Pinodytes sanjacinto Peck & Cook, 2011

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 33-34

Pinodytes sanjacinto Peck & Cook , new species

( Figs. 22 , 184 , 209–215 )

Type material. Holotype male ( SBPC). UNITED STATES. California: Riverside Co., San Jacinto Mts., Lake Fulmor , ± 15 km NW Idyllwild, 26.III.1999, 1530m, oak forest litter, S. & J. Peck, 99–112 . Paratypes (2). UNITED STATES. California: same data as holotype, 1 ( SBPC); Riverside Co., San Jacinto Mts., Vista Grande , ± 20 km NW Idyllwild, 26.III.1999, 1450m, S. & J. Peck, 99–111, scrub oak litter, 1 ( SBPC) .

Material examined. We have examined 3 specimens.

Distribution. Specimens ( Fig. 184 ) are known only from Rivereside County, in the San Jacinto Mountains of southern California.

Diagnostic description. Total length 1.52–1.62 mm; greatest width 0.70–0.76 mm. Color reddish brown; elongate-oval in shape ( Fig. 22 ). Head. Finely, sparsely punctate; a pair of widely spaced larger punctures on front; shining, with transverse substriate microsculpture on vertex and laterally. Eyes absent. Antenna ( Fig. 209 ) with antennomeres 2 and 3 subequal in length; antennomere 5 larger than 4, longer than 6; antennomere 7 longer but not wider than 8; antennomeres 9 and 10 each with a single sensory vesicle indicated apically by a protruding flange.

Pronotum. Finely punctate, punctures separated by 2–3 diameters; shining; with weak transverse substriate microsculpture basally and laterally. Widest near base, sides curving to apex; apical margin weakly emarginate, basal margin nearly straight; apical angles broadly rounded, basal angles narrowly rounded. Elytra. Punctures larger than on pronotum; in longitudinal rows on basal one-half; shining; punctures joined by fine, curved striae giving an imbricate appearance at base. Joined elytra slightly wider than pronotum, widest in basal one-third, roundly narrowing to apex. Legs. Protibia ( Fig. 210 ) slender, straight, moderately widened from base to apex; spinose on outer margin and apically; finely spinose on apical two-thirds of inner margin. Mesotibia ( Fig. 211 ) slender, straight; strongly spinose on outer margin and apically; spinose on apical one-third of inner margin. Metatibia ( Fig. 212 ) elongate, slender, nearly straight; strongly spinose apically; smaller spines on apical one-half. Metafemur ( Fig. 212 ) slender. Male protarsomeres ( Fig. 210 ) little expanded, bearing elongate setae laterally and thin, broad, colorless phanerae ventrally. Mesotarsomeres without phanerae. Venter. Mesoventrite ( Fig. 215 ) carinate; longitudinal carina with tooth-like expansion near middle; not excavated behind transverse carina. Male genitalia. Median lobe of aedeagus ( Figs. 213, 214 ) broad, with narrow apex. Inverted internal sac ( Fig. 214 ) with elongate structures. Parameres ( Figs. 213, 214 ) elongate, with flattened apices; extending beyond apex of median lobe; each bearing two setae before apex. Spermatheca. Tubular, slender, curved.

Etymology. The name sanjacinto , a noun in apposition, refers to the occurrence of this species in the San

Jacinto Mountains of California .

Figures

FIGURE 184. Distributions of species in the Pinodytes pusio species group in the southwestern United States. P.marinensis, open squares in the Coastal Ranges northwest of San Francisco Bay. P. minutus, open circles in the Coast and Sierra Nevada Mountains of central California, on both sides of the Napa River Valley. P. monterey, closed circles in the coastal Santa Lucia Mountains. P. parvus, closed triangles of the Coastal Ranges of northwest California., with disjunct populations west of San Francisco Bay. P. sanjacinto, open triangles in the San Jacinto Mountains of southern California.

FIGURES 19–27. Pinodytes habitus photos. 19. P.minutus, total length 1.08 mm. 20. P. monterey, total length 1.52 mm. 21. P. parvus, total length 1.22 mm. 22. P. sanjacinto, total length 1.60 mm. 23. P. sequoia, total length 2.92 mm. 24. P.shasta, total

FIGURES 209–215. Pinodytes sanjacinto. 209. Male right antenna, dorsal. 210. Male right protibia and protarsus. 211. Male right mesotibia. 212. Male right metafemur and metatibia. 213. Aedeagus, lateral. 214. Aedeagus, dorsal. 215. Mesoventrite, lateral. Scale bar = 0.3 mm.