracarus gobiensis Mahunka, 1981 syn. by Niedbała 1986b.

Archiphthiracarus gobiensis: Balogh & Mahunka 1983 .

Phthiracarus lanatus Feider et Suciu, 1957 syn. by Niedbała 1986b.

Phthiracarus lanatus: Krivolutsky 1975 , Mahunka & Mahunka-Papp 2004.

Paraphthiracarus lanatus: Aoki 1980a .

Archiphthiracarus lanatus: Balogh & Mahunka 1980 a, 1983 .

Phthiracarus lanatus sensu Feider et Suciu, 1957

Phthiracarus opacus Niedbała, 1986 : Beck et al. 2014 (see note of P. opacus )

Phthiracarus pilosus Sergienko, 1987 syn. by Niedbała 1994a.

Phthiracarus pilosus: Niedbała 1992 .

Archiphthiracarus hungaricus Balogh et Mahunka, 1979 syn. by Niedbała 1992.

Archiphthiracarus hungaricus: Balogh & Mahunka 1980 a, 1983 , Niedbała 1986b, 1992, Mahunka & Mahunka-Papp 2004, Subías 2004.

Nearctic: Canada (British Columbia), USA (Connecticut, New York, California, Wyoming, South Dakota, Wisconsin, Iowa, New York, North Carolina, Utah). Neotropic: Costa Rica, Bolivia. Palaearctic: Portugal, Spain, Norway, Finland, Denmark, Germany, Switzerland, Poland, Czech Republic, Austria,

Slovakia, Hungary, Romania, Slovenia, Latvia, Lithuania, Ukraine (western and eastern part and Crimea), Georgia, Russia (North, Central and South European Territories, Western and Eastern Siberia), Turkey, Uzbekistan, Kyrgyzstan, Mongolia, Russia (Far East), Nepal, China (Beijing), South Korea, China (Beijing, Heilongjiang), Japan ( Aoki 1980a, Balogh & Mahunka 1980 a, 1983, Beck et al. 2014, Feider & Suciu 1957, Jacot 1930, 1938, Kaczmarek & Niedbała 2002, Liu 2021 b, Mahunka 1981, 1990b, 1993, Mahunka & Mahunka-Papp 1999, 2003, 2004, Mahunka et al. 2013, Marshall et al. 1987, Ni & Liu 2014, Niedbała 1986b, 1992, 1994a, 2000, 2002a, 2008e, f, 2011b, 2012b, 2015b, Niedbała & Błoszyk 2022, Niedbała & Ermilov 2014a, Niedbała et al. 2020, Niedbała & Starý 2015b, 2015e, Olszanowski et al. 1996, Ryabinin et al. 2018, Seniczak et al. 2021, Sergienko 1987, Subías 2004, Weigmann et al. 2015, Willmann 1939, Subías 2022).

Holarctic, Panpalaearctic, probably introduced to Bolivia and Costa Rica

Notes. Subías (2022) recognized Hoplophora straminea C.L. Koch as probably congeneric with this species. On the basis of the redescription by Jacot (1936), H. straminea cannot be unambiguously classified.

Subías (2022) considered Phthiracarus clavatus Parry, 1979 as congeneric with this species. See note of P. clavatus .

Subías (2022) recognized Archiphthiracarus tzanoudakisi Mahunka, 1979 with this species. See note of Phthiracarus montanus Pérez-Íñigo, 1969 .

Subías (2022) considered Phthiracarus wallworki Niedbała, 1984 as congeneric with this species. See note of P. wallworki .

Subías (2022) identified this species as congeneric with P. opacus Niedbała, 1986 , possibly agreeing with the opinion of Beck et al 2014. See note of P. opacus .

buffaloensis Atropacarus (Hoplophorella) Niedbała, 2006

Atropacarus (Hoplophorella) buffaloensis: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) buffaloensis: Subías 2022 .

Australasia: Australia (Victoria) ( Niedbała 2006d, 2012a, Subías 2022).

Australasian, Australia, endemic buffaloensis Notophthiracarus Niedbała, 2006

Notophthiracarus buffaloensis: Liu & Zhang 2014 d, Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) diversisetosa Subías, 2010 : Niedbała & Liu 2018.

Australasia: Australia (Victoria) ( Niedbała 2006d, 2012a, Subías 2022).

Australasian, Australia, endemic bulbifer Oribotritia (Oribotritia) (Mahunka, 1987)

Berndotritia bulbifer Mahunka, 1987

Berndotritia bulbifer: Mahunka 1988d, 1990a , 1991b.

Oribotritia bulbifer: Mahunka 1990, 2009b , Niedbała 2000, 2004c, Subías 2004, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) bulbifer: Subías 2022 .

Orient: Thailand, Malaysia (Sabah, Sarawak, West Malaysia), Southern Vietnam, Indonesia (Kalimantan, West Sumatra), Andaman Islands ( Balogh & Balogh 2002, Ermilov et al. 2012, Mahunka 1987f, 1988d, 1990a, 1991b, 2008b, Niedbała 2000, 2004c, 2008b, 2011a, Niedbała & Ermilov 2013 a, 2014b, Niedbała & Starý 2015e, Subías 2022, Vu 2013).

Oriental, broad bulbiferus Phthiracarus (Mahunka, 1996)

Archiphthiracarella bulbifera Mahunka, 1996

Archiphthiracarella baloghorum Mahunka, 1997 : nom. nov. for Phthiracarus insularis Balogh, 1962 nom. praeoc. by Phthiracarus insularis Jacot, 1934

Phthiracarus baloghorum: Niedbała 2008c , Niedbała & Liu 2018.

Phthiracarus minor Niedbała, 2001 : nom. nov. for Phthiracarus insularis Balogh, 1962 nom. praeoc. by Phthiracarus insularis Jacot, 1934

Phthiracarus minor Niedbała, 2001 syn. by Mahunka 2002.

Phthiracarus minor Niedbała, 2001 : Niedbała 2004d, 2008c.

Phthiracarus insularis Balogh, 1962 nov. nom. by Mahunka 1997a.

Phthiracarus insularis Balogh, 1962 nov. nom. by Niedbała 2001a.

Phthiracarus insularis: Niedbała 1986b, 1992 , 2001, Mahunka 1997a, b.

Archiphthiracarella insularis: Mahunka 2002 .

Archiphthiracarella bulbifera Mahunka, 1996 syn. by Niedbała 2001a.

Phthiracarus (Archiphthiracarus) bulbiferus: Subías 2022 .

Microphthiracarus baloghi Mahunka, 1982 syn. by Niedbała 2017a.

Microphthiracarus baloghi Mahunka, 1982 nom. praeoc. by Subías 2004.

Microphthiracarus baloghi: Niedbała 1986b, 1992 .

Phthiracarus baloghi: Niedbała 2001a .

Phthiracarus neominatus (lapsus linguae) Subías, 2004 nov. nom. by Niedbała 2017a.

Phthiracarus neonominatus: Niedbała 2008c .

Afrotropic: Ethiopia, Madagascar ( Balogh 1962, Mahunka 1982d, 1996b, 1997a, b, 2002, Niedbała 1986b, 1992, 2001a, 2004d, 2008c, 2017a, Niedbała & Starý 2016a, Subías 2022).

Afrotropical, central and eastern parts

Note. The name baloghorum was proposed by Mahunka (1997a) to replace Phthiracarus insularis Balogh, 1962 which was preoccupied by Phthiracarus insularis Jacot, 1934 . Niedbała (2001a) proposed the name minor for Phthiracarus insularis Balogh, 1962 , and synonymized this species with Archiphthiracarus bulbiferus Mahunka, 1996 . Mahunka (2002) treated Phthiracarus minor Niedbała, 2001 as synonym of Archiphthiracarella baloghorum Mahunka, 1997 but considered Archiphthiracarella bulbifera Mahunka, 1966 as valid species. In this catalogue, we accept the view of Niedbała (2001a) and Subías (2004) that Archiphthiracarella bulbifera Mahunka, 1966 is identical to Phthiracarus insularis Balogh, 1962 . So, the name P. bulbiferus ( Mahunka, 1996b) for P. insularis Balogh, 1962 has priority over P. baloghorum ( Mahunka, 1997a) and P. minor Niedbała, 2001 .

bulbosus Arphthicarus Niedbała, 2006

Arphthicarus bulbosus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) bulbosus: Subías 2022 .

