Hoplophthiracarus nitidus, : Subias, 2022

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 153-158

Neotropical, Brazil, endemic nitidus Phthiracarus Niedbała, 1986

Phthiracarus nitidus: Niedbała 2002a, 2007c , Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) nitidus: Subías 2004 .

Phthiracarus (Phthiracarus) nitidus: Subías 2022 .

Nearctic: Canada (British Columbia, Nova Scotia , Quebec, Ontario), USA (Utah) ( Niedbała 1986b, 1992, 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022).

Nearctic, disjunctive notatus Arphthicarus Niedbała, 2001

Arphthicarus notatus: Niedbała & Liu 2018 .

Notophthiracarus (Notophthiracarus) notatus: Subías 2004 .

Hoplophorella (Hoplophorella) notata: Subías 2022 .

Afrotropic: Ghana, Gabon ( Niedbała 2001a, 2008 d, Subías 2022).

Afrotropical, central west Africa notoporosus Austrophthiracarus Liu et Zhang, 2014

Austrophthiracarus notoporosus: Liu & Zhang 2015a, 2016a , Liu & Wu 2016, Niedbała & Liu 2018. Hoplophthiracarus (Protophthiracarus) notoporosus: Subías 2022 .

Australasia: New Zealand ( Liu & Zhang 2014b, Subías 2022).

Australasian, New Zealand, endemic nova Mesotritia Starý, 1992

Mesotritia nova: Niedbała 2004a , Liu & Chen 2010a, Niedbała & Starý 2010b, Niedbała & Liu 2018. Mesotritia (Mesotritia) nova: Subías 2022 .

Neotropic: Antilles ( Cuba) ( Niedbała 2004c, Niedbała & Starý 2010b, Starý 1992a, Subías 2022). Neotropical, Cuba, endemic novazelandicus Notophthiracarus (Steganacarellus) Subías, 2010 synonym of Arphthicarus heterotrichus Niedbała, 2000

novazealandiensis Microtritia Niedbała, 2006

nov. nom for Microtritia glabrata Niedbała, 1993 : Niedbała 2006d

Microtritia glabrata: Niedbała & Colloff 1997 , Niedbała 2000, 2006d.

Microtritia novazealandiensis: Niedbała 2006 d, Liu & Chen 2014 a, Liu & Zhang 2014a , Liu 2016b, Niedbała & Szywilewska-Szczykutowicz 2017, Niedbała & Liu 2018, Niedbała et al. 2021.

Australasia: Papua New Guinea, New Zealand, Australia (Queensland, New South Wales, Victoria and Tasmania) (Liu 2016b, Niedbała 1993b, 2000, 2006d, 2012a, 2014b, 2017b, Niedbała & Colloff 1997, Niedbała et al. 2021, Niedbała & Starý 2010b, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, broad

Note. Subías (2022) considered this species as congeneric with M. glabrata Niedbała, 1993 . See explication at Niedbała (2006d).

novosibiricus Euphthiracarus (Euphthiracarus) Niedbała, 2006

Euphthiracarus novosibiricus Niedbała, 2006

Euphthiracarus novosibiricus: Niedbała 2011b, 2012b , Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) novosibiricus: Subías 2022 ).

Palaearctic: Russia (Western and Eastern Siberia, Far East) ( Niedbała 2006a, 2011b, 2012b, 2015b, Subías 2022).

Central-Eastern Palaearctic, distributed in Western Siberia and Amurian areas nuda Indotritia (Afrotritia) Mahunka, 1988

Indotritia nuda Mahunka, 1988

Indotritia nuda: Niedbała & Starý 2010b .

Indotritia (Indotritia) nuda: Niedbała 1998a , Liu 2015c, Subías 2022).

Indotritia (Afrotritia) nuda: Niedbała 2017c , Niedbała & Liu 2018.

Indotritia usambarensis Starý, 1993 syn. by Niedbała 1998a.

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1988a, Niedbała 1998a, 2017c, Niedbała & Starý 2010b, Starý 1993a, Subías 2022).

Afrotropical, Tanzania, endemic nuda Mesotritia (Berlese, 1887)

Tritia nuda Berlese, 1887

Oribotritia nuda: Hammen 1959 .

Tritia nuda: Berlese 1887a , Sellnick, 1923, Castagnoli & Pegazzano 1985.

