CEROCOCCIDAE Balachowsky 1942

Hodgson, Chris, 2020, A review of neococcid scale insects (Hemiptera: Sternorrhyncha: Coccomorpha) based on the morphology of the adult males, Zootaxa 4765 (1), pp. 1-264: 169-170

publication ID

http://doi.org/ 10.11646/zootaxa.4765.1.1

publication LSID

lsid:zoobank.org:pub:urn:lsid:zoobank.org:pub:C442D94C-0EB4-4509-B762-913707214819

persistent identifier

http://treatment.plazi.org/id/03B2EA64-0A87-46FB-2CFC-F9CCFA57D3B1

treatment provided by

Carolina

scientific name

CEROCOCCIDAE Balachowsky 1942
status

 

CEROCOCCIDAE Balachowsky 1942  

Cerococcus Comstock 1882, 213   . Type species: Cerococcus quercus Comstock   by monotypy.

Antecerococcus Comstock   ; Green 1901, 560. Green, 1908, 41. Incorrect synonymy ( Hodgson & Williams 2016, 19).

Introduction. The family Cerococcidae   currently includes 5 genera and 84 species ( García Morales et al. 2019). It was recently revised based on adult female morphology by Hodgson and Williams (2016). The only molecular study in which the Cerococcidae   have been included is that of Yokogawa and Yahara (2009) who found the cerococcids to be sister to the Asterolecaniidae   in a clade with the Kermesidae   . More recently, Hodgson and Hardy (2013), based on adult male morphology, found that the Cerococcidae   were sister to the Asterolecaniidae   and close to the Lecanodiaspididae   . No males of this family have been described in detail previously. The following diagnosis is based on the males of the three species described below.

Family diagnosis based on adult male morphology ( Figs 67–69 View FIGURE 67 View FIGURE 68 View FIGURE 69 ). Body: abdomen basically parallel-side and then narrowing to penial sheath; with very few body setae, all hs; loculate pores absent. Head: mid-cranial ridge present; ocelli absent; with only two pairs of simple eyes; ventral simple eyes placed distinctly posterior to dorsal simple eyes; gena with polygonal reticulations; postocular ridge touching dorsal eyes posteriorly; interocular ridge present (but ventral to dorsal eyes on A. indicus   and A. ornatus   but dorsad on C. artemisiae   ); ocular sclerite with polygonal reticulations but without inner microridges; postoccipital ridge absent or poorly developed; antennae short, about half total body length or rather less; antennae 10 segmented; antennal segment X with 4 capitate setae (unknown on A. ornatus   ); most antennal segments with both fs and hs setae. Thorax: prosternum with a welldeveloped lateral ridge but median ridge absent or ill-defined (unclear on C. indicus   ); prothorax without setae; membranous area of scutum with few or no setae; scutellum with a well-developed, inverted U-shaped, scutellar ridge; postmesospiracular setae absent; basisternum without a median ridge; basisternum without basisternal setae; lateropleurite broad, with an extension from marginal ridge; metasternum without metasternal setae; metepisternum not sclerotised, and metepimeron sclerotised; alar setae absent; alar lobe absent; hamulohalteres absent; legs quite setose, with ( Antecerococcus   ) or without ( Cerococcus   ) fs; trochanter elongate, with sensilla in a line; tibia with 1 or 2 spurs; tarsi 1 segmented; tarsal campaniform pores present; tarsal digitules capitate; claw digitules capitate; claws with a denticle (uncertain on C. ornatus   ). Abdomen: dorsal, ventral and pleural abdominal setae few and hs; tergites at most slightly sclerotised; glandular pouches present ( Antecerococcus   ) or absent ( Cerococcus   ); glandular pouches very shallow; caudal extensions on segment VII absent, those on segment VIII absent or small and rounded; penial sheath short, broad anteriorly, narrowing to a sharp point posteriorly; segment IX fused to style and not differentiated from it; basal membranous area at posterior end of segment IX present or absent; both surfaces of penial sheath with long setae; basal rod short; aedeagus blade-like.

Key to adult male Cerococcidae   (based on the three species studied here)

1. Glandular pouches and glandular pouch setae absent. Interocular ridge extending around dorsal side of each dorsal simple eye................................................................. Cerococcus artemisiae (Cockerell)   ( Fig. 67 View FIGURE 67 )

- Glandular pouches and glandular pouch setae present. Interocular ridge extending around ventral side of each dorsal simple eye … Antecerococcus   sp.................................................................................. 2

2. Antennal setae shorter than width of antennal segment. Metapleural ridge poorly developed or absent. Abdominal sternites not sclerotised........................................................ Antecerococcus indicus (Maskell)   ( Fig. 68 View FIGURE 68 )

- Antennal setae longer than width of antennal segments. Metapleural ridge well developed, extending medially to metasternum. Abdominal sternites sclerotised.......................................... Antecerococcus ornatus (Green)   ( Fig. 69 View FIGURE 69 )

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Cerococcidae

Loc

CEROCOCCIDAE Balachowsky 1942

Hodgson, Chris 2020
2020
Loc

Cerococcus

Cerococcus Comstock 1882, 213
Loc

Antecerococcus

Green 1901, 560
Green, 1908, 41
Hodgson & Williams 2016, 19