South Africa: Limpopo ( Niedbała 2006c, Subías 2022).

South African, endemic bulbus Arphthicarus Niedbała, 2004

Arphthicarus bulbus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) bulbus: Subías 2022 .

Neotropic: Antilles ( Cuba) ( Niedbała 2004a, Subías 2022).

Neotropical, Cuba, endemic cacainus Hoplophthiracarus Niedbała, 2004

Hoplophthiracarus cacainus: Niedbała & Liu 2018 .

Hoplophthiracarus (Hoplophthiracarus) cacainus: Subías 2022 .

Neotropic: Guatemala ( Niedbała 2004a, Subías 2022).

Neotropical, Guatemala, endemic cadizi Phthiracarus Niedbała, 1986

Phthiracarus cadizi: Niedbała 1992, 2011b , 2012b, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) cadizi: Subías 2004 .

Phthiracarus (Phthiracarus) cadizi: Subías 2022 .

Palaearctic: Spain ( Niedbała 1986 d, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, Spain, endemic caelestis Steganacarus (Steganacarus) Niedbała, 1984

Steganacarus caelestis Niedbała, 1984

Steganacarus (Steganacarus) caelestis: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Palaearctic: Iran ( Niedbała 1984f, 1986b, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, Iran, endemic cajanumaensis Austrophthiracarus Niedbała et Illig, 2007

Hoplophthiracarus (Protophthiracarus) cajanumaensis: Subías 2022 .

Austrophthiracarus cajanumaensis: Liu & Zhang 2016 a, Niedbała & Liu 2018 .

Neotropic: Ecuador ( Niedbała & Illig 2007 a, Niedbała & Roszkowska 2017, Subías 2022).

Neotropical, Ecuador, endemic caliginosus Notophthiracarus Niedbała, 1989 synonym of Notophthiracarus australis Ramsay, 1966

callainii Steganacarus (Tropacarus) Bernini, Bernini et Avanzati, 1989

Steganacarus (Tropacarus) callainii: Niedbała 1992, 2011b , 2012b, Niedbała & Liu 2018, Subías 2022. Palaearctic: Tunisia, Algeria ( Bernini et al. 1989, Niedbała 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, only in Maghreb countries calotropicus Atropacarus (Hoplophorella) Haq et Xavier, 2005

Atropacarus (Hoplophorella) calotropicus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) calotropica: Subías 2022 .

Orient: India (Kerala) ( Haq & Xavier 2005, Subías 2022).

Oriental, India, endemic

Note. Questionable species with strange shape of rostral setae. It is similar to Atropacarus (Hoplophorella) hamatus (Ewing, 1909) by the shape of interlamellar, notogastral and ad 2 setae.

calugarae Notophthiracarus Niedbała, 1987

Notophthiracarus calugari: Niedbała 1992, 2000 , Liu & Zhang 2014 d, Niedbała & Liu 2018, Niedbała et al. 2021.

Notophthiracarus calugarae: Niedbała & Szywilewska-Szczykutowicz 2017 .

Notophthiracarus (Notophthiracarus) calugari: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) calugarae: Subías 2022 .

Australasia: Australia (Victoria) ( Balogh & Balogh 2002, Niedbała 1987, 1992, 2000, 2012a, Niedbała et al. 2021, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, Australia, indigenous canadaris Protoribotritia Jacot, 1938

Protoribotritia canadaris: Niedbała 2002a, 2007c , Niedbała & Liu 2018, Subías 2022.

Nearctic: Canada (Yukon, Alberta, Ontario, Nova Scotia ), USA (Oregon, Connecticut, New York, New Mexico) ( Jacot 1938, Marshall et al. 1987, Niedbała 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022).

Nearctic, north and eastern parts with southern disjunction (New Mexico)

canariensis Calyptophthiracarus Pérez-Íñigo et Peña,1996 synonym of Austrophthiracarus heterotrichus ( Mahunka, 1979)

candidulus Austrophthiracarus (Niedbała, 1983)

Hoplophthiracarus candidulus Niedbała, 1983

Calyptophthiracarus candidulus: Niedbała 1986b, 1992 .

Notophthiracarus (Calyptophthiracarus) candidulus: Subías 2004 .

Hoplophthiracarus (Calyptophthiracarus) candidulus: Subías 2022 .

Hoplophthiracarus meridionalis Sergienko, 1992 syn. by Niedbała 2011b.

Hoplophthiracarus meridionalis: Subías 2012

Notophthiracarus (Calyptophthiracarus) meridionalis: Subías 2004 .

Austrophthiracarus candidulus: Niedbała 1994a, 2011b , 2012b, Liu & Zhang 2016 a, Niedbała & Liu 2018.

Palaearctic: Ukraine (western part and Crimea), Georgia, Russia (South European Territory) ( Niedbała 1983a, 1986b, 1992, 2011b, 2012b, 2015b, Sergienko 1992, Subías 2022).

Western Palaearctic, known from Black Sea-Pontic area canestrini Tritia Michael, 1898 species inquirenda, incertae sedis

Tritia canestrini: Niedbała 2011b , Niedbała & Liu 2018.

synonym of Acrotritia ardua: Hammen (1959) , Castagnoli & Pegazzano 1985, Subías 2022.

Palaearctic: Italy ( Michael 1898)

capevidalensis Notophthiracarus Niedbała, 2006

Notophthiracarus capevidalensis: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) capevidalensis: Subías 2022 .

South Africa: KwaZulu-Natal ( Niedbała 2006c, Subías 2022).

South African, endemic capillatus Notophthiracarus Niedbała, 1989

Notophthiracarus capillatus: Niedbała 1992, 2000 , 2006d, Niedbała & Penttinen 2007, Liu & Zhang 2014 d, Niedbała & Szywilewska-Szczykutowicz 2017, Niedbała & Liu 2018, Niedbała et al. 2021. Notophthiracarus (Notophthiracarus) capillatus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) capillatus: Subías 2022 .

Australasia: Australia (Victoria, New South Wales, Tasmania) ( Balogh &

Balogh 2002, Niedbała 1989b, 1992, 2000, 2006d, 2012a, 2014b, 2017b, Niedbała et al. 2021, Niedbała & Penttinen 2007, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022). Australasian, south-eastern part of Australia capitanea Oribotritia (Oribotritia) Niedbała et Corpuz-Raros, 1998

Oribotritia capitanea Niedbała et Corpuz-Raros, 1998

Oribotritia capitanea: Niedbała 2000 , Subías 2004, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) capitanea: Subías 2022 .

Orient: Philippines ( Niedbała 2000, Niedbała & Corpuz-Raros 1998, Subías 2022).

Oriental, Philippines, endemic capitata Hoploderma Hall, 1911 species inquirenda, incertae sedis

Hoploderma capitata Hall, 1911 : Marshall et al. 1987, Niedbała & Liu 2018.

Phthiracarus (Phthiracarus) capitatus ( Hall, 1911) : Subías 2004.

Nearctic: USA (Connecticut) ( Hall 1911, Marshall et al. 1987, Subías 2004).

carinata Austrotritia Niedbała, 2000

Austrotritia carinata: Niedbała 2006 d, Liu et al. 2009b , Liu & Zhang 2014 a, Niedbała & Liu 2018, Subías 2022.

Australasia: Papua New Guinea, Australia (northern and western parts) ( Niedbała 2000, 2006d, 2012a, Subías 2022).