Mesotritia nuda: Märkel 1964 , Aoki 1980a, Balogh & Mahunka 1980 a, 1983, Balogh & Mahunka 1983, Niedbała 1993 a, 2001b, 2002a, b, 2003a, 2004a, 2006c, 2007c, 2008e, f, 2010b, 2011b, 2012b, Mahunka 1994c, Mahunka & Mahunka-Papp 1995, 2004, Weigmann 2006, Liu & Chen 2010 a, Niedbała & Liu 2018, Liu 2021b, Niedbała et al. 2021.

Mesotritia cf. nuda: Ryabinin et al. 2018 .

Mesotritia (Mesotritia) nuda: Subías 2022 .

Oribotritia brachytrix Walker, 1965 syn. by Niedbała 2001b (lapsus linguae).

Oribotritia brachytrix (lapsus linguae): Niedbała 2002.

Mesotritia brachythrix: Marshall et al. 1987 .

Oribotritia grandjeani Feider et Suciu, 1957 syn. by Niedbała 2011b.

Mesotritia grandjeani: Balogh & Mahunka 1980, 1983 .

Mesotritia (Entomotritia) grandjeani: Märkel 1964 , Krivolutsky 1975.

Mesotritia (Mesotritia) grandjeani: Subías 2004 .

Mesotritia piffli Märkel, 1964 syn. by Niedbała 1993a.

Mesotritia piffli: Balogh & Mahunka 1980 a, 1983 , Mahunka & Mahunka-Papp 2004.

Mesotritia (Entomotritia) piffli: Märkel 1964 , Krivolutsky 1975.

Entomotritia piffli: Olszanowski et al. 1996 , Niedbała & Olszanowski 1997, 2004, 2008.

Mesotritia elastica Sergienko, 1988 syn. by Niedbała 1993a.

Nearctic: Canada (British Columbia, New Brunswick, Alberta, Quebec, Nova Scotia ), USA (Washington, Wyoming, Utah, Nevada, Arizona, New Mexico, South Dacota, Texas, Minnesota, Wisconsin, Iowa, Indiana, Vermont, Louisiana, Tennessee, South Carolina, California, Florida), Bermuda. Palaearctic:Azores, Canary Islands, Gibraltar, Sweden, Finland, Switzerland, Austria, Italy, Germany, Poland, Slovakia, Hungary, Romania, Belarus (Brest, Palm house), Ukraine (western and eastern parts and Crimea), Azerbaijan, Russia (South European Territory, Western Siberia, Far East), Iran, India (Kashmir), China (Gansu, Hebei, Beijing). Orient: China (Sichuan). Neotropic: Mexico, Costa Rica, El Salvador, Antilles, Ecuador. South Africa: Western Cape ( Ahaniazad et al 2016, Balogh & Mahunka 1980 a, 1983, Berlese 1887a, Castagnoli & Pegazzano 1985, Chen et al. 2013, Hammen 1959, Liu 2021b, Liu & Chen 2010a, Mahunka 1994c, Mahunka et al. 2013, Mahunka & Mahunka-Papp 1995, 2003, 2004, Marshall et al. 1987, Märkel 1964, Niedbała 1993 a, 2001b, 2002a, b, 2003a, 2004a, 2006c, 2007c, 2008e, f, 2010b, 2011b, 2012b, 2014c, 2015a, b, 2018, Niedbała & Buczkowska 2020, Niedbała & Roszkowska 2017, Niedbała & Starý 2010b, 2015e, Olszanowski et al. 1996, Ryabinin et al. 2018, Schatz 1983, Sergienko 1988, Subías 2022, Walker 1965, Weigmann et al. 2015).

Semicosmopolitan, Nearctic, Panpalaearctic, North-Western Oriental, Mesoamerica, South Africa but absent in Australasian Region nunomurai Indotritia (Indotritia) Hirauchi et Aoki, 2011

Indotritia nunomurai Hirauchi et Aoki, 2011

Indotritia nunomurai: Niedbała & Liu 2018 .

Indotritia (Indotritia) nunomurai: Liu 2015c , Subías 2022.

Palaearctic: Japan ( Hirauchi & Aoki 2011, Niedbała 2015b, Subías 2022).

