Cadoderus weisei Yunakov, 2012

Yunakov, Nikolai, 2012, 3535, Zootaxa 3535, pp. 1-104 : 13-14

Cadoderus weisei Yunakov , sp. n.

(figs 64–75, 365, 368)

Diagnosis. Very similar to C. bellus , differs from it by funicular segments 1–3 without scales; epifrons wider, at the level of antennal articulation 1.8 x narrower than vertex; eyes larger [FW/ELD: 0.67–0.87 (0.8)]; metatibiae with large teeth on interior edge; 2nd ventrite 2 x as long as 3rd one; apex of median lobe narrowly rounded, not flattened.

Description.

Measurements. BL: 2.79–3.22 (2.97) mm, BW: 1.34–1.66 (1.44) mm, BH: 1.20–1.52 (1.35) mm.

Vestiture. Body densely covered only with overlapping, round scales, setose. Setae of head form rows along lateral edges of epifrons (recumbent or subrecumbent) and above the eyes (erect). Subocular row consists of 3 setae. Elytral intervals 1–7 with distinct row of setae. Anterior half of elytra with rather short suberect setae (half as long as interval width), posterior declivity with rather long, strongly erect, thick, acute or truncate spatulate setae (as long as interval width). Ventrites densely squamose. Antennal scape setose and densely squamose; scales round; setae long, slender, acute, and erect. Funicle setose, funicular segments 1–3 with elongate scales.Male metatibiae with small grooming brush; hairs short. Tarsi setose and squamose.

Coloration. Background scaling deep-brown; striped pattern formed by green and cupreous scales. Dark and pale scales both with slightly pronounced pearl shine. Head dorsal surface brown, with green stripes around eyes and antennal sockets; anterior portion of epifrons green. Pronotum with deep-brown background scaling and 3 elongate cupreous stripes. Elytra background scaling deep-brown. Intervals 1, 7–11 green. Ventral side of head and pronotum green, meso- and metanotum cupreous. Femora brown-cupreous, proximal portion and ventral surface with green scales. Tibiae external surface cupreous.

Head. Rostrum weakly elongate [RL/RW: 1.14–1.17 (1.16)], parallel-sided. Pterygia not extended beyond contour of rostrum. Lateral carinae broadly weakly convex. Epifrons very narrow, parallel-sided, at the level of antennal articulation 2 x narrower than vertex, distinctly sloping at sides, with convex median carina. Transverse sulcus deep, concealed by dense scaling at dorsal surface and at sides. Anterior portion of epifrons not steep, without median sulcus. Frons horizontal, glabrous, bare. Epistome convex, broad U-shaped, delimited by narrow carina. Eyes large, orbicular, sublateral, moderately convex, highest at the middle [FW/ELD: 0.67–0.87 (0.80)]. Vertex broad, flat. Frontal fovea concealed by scales.

Antennae. Scape extended behind anterior edge of pronotum, evenly curved and widened. 1st funicular segment as long as 2nd 3rd–7th oblong.

Thorax. Pronotum strongly transverse [PL/PW: 0.79–0.84 (0.81)], evenly slightly convex at sides, constricted at apex, widest at middle. Disc weakly convex longitudinally and transversally. Tergosternal suture complete. Metanepisternal suture obsolete posteriorly.

Elytra. Broadly-ovate [EL/EW: 1.26–1.34 (1.31)], anterior edge arcuate, vertical, narrowly rounded; disc strongly convex [EL/BH: 1.38–1.50 (1.44)]. Subscutellar callosity hidden by scales, with 2 tiny tubercles. Intervals flat. Anterior portion of 1st striae with regular row of tiny tubercles.

Legs. Protibiae and mesotibiae curved, not widened at the apex. Interior edge C-shaped without teeth in distal portion. Metatibiae with small teeth on interior edge. Mucro well developed, thorn-shaped, acute. Setal comb of protibiae consists of sparse 9–10 black setae not extending beyond external edge of tibiae. Bevel of metatibiae narrowly enclosed; its surface bare. Tarsi robust; tarsomere 2 elongate, 1.5 x as long as wide; tarsomere 3 with two wide lobes; tarsomere 5 of metatarsus extending beyond apical lobes of 3rd by length of the last one.

Abdomen. Posterior margin of 1st ventrite straight. 2nd ventrite 2 x as long as 3rd one, posterior margin of 2nd ventrite straight. 5th ventrite in male flat, in female weakly convex near the apex; apical edge in male broadly rounded, in female acute.

Male genitalia. Median lobe parallel-sided, tubular, in apical 1/3 evenly narrowed anteriorly; apex attenuate, narrowly rounded; 0.7 x as long as apodeme. Internal sac without spiculate fields, with large, long aggonoporium consisting of two stick-shaped symmetrical lobes connected proximally by transverse bridge and long ventral slender process. Ostium triangular, moderately sclerotized. Ligulae membranous. Parameres long, fused in base, basal piece of tegmen narrow, tegminal apodeme 0.5 x as long as apodemes of median lobe.

Female genitalia. Coxites heavily sclerotized dorso-ventrally curved, with heavily sclerotized dorsal baculi, evenly covered with pores, without sensilla. Styli well developed, stick-shaped, bearing 1 seta. Spermatheca moderately sclerotized, with short broad ramus, collum slender, as long as ramus. Corpus large, slightly swollen. cornus thick, extended beyond corpus. Tergite 8 subtrapezoid, with curved fringe of multiple, tenuous, short setae at the apex. Sternite 8 thick. Lamella sharply narrowed and acute, without setae, knife-shaped, heavily sclerotized; apodeme thick; caput small.

Distribution. Tanzania: West Usambara Mountains (Kwai) ( Figs 365, 368).

Bionomics. Unknown.

Material examined. Tanzania: Holotype, ♂, dissected ( MTD) “Kwai, Weise” ; Paratypes. 1♂, 1♀, ( MTD) labeled as holotype; 1♀ ( DEI) “Usambara”; “ Ellimenistes / bellus Fst. ; “coll. Pape ”; “ Cadoderus / bellus Fst. / Hustache det.”

Etymology. The name is dedicated to the collector Paul Weise, a son of famous German coleopterologist Julius Weise, in recognition of his contribution to the knowledge of beetles of the Usambara Mountains.