Australasian, Papua New Guinea and northern and western parts of Australia carinata var. pulcherrima Hoplophora Berlese, 1887 = Steganacarus (Tropacarus)

pulcherrimus (Berlese, 1887)

carinatus Arphthicarus Niedbała, 2004

Arphthicarus carinatus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) carinatus: Subías 2022 .

Neotropic: Antilles ( Trinidad) ( Niedbała 2004a, Niedbała & Ermilov 2017b, Subías 2022).

Neotropical, Trinidad, endemic carinatus Euphthiracarus (Euphthiracarus) Liu, Wu et Chen, 2011

Euphthiracarus carinatus Liu, Wu et Chen, 2011

Euphthiracarus carinatus: Liu et al. 2011 d, 2012 , Niedbała & Ermilov 2018 a, Niedbała & Liu 2018, Liu 2021b.

Euphthiracarus (Euphthiracarus) carinatus: Subías 2022 .

Palaearctic: Nepal. Orient: China (Guizhou) (Liu 2021b, Liu et al. 2011 d, 2012, 2013, Niedbała & Ermilov 2018a, Subías 2022).

Palaearctic, Nepal and Oriental, China carinatus Steganacarus (Tropacarus) (C.L. Koch, 1841)

Hoplophora carinata C.L. Koch, 1841

Hoplophora carinata: Berlese 1887b , Oudemans 1937

Hoplophora carinatum: Ewing 1917b .

Hoploderma carinatum: Michael 1898 .

non Hoploderma carinatum: Michael 1890 .

Calhoplophora carinata: Jacot 1930 .

Tropacarus carinatus: Sellnick 1928, 1960 , Willmann 1931, Balogh 1943, Feider & Suciu 1957, Hammen 1959, Bernini 1971, Lions 1973, Krivolutsky 1975, Niedbała et al. 1981, 1982, 1990, Balogh & Mahunka 1983.

Steganacarus carinatus: Balogh & Mahunka 1980 a, 1983 , Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008.

Steganacarus (Steganacarus) carinatus: Niedbała 1986b, 1992 .

Steganacarus (Tropacarus) carinatus: Bernini & Avanzati 1989a , Mahunka 1990b, Avanzati et al. 1995, Niedbała 1994b, 2000, 2008f, 2011b, 2012b, Olszanowski et al. 1996, Mahunka & Mahunka-Papp 2004, Subías 2022, Weigmann 2006, Niedbała & Liu 2018, Niedbała et al. 2021.

Steganacarus (Tropacarus) carinatus carinatus: Subías 2022 .

Steganacarus lucidus Sergienko, 1994 (?): Niedbała & Liu 2018.

Palaearctic: Spain, France, England, Sweden, Finland, Denmark, Germany, Switzerland, Austria, Italy (continental and Sicily), Poland, Czech Republik, Slovakia, Hungary, Slovenia, Croatia, Macedonia, Greece, Bulgaria, Romania, Estonia, Latvia, Lithuania, Ukraine (western and eastern parts and Crimea), Azerbaijan, Caucase, Russia (North, Central and South European Territory, Western Siberia), Algeria, Turkey, Syria, Iran, India (Kashmir) Ahaniazad et al. 2016, Avanzati et al. 1995, Balogh 1943, Balogh & Mahunka 1980 a,1983, Berlese 1887b, Bernini & Avanzati 1988b, 1989a, Castagnoli & Pegazzano 1985, Ewing 1917b, Feider & Suciu 1957, Hammen 1959, C.L. Koch 1841, Krivolutsky 1975, Lions 1973, Mahunka 1979, 1982c, 1990b, 1993b, Mahunka et al. 2013, Mahunka & Mahunka-Papp 2003, 2004, Mahunka & Paoletti 1984, Michael 1898, Niedbała 1986b, 1992, 2000, 2008f, 2011b, 2012b, Niedbała et al. 2021, Olszanowski et al. 1996, Oudemans 1915a, 1937, Pigliucci et al. 1990, Schatz 1983, Schuster 1956, Sellnick 1928, 1960, Subías 2022, Weigmann et al. 2015, Willmann 1931.

South-Western Palaearctic, Paneuropean and also known from Maghrebian, Black Sea-Pontic, Anatolian, Iranian and Kashmiran areas

Notes. Steganacarus lucidus Sergienko, 1994 is probably conspecific with Steganacarus (T.) carinatus , because three different characters (structure of surface of body, shape of prodorsal setae, position of lyrifissures ia) are within the range of variability of S. (T.) carinatus .

Subías (2022) considered Steganacarus (Tropacarus) lucidus Sergienko, 1994 (synonym of Steganacarus (Tropacarus) carinatus ) as congeneric with Steganacarus (Tropacarus) malatestai Avanzati, Baratti et Bernini, 1994 . See the comparison of S. (T.) malatestai in Niedbała 2011b.

carlosi Steganacarus (Steganacarus) Niedbała, 1984

Steganacarus striculus: Pérez-Íñigo 1972 .

Steganacarus (Steganacarus) carlosi: Niedbała 1985b, 1986b , 1992, 2001a, 2011b, 2012b, Avanzati et al. 1994, Niedbała & Liu 2018, Subías 2022.

Palaearctic: Canary Islands (La Gomera, Tenerife) ( Avanzati et al. 1994, Niedbała 1984e, 1986b, 1992, 2001a, 2011b, 2012b, Pérez-Íñigo 1972, Pérez-Íñigo & Peña,1996, Subías 2022).

Western Palaearctic, Canary Islands, endemic caroli Synichotritia Walker, 1965

Synichotritia caroli: Mahunka 1990a , Lions & Norton 1999, Niedbała 2002a, 2007c, Niedbała & Liu 2018, Subías 2022.

Nearctic: Canada (British Columbia), USA (Oregon, Mississippi, North Carolina, California) ( Lions & Norton 1999, Mahunka 1990a, Marshall et al. 1987, Niedbała 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022, Walker 1965).

Nearctic, western part with south-eastern disjunction (Mississippi)

carolinae Oribotritia (Oribotritia) Jacot, 1930

Oribotritia carolinae Jacot, 1930

Indotritia carolinae ( Jacot, 1930) : Märkel 1964, Marshall et al. 1987.

Oribotritia carolinae: Niedbała 2002a , Subías 2004, Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) carolinae: Subías 2022 .

Nearctic: USA (North Carolina, Virginia) ( Jacot 1930, 1933, Marshall et al. 1987, Niedbała 2002a, Subías 2022).

Nearctic, eastern part carusoi Steganacarus (Steganacarus) Bernini et Avanzati, 1989

Steganacarus cf. applicatus: Bernini 1984 .

Steganacarus (Steganacarus) carusoi: Niedbała 1992, 2011b , 2012b, Avanzati et al. 1995, Niedbała & Liu 2018, Niedbała et al. 2022, Subías 2022

Palaearctic: Spain, France, Corsica, Tuscan Archipelago, Italy, Sardinia, Morocco, Algeria, Tunisia ( Avanzati et al. 1995, Bernini & Avanzati 1989 c, Niedbała 1992, 2011b, 2012b, 2015b, Niedbała et al. 2022, Subías 2022).

Western Palaearctic, reported from Maghrebian and Atlanto-Mediterranean subregions catalaucus Arphthicarus Niedbała, 2004

Arphthicarus catalaucus: Niedbała & Liu 2018 .

Neotropic: Dominican Republic ( Niedbała 2004a, Subías 2022).

Neotropical, Dominican Republic, endemic catalaucus Phthiracarus Niedbała, 2004

Phthiracarus catalaucus: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) catalaucus: Subías 2022 .

Neotropic: Jamaica ( Niedbała 2004a, Subías 2006).

Neotropical, Dominican Republic, endemic cataracta Neosteganacarus Balogh et Mahunka, 1992 synonym of Steganacarus (Steganacarus) rafalskii (Niedbała, 1982)

cathayanus Euphthiracarus (Euphthiracarus) Mahunka, 2000

Euphthiracarus cathayanus Mahunka, 2000

Euphthiracarus cathayanus: Wang et al. 2002 , Niedbała 2004c, Chen et al. 2010, Liu et al. 2012, Niedbała & Liu 2018, Liu 2021b.