Eastern Palaearctic, Japan, endemic obesus Atropacarus (Atropacarus) (Niedbała, 1983)

Steganacarus (Atropacarus) obesus Niedbała, 1983

Atropacarus obesus: Subías 2004 .

Atropacarus (Atropacarus) obesus: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Atropacarus (Atropacarus) obesus obesus: Subías 2022 .

Atropacarus obesus minimus Shtanchaeva et Subías, 2012 syn. by Niedbała 2015a.

Atropacarus obesus minimus: Subías 2022 .

Palaearctic: Ukraine (Crimea), Georgia, Russia (South European Territory), Abkhazia ( Niedbała 1983a, 1986b, 1992, 2011b, 2012b, 2015 a, Shtanchaeva & Subías 2012, Subías 2022).

Western Palaearctic, reported from Black Sea-Pontic region obesus minimus Atropacarus Shtanchaeva et Subías, 2012 = Atropacarus (Atropacarus) obesus (Niedbała, 1983) obliquus Notophthiracarus Niedbała et Starý, 2015

Notophthiracarus obliquus: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) obliquus: Subías 2022 .

Afrotropic: Madagascar (Niedbała 2017a, Niedbała & Starý 2015 d, Subías 2022.

Afrotropical, Madagascar, endemic oblongus Atropacarus (Hoplophorella) (Niedbała, 1983)

Hoplophorella oblonga Niedbała, 1983

Atropacarus (Hoplophorella) oblongus: Niedbała 1986b, 1992 , 2001 a, Niedbała & Liu 2018. Hoplophorella (Hoplophorella) oblonga: Subías 2022 .

Afrotropic: Kenya, Madagascar ( Balogh & Balogh 2002, Niedbała 1983i, 1986b, 1992, 2001a, 2017a, Niedbała & Starý 2016a, Subías 2022).

Afrotropical, central and eastern parts oblongus Euphthiracarus (Euphthiracarus) Liu, Wu et Chen, 2012

Euphthiracarus oblongus Liu, Wu et Chen, 2012

Euphthiracarus oblongus: Liu 2021b , Liu et al. 2012, Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) oblongus: Subías 2022 .

Orient: China (Hubei) (Liu 2021b, Liu et al. 2012, 2013, Subías 2022).

Oriental, China, endemic obscurus Phthiracarus Niedbała, 1986

Phthiracarus obscurus: Niedbała 1992, 2000 , 2004c, Niedbała & Corpuz-Raros 1998, Liu & Zhang 2016b, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) obscurus: Subías 2022 .

Palaearctic: India (Assam). Orient: Northern Vietnam, Indonesia (Java, Sumatra), Malaysia (Sabah), Philippines. Australasia: Australia (Queensland) ( Balogh & Balogh 2002, Niedbała 1986b, 1992, 2000, 2004c, 2012a, Niedbała & Corpus-Raros 1998, Niedbała & Ermilov 2013b, 2018b, Subías 2022). Oriental (eastern part) introduced to southern part of Palaearctic and to Australia (Queensland)

obsessus Notophthiracarus (Notophthiracarus) Subías, 2004 synonym of Notophthiracarus admirabilis Niedbała et Colloff, 1997

obsessus Phthiracarus (Neophthiracarus) Subías, 2004 synonym of Austrophthiracarus sellnicki ( Niedbała, 1987) obturatus Notophthiracarus Niedbała, 2009

Hoplophthiracarus (Steganacarellus) obturatus: Subías 2022 .

Neotropic: Antilles ( Cuba) ( Niedbała 2009a, Subías 2022).

Neotropical, Cuba, endemic obtusus Austrophthiracarus Niedbała, 2017

Austrophthiracarus obtusus: Niedbała & Roszkowska 2017 , Niedbała & Liu 2018.

Hoplophthiracarus (Protophthiracarus) obtusus: Subías 2022 .

Neotropic: Ecuador ( Niedbała & Roszkowska 2017, Subías 2022).

Neotropical, Ecuador, endemic occultus Phthiracarus Niedbała, 1981 synonym of Phthiracarus pallidus Feider et Suciu, 1958

ochraceus Atropacarus (Atropacarus) (Niedbała, 1983)

Steganacarus (Atropacarus) ochraceus Niedbała, 1983

non Steganacarus (Atropacarus) ochraseus (lapsus linguae): Sergienko 1985.

Atropacarus ochraceus: Subías 2004 .