Euphthiracarus (Euphthiracarus) cathayanus: Subías 2022 .

Orient: China ( Hong Kong) ( Chen et al. 2010, Liu 2021b, Liu et al. 2012, Mahunka 2000, Niedbała 2004c, Subías 2022, Wang et al. 2002).

Oriental, China ( Hong Kong), endemic caudatus Austrophthiracarus ( Balogh et Mahunka, 1977)

Phthiracarus caudatus Balogh et Mahunka, 1977

Phthiracarus caudatus: Niedbała 1986b, 1992 .

Austrophthiracarus caudatus: Niedbała 1994a, 2003a , 2004a, Liu & Zhang 2016 a, Niedbała & Liu 2018. Notophthiracarus (Notophthiracarus) caudatus: Subías 2004

Hoplophthiracarus (Notophthiracarus) caudatus: Subías 2022 .

Neotropic: Costa Rica, Bolivia ( Balogh & Balogh 1987, 1988, 2002, Balogh & Mahunka 1977, Niedbała 1986b, 1992, 1994a, 2003a, 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Mesoamerica and Guyano-Brasilian caulis Notophthiracarus Niedbała, 2017

Notophthiracarus caulis: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) caulis: Subías 2022 .

Australasia: Australia (Queensland) (Niedbała 2017b, Subías 2022).

Australasian, Australia, endemic cavernosus Notophthiracarus (Mahunka, 1987)

Hoplophorella cavernosa Mahunka, 1987

Notophthiracarus cavernosus: Mahunka 1990c , Niedbała 1994 a, Niedbała & Liu 2018.

Hoplophorella cavernosa: Niedbała 1992 .

Hoplophorella (Kakophthiracarus) cavernosa: Subías 2004 .

Hoplophthiracarus (Steganacarellus) cavernosus: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1987c, 1990 c, Niedbała 1992, 1994a, 2017c, Subías 2022).

Afrotropical, Tanzania, endemic cavernosus Phthiracarus ( Wallwork, 1977)

Hoplophthiracarus cavernosus Wallwork, 1977

Phthiracarus cavernosus: Niedbała 1985b, 1992 , Niedbała & Liu 2018.

Notophthiracarus cavernosus: Niedbała 2001a .

Phthiracarus (Archiphthiracarus) cavernosus: Subías 2022 .

Afrotropic: Saint Helena Island ( Balogh & Balogh 2002, Niedbała 1985b, 1992, 2001a, Subías 2022, Wallwork 1977).

Afrotropical, Saint Helena Island, endemic cavernosus Protophthiracarus Niedbała, 2002

Protophthiracarus cavernosus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) neonominata Subías 2004 : Niedbała & Liu 2018.

Nearctic: USA (Arizona) ( Niedbała 2002a).

Nearctic, USA, endemic cavernus Arphthicarus Niedbała, 2012

Arphthicarus cavernus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) caverna: Subías 2022 .

Afrotropic: Ethiopia ( Niedbała & Ermilov 2012, 2014a, 2022c, Subías 2022).

Afrotropical, Ethiopia, endemic cazanicus Plonaphacarus ( Feider et Călugăr, 1969)

Hoplophthiracarus cazanicus: Feider et al. 1969 , Feider & Călugăr 1970, Balogh & Mahunka 1983, Niedbała 1986b, 1992, Subías 2004.

Plonaphacarus cazanicus: Niedbała & Liu 2018 .

Hoplophthiracarus (Hoplophthiracarus) cazanicus: Subías 2022 .

Palaearctic: Romania ( Balogh & Mahunka 1983, Feider & Călugăr 1969, 1970, Feider et al. 1969, Mahunka et al. 2013, Niedbała 1986b, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, Romania, endemic cernuus Euphthiracarus (Euphthiracarus) Walker, 1965

Euphthiracarus cernuus Walker, 1965

Euphthiracarus cernuus: Niedbała 2002a, 2007c , Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) cernuus: Subías 2022 .

Nearctic: Canada (British Columbia, Nova Scotia, Vancouver ), USA (Oregon, Utah, California, Washington) ( Marshall et al. 1987, Niedbała 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022, Walker 1965). Nearctic, western part chaliyamensis Atropacarus (Hoplophorella) Haq et Xavier, 2005

Atropacarus (Hoplophorella) chaliyamensis: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) chaliyamensis: Subías 2022 .

Orient: India (Kerala) ( Haq & Xavier 2005, Subías 2022).

Oriental, India, endemic

Note. Questionable species with strange shape of rostral setae. It is similar to Atropacarus (Hoplophorella) singularis ( Sellnick, 1959) by the shape of setae ad 3.

chatham Notophthiracarus Liu et Zhang, 2015

Notophthiracarus chatham: Niedbała & Liu 2018 .

Hoplophthiracarus (Protophthiracarus) chatham: Subías 2022 .

Australasia: New Zealand ( Liu & Zhang 2015b, Subías 2022).

Australasian, New Zealand, endemic chernovae Atropacarus (Atropacarus) Shtanchaeva et Subías, 2012

Atropacarus (Atropacarus) chernovae: Niedbała & Liu 2018 , Subías 2022.

Palaearctic: Abkhazia, Armenia ( Shtanchaeva & Subías 2012, Subías 2022).

Southern Palaearctic, Caucasus

Note. This species is probably conspecific with A. (A.) echinodiscus (Mahunka, 1982) (Niedbała 2015a) . cherokee Oribotritia (Oribotritia) Niedbała, 2002

Oribotritia cherokee Niedbała, 2002

Oribotritia cherokee: Subías 2004 , Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) cherokee: Subías 2022 .

Nearctic: USA (Arkansas) ( Niedbała 2002a, Subías 2022).

Nearctic, USA, endemic chiayiensis Oribotritia (Oribotritia) Niedbała, 2021

Oribotritia chiayiensis Niedbała, 2021

Oribotritia (Oribotritia) chiayiensis: Subías 2022 .

Orient: China (Taiwan) ( Niedbała & Ermilov 2021).

Oriental, China (Taiwan), endemic chichijimensis Oribotritia (Oribotritia) Aoki, 1980

Oribotritia chichijimensis Aoki, 1980

Oribotritia chichijimensis: Aoki 1980a , Suzuki 1982b, Niedbała, 2000, 2004c, 2011b, 2012b, Subías 2004, Liu et al. 2011a, Liu 2015a, 2021b, Niedbała & Liu 2018.

Oribotritia (Oribotritia) chichijimensis: Subías 2022 .

Oribotritia chichijimensis ryukyuensis Nakatamari, 1985 syn. by Niedbała 2000.

Orient: China ( Taiwan, Macau Peninsula, Fujian, Hainan). Palaearctic: Japan ( Aoki 1980a, Liu 2015a, 2021b, Liu et al. 2011a, Nakatamari 1985, Niedbała 2000, 2004c, 2011b, 2012b, Niedbała & Ermilov 2018b, 2020, 2021, Subías 2022, Suzuki 1982b).

Oriental (Chinese) introduced to Japanese subregion chichijimensis ryukyuensis Oribotritia Nakatamari, 1985 = Oribotritia (Oribotritia) chichijimensis Aoki, 1980 chilensis Protophthiracarus ( Balogh et Mahunka, 1967)

Notophthiracarus chilensis Balogh et Mahunka, 1967

Notophthiracarus chilensis: Niedbała 1986b, 1992 .

Protophthiracarus chilensis: Aoki 1980a , Balogh 1972, Niedbała 1994a, 2004 a, Niedbała & Liu 2018. Notophthiracarus (Protophthiracarus) chilensis: Subías 2004 .

Hoplophthiracarus (Protophthiracarus) chilensis: Subías 2022 .

Neotropic: Chile ( Balogh 1972, Aoki 1980a, Balogh & Balogh 1987, 1988, Balogh & Mahunka 1967, Niedbała 1986b, 1992, 1994a, 2004a, Subías 2022).

Neotropical, Chile, endemic chuxiongensis Plonaphacarus Liu, 2020

Plonaphacarus chuxiongensis: Liu 2021b .

Hoplophthiracarus (Hoplophthiracarus) chuxiongensis: Subías 2022 .

Orient: China (Yunnan) ( Liu 2020a, 2021b).

Oriental, China, endemic ciliosus Atropacarus (Atropacarus) Niedbała, 2002

Atropacarus ciliosus: Subías 2004 .

Atropacarus (Atropacarus) ciliosus: Niedbała & Liu 2018 , Subías 2022.

Nearctic: USA (Utah, Arizona) ( Niedbała 2002a, Subías 2022).

Nearctic, south-western part of USA clandestinus Protophthiracarus Niedbała, 2003

Protophthiracarus clandestinus: Niedbała 2004a, 2008e , Niedbała & Liu 2018.

Hoplophthiracarus (Protophthiracarus) clandestinus: Subías 2022 .

Neotropic: Costa Rica, Panama ( Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica clara Mesotritia Starý, 1992 synonym of Mesotritia maerkeli Sheals, 1965

clavata Acrotritia ( Märkel, 1964)

Rhysotritia clavata Märkel, 1964

Rhysotritia clavata: Niedbała & Schatz 1996 , Niedbała 2002b, 2003a, 2004a.

Acrotritia clavata: Subías 2004 , Niedbała & Illig 2007b, Niedbała 2008e, 2010b, 2012b, Liu 2015 d, Niedbała & Liu 2018, Niedbała & Ermilov 2022b.

Acrotritia clavata clavata: Subías 2004 .

Nearctic: Bermuda. Neotropic: Cocos Island, Mexico, Guatemala, Belize, Costa Rica, Panama, Galapagos Islands, Antilles, Colombia, Ecuador, Venezuela, Peru, Bolivia, Brazil, Argentina. Palaearctic: Poland (Palm House) ( Balogh & Balogh 1987, 1988, 2002, Behan-Pelletier et al. 1993, Ermilov & Niedbała 2013, Ermilov et al. 2016, Liu & Oconnor 2015, Märkel 1964, Niedbała 2002b, 2003a, 2004a, 2008e, 2010b, 2012b, 2014c, Niedbała & Ermilov 2015, 2017b, 2022b, Niedbała & Illig 2007b, Niedbała & Roszkowska 2017, Niedbała & Schatz 1996, Niedbała & Starý 2015b, Oliveira 2004, Oliveira et al. 2017, Schatz 1998, Subías 2022).

Nearctic and Neotropical, broad (large distribution), introduced to Palm house, Poland

Note. Subías (2022) synonymized this species with Acrotritia curticephala ( Jacot, 1938) . However, both species have completely different shape of trichobothria.

clavata Indotritia (Indotritia) Wallwork, 1977

Indotritia clavata Wallwork, 1977

Indotritia (Indotritia) clavata: Niedbała 1998a , Liu 2015c, Niedbała & Liu 2018, Subías 2022.

Afrotropic: Saint Helena Island ( Balogh & Balogh 2002, Niedbała 1998a, Subías 2022, Wallwork 1977).

Afrotropical, Saint Helena Island, endemic clavata sextiana Rhysotritia Lions, 1966 synonym of Acrotritia curticephala ( Jacot, 1938)

clavatosensillus Notophthiracarus (Notophthiracarus) Subías, 2010 synonym of Arphthicarus baloghi Niedbała, 2003

clavatus Arphthicarus Niedbała et Starý, 2014

Arphthicarus clavatus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) clavata: Subías 2022 .

Neotropic: Costa Rica ( Niedbała & Starý 2014e, Subías 2022).

Neotropical, Costa Rica, endemic clavatus Atropacarus (Atropacarus) Aoki, 1980

Atropacarus striculus var. clavatus Aoki, 1980

Atropacarus clavatus: Subías 2004 .

Atropacarus (Atropacarus) clavatus: Niedbała 1986b, 1992 , 2000, 2004c, 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Palaearctic: Japan. Orient: Thailand ( Aoki 1980a, Niedbała 1986b, 1992, 2000, 2004c, 2011b, 2012b, Subías 2022).

Oriental, introduced to Japanese subregion clavatus Euphthiracarus (Euphthiracarus) Niedbała, 2004

Euphthiracarus (Euphthiracarus) clavatus: Niedbała & Liu 2018 , Subías 2022.

Neotropic: Guatemala ( Niedbała 2004a, Subías 2022).

Neotropical, Guatemala, endemic clavatus Hoplophthiracarus Niedbała, 2014

Hoplophthiracarus clavatus: Niedbała & Liu 2018 .

Hoplophthiracarus (Hoplophthiracarus) clavatus: Subías 2022 .

Orient: Southern Vietnam ( Niedbała & Ermilov 2014b, Subías 2022).

Oriental, Southern Vietnam, endemic clavatus Phthiracarus Parry, 1979

Archiphthiracarus clavatus: Balogh & Mahunka 1983 .

Phthiracarus clavatus: Parry 1979a , Niedbała 1986b, 1992, 2008f, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Olszanowski & Niedbała 2000, Weigmann 2006, Niedbała & Liu 2018, Niedbała et al. 2021, Niedbała & Błoszyk 2022.

Phthiracarus (Archiphthiracarus) clavatus: Subías 2004 .

Phthiracarus stramineus sensu Berg et al., 1990

Nearctic: Canada. Palaearctic: Azores, England, Norway, Sweden, Finland, Denmark, Germany, Luxembourg, Switzerland, Italy, Poland, Czech Republic, Austria, Slovakia, Croatia, Serbia, Greece, Romania, Bulgaria, Lithuania, Ukraine (Crimea), Georgia, Russia (North and Central European Territories, Far East), Turkey ( Balogh & Mahunka 1983, Mahunka & Mahunka-Papp 2003, Niedbała 1986b, 1992, 2008f, 2011b, 2012b, Niedbała & Błoszyk 2022, Niedbała et al. 2020, Olszanowski et al. 1996, Parry 1979 a, Seniczak et al. 2021, Subías 2022).

Holarctic, Panpalaearctic

Notes. It is not understandable why Subías (2022) identified this species as Hoplophora straminea C.L. Koch, 1841 — species inquirenda whose taxonomical status is ambiguous. With regard to Jacot (1936) description of Phthiracarus stramineus based on topotypic material, P. clavatus differs by the trichobothria rounded distally (versus pointed distally in P. stramineus ), setae c 1-3 of notogaster situated in one row near anterior border (versus setae c 2 remote from anterior border more than setae c 1 and c 3), and setae ad 2 longer than other ano-adanal setae (versus ad 2 shorter).

Subías (2022) identified this species as P. bryobius Jacot, 1930 . Actually, these two species differ considerably by the length of notogastral setae: long, c 1> c 1 - d 1 or h 1>2 h 1 - ps 1 (versus these setae shorter than appriopiate distances in P. bryobius ).

Beck et al. (2014) identified this species as Phthiracarus borealis Trägårdh, 1910 . On the basis of two slides with specimens from the locus typicus deposited in the Rijksmuseum and a comparison with the description of Parry (1979), we conclude that P. clavatus may be synonymous with P. borealis . This opinion may be agreed with remark of Niedbała (1992, in page 52) that one of the recently described species could be synonymous with P. borealis .

clavellatus Hoplophthiracarus Niedbała et Corpuz-Raros, 1998

Hoplophthiracarus clavellatus: Niedbała 2000 , Niedbała et al. 2006, Subías 2004, Niedbała & Liu 2018.

Hoplophthiracarus (Hoplophthiracarus) clavellatus: Subías 2022 .

Orient: Philippines (Luzon, Leyte, Mindanao) ( Niedbała 2000, Niedbała & Corpuz-Raros 1998, Niedbała et al. 2006, Subías 2022).