Atropacarus (Atropacarus) ochraceus: Niedbała 1986b, 1992 , 2011b, 2012 b, Mortazavi et al. 2010, Niedbała & Liu 2018, Subías 2022.

Palaearctic: Macedonia, Ukraine (western part and Crimea), Georgia, Turkey, Syria, Iran ( Mortazavi et al. 2010, Niedbała 1983e, 1986b, 1992, 2011b, 2012b, 2015 b, Sergienko 1985, Subías 2022).

Western Palaearctic, distributed in Balkan-Black Sea, Anatolian, Pontic and Iranian areas ochthus Phthiracarus Niedbała, 2001

Phthiracarus ochthus: Niedbała 2004 d, Niedbała & Liu 2018 .

Phthiracarus (Archiphthtiracarus) ochthus: Subías 2022 .

Afrotropic: Ivory Coast, Cameroon, Tanzania, Madagascar ( Mahunka 2002, Niedbała 2001a, 2004d, 2017 a, Niedbała & Osiejuk 2018, Subías 2022).

Afrotropical, central and eastern parts, disjunctive octosetosus Phthiracarus Niedbała, 2004

Phthiracarus octosetosus: Niedbała & Liu 2018 .

Phthiracarus (Neophthiracarus) octosetosus: Subías 2022 .

Neotropic: Peru ( Niedbała 2004a, Subías 2022).

Neotropical, Peru, endemic oculus Protophthiracarus Niedbała, 2004

Protophthiracarus oculus: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) ocula: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic oenipontanus Austrophthiracarus (Mahunka, 1982)

Hauserophthiracarus oenipontanus Mahunka, 1982

Hauserophthiracarus oenipontanus: Balogh & Mahunka 1983 .

Calyptophthiracarus oenipontanus: Niedbała 1986b, 1992 .

Notophthiracarus (Calyptophthiracarus) oenipontanus: Subías 2004 .

Hoplophthiracarus (Calyptophthiracarus) oenipontanus: Subías 2022 .

Austrophthiracarus oenipontanus: Niedbała 1994a, 2011b , 2012b, Liu & Zhang 2016 a, Niedbała & Liu 2018.

Palaearctic: Greece, Israel ( Balogh & Mahunka 1983, Mahunka 1982a, Mahunka et al. 2013, Niedbała 1986b, 1992, 2011b, 2012b, Subías 2022).

Western Palaearctic, reported from boundary of Ponto-Mediterranean and Iranian regions oglasicola Protoplophora Bernini, 1983

Protoplophora oglasicola: Niedbała 2004b, 2011b , 2012b, Niedbała & Liu 2018, Subías 2022. Palaearctic: Italy ( Bernini 1983, Niedbała 2004b, 2011b, 2012b, Subías 2022).

Western Palaearctic, Italy, endemic ogmos Arphthicarus Niedbała, 2004

Arphthicarus ogmos: Liu & Oconnor 2014 , Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) venezolanus Subías, 2010 syn. by Niedbała & Liu 2018.

Neotropic: Venezuela ( Niedbała 2004a, Subías 2006, 2009).

Neotropical, Venezuela, endemic ogmos Notophthiracarus Niedbała, 2004

Notophthiracarus ogmos: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) ogmos: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic oidematos Protophthiracarus Niedbała, 2004

Protophthiracarus oidematos: Niedbała 2008e , Niedbała & Liu 2018.

Hoplophorella (Hoplophorella) oidematos: Subías 2022 .

Neotropic: Mexico, Belize, Panama ( Niedbała 2004a, 2008e, Subías 2022).

Neotropical, Mesoamerica okuyamai Mesotritia Aoki, 1980

Mesotritia okuyamai: Aoki 1980a , Niedbała 2000, 2011b, 2012b, Liu & Chen 2010 a, Niedbała & Liu 2018, Liu 2021b.

Mesotritia (Mesotritia) okuyamai: Subías 2022 .

Palaearctic: Japan. Orient: China (Fujian) ( Aoki 1980a, Liu 2021b, Liu & Chen 2010a, Niedbała 2000, 2011b, 2012b, Subías 2022).