Oriental, Philippinian clavifer Atropacarus (Hoplophorella) Mahunka et Mahunka-Papp, 2009

Hoplophorella clavifer Mahunka et Mahunka-Papp, 2009

Atropacarus (Hoplophorella) clavifer: Niedbała & Liu 2018 .

Hoplophthiracarus (Hoplophthiracarus) clavifer: Subías 2022 .

Afrotropic: Kenya ( Mahunka & Mahunka-Papp 2009a, Subías 2022).

Afrotropical, Kenya, endemic clavifer Notophthiracarus (Mahunka, 1988)

Phthiracarus clavifer Mahunka, 1988

Phthiracarus clavifer: Niedbała 1992, 2001a , 2004d.

Archiphthiracarus hamatus: Mahunka 1988b .

Notophthiracarus clavifer: Niedbała & Liu 2018 .

Phthiracarus (Phthiracarus) clavifer: Subías 2022 .

Afrotropic: Reunion Island ( Mahunka 1988b, Niedbała 1992, 2001a, 2004d, 2017a, Subías 2004).

Afrotropical, Reunion Island, indigenous claviger Hoploderma Pearce, 1906 species inquirenda, incertae sedis

Hoploderma claviger: Niedbała & Liu 2018 .

Phthiracarus (Phthiracarus) claviger: Subías 2004 .

Notophthiracarus pearcei Subías, 2010 syn. by Niedbała & Liu 2018.

Palaearctic: India (Sikkim) ( Pearce 1906, Subías 2022).

Note. It is not certain that this is a Phthiracarus species.

claviger Notophthiracarus Niedbała, 1993

Notophthiracarus claviger: Niedbała 2000, 2006 d , Liu & Zhang 2013 d, 2014d, 2015b, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) claviger: Subías 2022 .

Hoplophorella (Hoplophorella) claviger: Subías 2004 .

Australasia: Australia (Victoria, Tasmania), New Zealand ( Balogh & Balogh 2002, Niedbała 1993b, 2000, 2006d, 2012a, Subías 2022).

Australasian, southern part of Australia clavigerus Atropacarus (Atropacarus) ( Berlese, 1904)

Hoploderma clavigerum Berlese, 1904 : Castagnoli & Pegazzano 1985.

Steganacarus clavigerus: Hammen 1959 , Balogh & Mahunka 1979, 1980, 1983.

Atropacarus clavigerus: Kamill & Baker 1980 , Mahunka & Mahunka-Papp 1995, 2004, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Subías 2004.

Atropacarus (Atropacarus) clavigerus: Niedbała 1986b, 1992 , 1993b, 1994b, 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Steganacarus (Atropacarus) clavigerus: Weigmann 2006 .

Palaearctic: Germany, Austria, Hungary, Italy, Slovenia, Algeria ( Balogh & Mahunka 1979, 1980, 1983, Berlese 1904, Bernini et al. 1988, Castagnoli & Pegazzano 1985, Hammen 1959, Kamill & Baker 1980,

Mahunka & Mahunka-Papp 1995, 2004, Mahunka et al. 2013, Niedbała 1986b, 1992, 1994b, 2011b, 2012b, 2015b, Olszanowski et al. 1996, Schatz 1983, Subías 2022, Weigmann et al. 2015).

Western Palaearctic, known from Central Europe, North-Mediterranean and Maghrebian subregions clemens Phthiracarus Aoki, 1963

Phthiracarus clemens clemens: Suzuki 1982 a, Aoki 1980 .

Phthiracarus clemens: Krivolutsky 1975 , Suzuki 1982, Niedbała 1982a, 1986b, 1992, 2000, 2011b, 2012b, Wang et al. 2002, Chen et al. 2010, Ni & Liu 2014, Niedbała & Ermilov 2018a, 2022, Niedbała & Liu 2018, Liu 2021b, Niedbała et al. 2022.

Archiphthiracarus clemens: Balogh & Mahunka 1983 .

Phthiracarus (Phthiracarus) clemens: Subías 2022 .

Phthiracarus clemens kyushuensis Aoki, 1980 syn. by Niedbała 1986b.

Phthiracarus clemens kyushuensis: Niedbała 1992 .

Palaearctic: India (Kashmir), Nepal, South Korea, China (Heilongjiang, Jilin), Japan. Orient: China ( Taiwan, Zhejiang, Hunan, Guangdong) ( Aoki 1963, 1980, Balogh & Balogh 2002, Balogh & Mahunka 1983, Chen et al. 2010, Kaczmarek & Niedbała 2002, Liu 2021b, Ni & Liu 2014, Niedbała 1982a, 1986b, 1992, 2000, 2011b, 2012b, 2015b, Niedbała & Ermilov 2014a, 2018 a, 2021, 2022, Niedbała et al. 2022, Ryabinin et al. 2018, Subías 2022, Suzuki 1982a, Wang et al. 2002).

Oriental and Palaearctic species, known from Prooriental and Far Eastern provinces clemens kyushuensis Phthiracarus Aoki, 1980 = Phthiracarus clemens Aoki, 1963

closteros Acrotritia Niedbała, 2006

Acrotritia closteros: Niedbała 2011b, 2012b , Liu 2015 d, Niedbała & Liu 2018.

Palaearctic: Algeria, Italy ( Niedbała 2006a, 2011b, 2012b, 2015b, Subías 2022).

Western Palaearctic, Mediterranean subregion

Note. Subías (2022) considered this species as congeneric with A. curticephala ( Jacot, 1938) . However, the two species have completely different shape of trichobothria and different numbers of genital setae (7 pairs versus 9 pairs in A. curticephala ).

closteros Phthiracarus Niedbała, 2004

Phthiracarus closteros: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) closteros: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic cochlearia Hoplophorella Pérez-Íñigo et Baggio,1993 synonym of Atropacarus (Hoplophorella) hamatus (Ewing, 1909)

coetzeeae Notophthiracarus Niedbała, 2006

Notophthiracarus coetzeeae: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) coetzeeae: Subías 2022 .

South Africa: Northern Cape ( Niedbała 2006c, Subías 2022).

South African, endemic cognatus Phthiracarus Niedbała, 1988

Phthiracarus cognatus: Niedbała 1992, 2002a , 2007c, Niedbała & Liu 2018.

Phthiracarus rafalskii Niedbała, 1997 syn. by Niedbała 2002a.

Phthiracarus incertus from Canada: Niedbała 1992.

Phthiracarus (Archiphthiracarus) cognatus: Subías 2022 .

Nearctic: Canada (British Columbia, Saskatchewan, Ontario, Quebec), USA (Oregon, Washington, California, Tennessee, Vermont) ( Niedbała 1988e, 1992, 1997, 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022).

Nearctic, northern and western parts collaris Atropacarus (Hoplophorella) ( Balogh, 1958)

Steganacarus collaris Balogh, 1958

Steganacarus collaris: Niedbała 1986b, 1992 .

Hoplophorella collaris: Mahunka 1984b .

Atropacarus (Hoplophorella) collaris: Niedbała 2001 a, Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) collaris: Subías 2022 .

Afrotropic: Angola ( Balogh 1958, Balogh & Balogh 2002, Kamill & Baker 1980, Mahunka 1984b, Niedbała 1986b, 1992, 2001a, Subías 2022).

Afrotropical, Angola, endemic coloffi Indotritia Balogh et Balogh, 2002 synonym of Indotritia (Zeaotritia) brevisetosa Niedbała, 2000

colombianus Acrotritia Liu, 2015

Acrotritia colombiana: Niedbała & Liu 2018 , Subías 2022.

Neotropic: Colombia ( Liu 2015 d, Subías 2022).

Neotropical, Colombia, endemic colombianus Notophthiracarus Liu et Oconnor, 2015

Notophthiracarus colombianus: Niedbała & Liu 2018 .

Hoplophthiracarus (Besuchetacarus) colombianus: Subías 2022 .

Neotropic: Colombia ( Liu & Oconnor 2015, Subías 2022).

Neotropical, Colombia, endemic comatus Notophthiracarus Niedbała, 2000

Notophthiracarus comatus: Niedbała & Penttinen 2007 , Liu & Zhang 2013d, 2015b, Niedbała & Liu 2018.

Notophthiracarus (Calyptophthiracarus) comatus: Subías 2004 .

Hoplophthiracarus (Calyptophthiracarus) comatus: Subías 2022 .

Australasia: New Zealand ( Niedbała 2000, 2012a, Niedbała & Penttinen 2007, Subías 2022). Australasian, New Zealand, endemic comatus Phthiracarus Niedbała, 1983

Phthiracarus comatus: Suzuki 1982 , Niedbała 1986b, 1992, 2000, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) comatus: Subías 2022 .

Palaearctic: Russia (Far East) ( Niedbała 1983d, 1986b, 1992, 2000, 2011b, 2012b, Subías 2022).

Eastern Palaearctic, Russia, endemic commutabilis Phthiracarus Niedbała, 1983

Phthiracarus commutabilis: Suzuki 1982 , Niedbała 1986b, 1992, Niedbała & Liu 2018.

Phthiracarus (Phthiracarus) commutabilis: Subías 2004 .

Palaearctic: Kyrgyzstan ( Niedbała 1983b, 1986b, 1992, 2011b, 2012b, 2015b, Subías 2022).

Central Palaearctic, Kyrgyzstan, endemic

Note. Subías (2022) considered this species as congeneric with P. compressus Jacot, 1930 . However, both species have different position of lyrifissures ia and im on notogaster. In addition, setae d of femora I situated anterior of level of setae v´ and v˝, setae l´ of genua IV absent (versus situated at middle of article posterior of setae v´ and v˝, setae l´ of genua IV present in P. commutabilis ).

comosus Austrophthiracarus (Aoki, 1980)

Neophthiracarus comosus Aoki, 1980

Phthiracarus comosus: Niedbała 1986b, 1992 , 2000.

Phthiracarus (Neophthiracarus) comosus: Subías 2022 .

Austrophthiracarus comosus: Niedbała 2011b, 2012b , Liu & Zhang 2016 a, Niedbała & Liu 2018. Palaearctic: Japan ( Aoki 1980a, Niedbała 1986b, 1992, 2000, 2011b, 2012b, Subías 2022).

Eastern Palaearctic, Japan, endemic compacta Indotritia (Afrotritia) Mahunka, 1988

Indotritia (Afrotritia) compacta: Niedbała 1998 a, 2001a , Liu 2015c, Niedbała & Liu 2018, Subías 2022. Afrotropic: Cameroon, Democratic Republic Congo, Tanzania, Malawi ( Balogh & Balogh 2002, Mahunka 1988a, Niedbała 1998 a, 2001a, 2017c, Niedbała & Starý 2014 d, 2022, Subías 2022).

Afrotropical, central part comparativus Notophthiracarus Niedbała et Colloff, 1997

Notophthiracarus comparativus: Niedbała 2000, 2006 d , Liu & Zhang 2014d, Niedbała & SzywilewskaSzczykutowicz 2017, Niedbała & Liu 2018, Niedbała et al. 2021.

Notophthiracarus (Notophthiracarus) comparativus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) comparativus: Subías 2022 .

Australasia: Australia (Victoria, Tasmania) ( Balogh & Balogh 2002, Niedbała 2000, 2006d, 2012a, Niedbała & Colloff 1997, Niedbała et al. 2021, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, southern part of Australia completa Indotritia Mahunka, 1987 synonym of Indotritia (Indotritia) javensis ( Sellnick, 1923)

complicatus Steganacarus (Steganacarus) Mahunka, 1993

Steganacarus complicatus Mahunka, 1993

Steganacarus complicatus: Subías 2004 .

Steganacarus (Steganacarus) complicatus: Niedbała 2001 a, Niedbała & Liu 2018 .

Afrotropic: Rwanda ( Balogh & Balogh 2002, Mahunka 1993b, Niedbała 2001a).

Note. Subías (2004) recognized this species as congeneric with S. (S.) vestitus Niedbała, 1983 , but both species differ by some characters: rostral setae procumbent in S. (S.) vestitus (versus semierect in S. (S.) complicatus ), setae d 1, d 2, e 2, h 3, ps 3 considerably shorter (ca 2.5 times) (versus less than 2 times shorter), and setae h 1, ps 1, ps 2 flagelliform (versus filiform).

Afrotropical, Rwanda, endemic compressus Phthiracarus Jacot, 1930

Phthiracarus compressum Jacot, 1930

Phthiracarus compressus: Niedbała 1986b, 1992 , 2000, 2002a, 2008f, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Weigmann 2006, Niedbała & Liu 2018, Niedbała & Błoszyk 2022.

non Phthiracarus pallidus: Niedbała 1986b, 1992 .

Phthiracarus (Phthiracarus) compressus: Subías 2022 .

Nearctic: USA (Connecticut, Maine, New York, California, Texas, Wisconsis, Iowa, Vermont, New Hampshire, Mississippi, North Carolina). Orient: Banglades. Palaearctic: Gibraltar, France, Norway, Sweden, Denmark, Germany, Luxembourg, Switzerland, Austria, Poland, Czech Republic, Slovakia, Romania, Ukraine (western part and Crimea), Georgia, Russia (North and Central European Territories, Eastern Siberia), India (Kashmir), South Korea ( Beck et al. 2014, Jacot 1930, 1938, Kaczmarek & Niedbała 2002, Mahunka & Mahunka-Papp 2003, Mahunka et al. 2013, Marshall et al. 1987, Niedbała 1986b, 1992, 2000, 2002a, 2008f, 2011b, 2012b, 2015b, Niedbała & Błoszyk 2022, Niedbała et al. 2020, Niedbała & Starý 2015e, Olszanowski et al. 1996, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Holarctic, Panpalaearctic, introduced to Bangladesh (?)

Notes. Subías (2022) considered Phthiracarus commutabilis Niedbała, 1983 as congeneric with this species. See note of P. commutabilis .

Subías (2022) considered Phthiracarus pallidus Feider et Suciu, 1958 (= P. occultus Niedbała, 1981 ) as congeneric with this species. See note of P. pallidus .

comteae Rhysotritia Mahunka, 1983 synonym of Acrotritia vestita ( Berlese, 1913)

comteae Euphthiracarus (Euphthiracarus) Mahunka, 1982

Euphthiracarus comteae Mahunka, 1982

Euphthiracarus comteae: Niedbała 2003 a, Niedbała & Liu 2018 .

Euphthiracarus (Euphthiracarus) comteae: Niedbała 2004a, 2008e , Subías 2022.

Neotropic: Costa Rica, Panama ( Balogh & Balogh 1987, 1988, 2002, Mahunka 1982e, Niedbała 2003a, 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica concavus Plonaphacarus Liu, Wu et Chen, 2011

Plonaphacarus concavus: Niedbała & Liu 2018 , Liu 2021b.

Hoplophthiracarus (Hoplophthiracarus) concavus: Subías 2022 .

Orient: China (Hubei) (Liu 2021b, Liu et al. 2011b, 2013, Subías 2022).

Oriental, China, endemic concinuus Hoplophthiracarus Niedbała, 1982

Hoplophthiracarus concinuus: Niedbała 1986b, 1992 , 2000, 2011b, 2012b, Subías 2004, Niedbała & Ermilov 2018 a, Niedbała & Liu 2018, Liu 2021b.

Hoplophthiracarus (Hoplophthiracarus) concinuus: Subías 2022 .

Hoplophthiracarus tropicus Mondal et Kundu, 1988 syn. by Subías 2004 (updated in 2014).

Hoplophthiracarus tropicus: Niedbała 2004c , Liu et al. 2011.