Oriental (Chinese) introduced to Palaearctic Japanese subregion oligotricha Protoribotritia Märkel, 1963

Protoribotritia oligotricha: Krivolutsky 1975 , Balogh & Mahunka 1980 a, 1983, Niedbała 2002a, 2011b, 2012b, Subías 2004, Liu et al. 2011c, Niedbała & Liu 2018, Liu 2021b, Subías 2022.

Nearctic: Canada (Yukon, British Columbia, Alberta), USA (Wyoming, Utah, South Dakota, Maine, New Hampshire). Palaearctic: Norway, Sweden, Switzerland, Poland, Italy, Russia (North European Territory, Western and Eastern Siberia, Far East), North and South Korea, China (Tibet, Jilin, Qinghai), Nepal. Orient: China (southern part of Anhui) ( Balogh & Mahunka 1980 a, 1983, Forsslund & Märkel 1963, Liu 2021b, Liu et al. 2011c, Mahunka & Mahunka-Papp 2003, Märkel 1963, Niedbała 2002a, 2011b, 2012b, 2015b, Niedbała & Ermilov 2014 a, Niedbała et al. 2006, Ryabinin et al. 2018, Subías 2022).

Holarctic and Oriental. Panpalaearctic, disjunctive rather in northern and eastern parts olivaceus Austrophthiracarus ( Jacot, 1929)

Phthiracarus olivaceus Jacot, 1929

Phthiracarus olivaceum: Jacot 1930 .

Calyptophthiracarus olivaceus ( Jacot, 1929) : Niedbała 1986b, 1992.

Phthiracarus erinaceum Jacot, 1930 syn. by Niedbała 1986b.

Austrophthiracarus olivaceus: Niedbała 1994a, 2002a , Liu & Zhang 2016 a, Niedbała & Liu 2018.

Notophthiracarus (Calyptophthiracarus) olivaceus: Subías 2004 .

Hoplophthiracarus (Calyptophthiracarus) olivaceus: Subías 2022 .

Nearctic: Canada (Ontario, Quebec), USA (Connecticut, Oregon, Texas, Minnesota, Wisconsin, Iowa, Missouri, Illinois, Michigan, Indiana, Ohio, Kentucky, New Hampshire, Pennsylvania, New York, North Karolina, West Virginia) ( Jacot 1930, 1938, Marshall et al. 1987, Niedbała 1986b, 1992, 2002a, Subías 2022).

Nearctic, broad, continuous olszanowskii Arphthicarus Liu, 2020

Arphthicarus olszanowskii: Liu 2021b .

Hoplophthiracarus (Notophthiracarus) olszanowskii: Subías 2022 .

Palaearctic: China (Tibet) ( Liu 2020c, 2021b).

Palaearctic, China, endemic olszanowskii Plonaphacarus Niedbała, 2020

Hoplophorella (Rhacaplacarus) olszanowskii: Subías 2022 .

Afrotropic: Uganda (Niedbała 2020)

Afrotropical, Uganda, endemic onkos Atropacarus (Hoplophorella) Niedbała, 2017

Atropacarus (Hoplophorella) onkos: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) onkos: Subías 2022 .

Afrotropic: Tanzania (Niedbała 2017c, Subías 2022).

Afrotropical, Tanzania, endemic opacus Phthiracarus Niedbała, 1986

Phthiracarus opacus: Niedbała 1992, 2000 , 2008f, 2011b, 2012b, Olszanowski et al. 1996, Weigmann 2006, Beck et al. 2014, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) opacus: Subías 2004 .

Palaearctic: Finland, Spain, Norway, Denmark, Germany, Switzerland, Italy, Poland, Czech Republic, Slovenia, Croatia, Slovakia, Romania, Moldova Republic, Ukraine (western and eastern parts and Crimea), Georgia, Russia (North and South European Territory, Eastern Siberia), Turkey, North Korea ( Mahunka et al. 2013, Mahunka & Mahunka-Papp 2003, Niedbała 1986b, 1992, 2000, 2008f, 2011b, 2012b, 2015b, Niedbała et al. 2006, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Panpalaearctic

Note. Subías (2022) identified this species as congeneric with P. bryobius Jacot, 1930 , possibly agreeing with the opinion of Beck et al. (2014). However, these two species differ considerably by the length of notogastral setae: long, c 1> c 1 - d 1 or h 1>2 h 1 - ps 1 in P. opacus (versus these setae shorter than appropriate distances in P. bryobius ).