Orient: India (Bengal), China (Fujian). Palaearctic: Nepal, India (Arunachal Pradesh) ( Balogh & Balogh 2002, Liu 2021b, Liu et al. 2011, Mondal & Kundu 1988, Niedbała 1982a, 1986b, 1992, 2000, 2004c, 2011b, 2012b, Niedbała & Ermilov 2013b, 2018a, Subías 2022).

Oriental (Indian and Chinese), introduced to Tibetan region concolor Austrophthiracarus ( Sergienko, 2000)

Calyptophthiracarus concolor Sergienko, 2000

Notophthiracarus (Calyptophthiracarus) concolor: Subías 2004 .

Hoplophthiracarus (Hoplophthiracarus) concolor: Subías 2022 .

Austrophthiracarus concolor: Niedbała 2011b, 2012b , Liu & Zhang 2016 a, Niedbała & Liu 2018. Palaearctic: Ukraine (Crimea) ( Niedbała 2011b, 2012b, Sergienko 2000, Subías 2022).

Western Palaearctic, Crimea, endemic conformis Phthiracarus Sergienko, 1987 synonym of Phthiracarus incertus Niedbała, 1983

coniunctus Steganacarus (Steganacarus) Niedbała, 1983

Steganacarus (Steganacarus) coniunctus: Niedbała 1986b, 1992 , 1994a, 2011b, 2012b, Subías 2004, Niedbała & Liu 2018.

Steganacarus (Tropacarus) coniunctus: Subías 2022 .

Steganacarus danae Mahunka et Miko, 1989 syn. by Niedbała 1994a.

Steganacarus danae: Niedbała 1992 .

Palaearctic: Ukraine (western part), Georgia, Russia (South European Territory, Far East), Turkey ( Mahunka & Miko 1989, Niedbała 1983a, 1986b, 1992, 1994a, 2011b, 2012b, 2015a, Subías 2022).

Panpalaearctic, distributed in Pontic, Anatolian and Amurian areas, disjunctive consimilis Indotritia (Indotritia) Märkel, 1964

Indotritia krakatauensis consimilis Märkel, 1964

Indotritia krakatauensis consimilis: Balogh & Mahunka 1983 .

Indotritia consimilis: Balogh & Mahunka 1980a , Mahunka 1990b, 2001, Mahunka & Mahunka-Papp 2004, Niedbała 2011b, 2012b, Niedbała & Liu 2018.

Indotritia (Indotritia) consimilis: Liu 2015c , Subías 2022.

Indotritia krakatauensis consimilis Märkel, 1964

Palaearctic: Algeria, Croatia, Hungary ( Balogh & Mahunka 1980 a, 1983, Mahunka 1990b, 2001, Mahunka & Mahunka-Papp 1991, 2004, Mahunka et al. 2013, Märkel 1964, Niedbała 2011b, 2012b, Subías 2022). Western Palaearctic, reported from Central Europe and Holo-Mediterranean areas consimilis Notophthiracarus Niedbała et Colloff, 1997

Notophthiracarus consimilis: Niedbała 2000, 2006 d , Liu & Zhang 2014 d, Niedbała & Liu 2018, Niedbała et al. 2021.

Notophthiracarus (Notophthiracarus) consimilis: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) consimilis: Subías 2022 .

Australasia: Australia (Queensland, New South Wales, Victoria, Tasmania) ( Balogh & Balogh 2002, Niedbała 2000, 2006d, 2012a, Niedbała & Colloff 1997, Niedbała et al. 2021, Niedbała & Liu 2018, Subías 2022).

Australasian, disjunctive in Australia conspersus Notophthiracarus Niedbała, 2006

Notophthiracarus conspersus: Niedbała 2008e , Niedbała & Liu 2018.

Phthiracarus (Metaphthiracarus) conspersus: Subías 2022 .

Neotropic: Costa Rica, Panama ( Niedbała 2006e, 2008e, Subías 2022).

Neotropical, Mesoamerica conspicuus Notophthiracarus Niedbała, 1989

Notophthiracarus conspicuus: Niedbała 1992, 2000 , 2006 d, Liu & Zhang 2013 d, 2014d, 2015b, Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) conspicuus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) conspicuus: Subías 2022 .

Australasia: Australia (Tasmania), New Zealand ( Balogh & Balogh 2002, Niedbała 1989b, 1992, 2000, 2006d, 2012a, Niedbała & Ermilov 2016, Subías 2022).

Australasian, New Zealand and southern part of Australia contiguus Protophthiracarus Niedbała, 2004

Protophthiracarus contiguus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) contigua: Subías 2022 .

Neotropic: Antilles ( Cuba) ( Niedbała 2004a, Subías 2022).

Neotropical, Cuba, endemic contortula Oribotritia (Oribotritia) Niedbała, 1993

Oribotritia contortula Niedbała, 1993

Oribotritia contortula: Niedbała 1993b, 2000 , 2006d, Subías 2004, Liu & Zhang 2013a, Liu 2015a, 2016b, Niedbała & Liu 2018.

Oribotritia (Oribotritia) contortula: Subías 2022 .

Australasia: South Australia, New Zealand, New Caledonia ( Niedbała 1993b, 2000, 2006d, 2012a, Niedbała & Colloff 1997, Niedbała & Ermilov 2016, Subías 2004).

Australasian, eastern part contortulus Notophthiracarus Niedbała, 1991

Notophthiracarus contortulus: Niedbała 2006c , Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) contortulus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) contortulus: Subías 2022 .

South Africa: Mpumalanga, KwaZulu-Natal ( Balogh & Balogh 2002, Niedbała 1991b, 2006c, Subías 2022).

South African, indigenous contractilis Phthiracarus Perty, 1841 species inquirenda, incertae sedis

Phthiracarus contractilis: Oudemans 1937 , Parry 1979a, Niedbała 1986b, 1992, 20011b, Marshall et al. 1987, Subías 2004, Niedbała & Liu 2018.

Oribata dasypus Dugès, 1834

Phthiracarus dasypus: Castagnoli & Pegazzano 1985 .

Phthiracarus (Phthiracarus) dasypus: Subías 2004 .

Palaearctic: Germany (?).

Notes. Phthiracarus contractilis Perty, 1841 , figured by Claparède (1868), was synonymized by Jacot (1930) with Phthiracarus laevigatus (C.L. Koch, 1841) . It is not clear whether the figure of Claparède concerns Perty’s species. It is also not certain that the figure of Claparède concerns Phthiracarus laevigatus .

Subías (2022) considered this species as congeneric with Phthiracarus (P.) laevigatus (C.L. Koch, 1844) . However, there is no certainty that these species are congeneric.

contraria Microtritia Niedbała, 1993

Microtritia contraria: Niedbała 2000, 2006d , Niedbała & Penttinen 2007, Liu & Chen 2014 a, Liu & Zhang 2014 a, Niedbała & Szywilewska-Szczykutowicz 2017, Niedbała & Liu 2018, Niedbała et al. 2021, Subías 2022.

Australasia: Australia (broad, Tasmania), New Zealand ( Niedbała 1993b, 2000, 2006d, 2012a, 2017b, Niedbała & Colloff 1997, Niedbała et al. 2021, Niedbała & Penttinen 2007, Niedbała & SzywilewskaSzczykutowicz 2017, Subías 2022).

Australasian, broad contraria Oribotritia (Oribotritia) Niedbała, 1993

Oribotritia contraria Niedbała, 1993

Oribotritia sp. : Niedbała & Colloff 1997.

Oribotritia contraria: Niedbała 1993b, 2000 , 2006d, Subías 2004, Niedbała & Penttinen 2007, Liu & Zhang 2013a, Liu 2015a, 2016b, Niedbała & Liu 2018, Niedbała et al. 2021.

Oribotritia (Oribotritia) contraria: Subías 2022 .

Australasia: Australia (northern and southern parts), New Caledonia, New Zealand ( Liu & Zhang 2013a, Niedbała 1993b, 2000, 2006d, 2012a, 2017b, Niedbała & Colloff 1997, Niedbała et al. 2021, Niedbała & Penttinen 2007, Subías 2022).