Pinodytes garibaldi Peck & Cook, 2011

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 45

Pinodytes garibaldi Peck & Cook , new species

( Figs. 37 , 310 , 335–342 )

Type material. Holotype: male ( OSAC). UNITED STATES. Oregon: Garibaldi , 2 mi. N., duff, rotting wood, Mar. 15, 1955, V. Roth . Paratypes (5). UNITED STATES. Oregon: with same data as holotype, 3 ( OSAC); with same data except ex moss, 2 ( OSAC) .

Material examined. We have examined 6 specimens.

Distribution. Specimens ( Fig. 310 ) are known only from Tillamook County, in northwestern Washington.

Diagnostic description. Total length 2.48–2.80 mm; greatest width 1.04–1.24 mm. Reddish brown; elongate in shape ( Fig. 37 ). Head. Finely punctate, punctures separated by 2–4 diameters; with faint reticulate and substriate microsculpture. Eyes absent. Antenna ( Fig. 335 ) with antennomere 3 slightly larger than 2; antennomere 5 larger than 4 and 6; antennomere 7 clearly larger than 8; antennomeres 9 and 10 lack visible sensory vesicles. Pronotum. Finely punctate, punctures separated by 2–5 diameters; a few large punctures subapically and subbasally; with reticulate microsculpture. Sides subparallel in basal one-half, narrowing apically; apical margin emarginate, basal margin straight; apical angles rounded, basal angles subrectangular. Elytra. Strial punctures moderately coarse; striae impressed in basal one-half; interstrial punctures smaller; punctures joined by fine transverse strioles forming a weak imbricate pattern. Joined elytra slightly wider than pronotum; sides subparallel in basal one-half, narrowing apically. Legs. Protibia ( Fig. 336 ) broad at apex in male, slightly narrower in female; two curved spines at apex of outer margin; apical one-half of inner margin with fine, dense spines. Mesotibia ( Fig. 337 ) evenly widened from base to apex; with strong spines on outer margin and apically; fine spines on apical one-half of inner margin. Metatibia ( Fig. 338 ) slender basally, widened in apical two-fifths; apical two-fifths spinose. Metafemur ( Fig. 338 ) slender. Male protarsomeres ( Fig. 336 ) weakly expanded; protarsomere 1 as long as 2–4 combined; bearing elongate setae laterally and two rows of thin, broad, colorless phanerae ventrally. Mesotarsomeres without phanerae.

Venter. Mesoventrite ( Fig. 342 ) carinate; longitudinal carina with a median tooth; depressed anterior to median tooth; excavation behind transverse carina. Male genitalia. Median lobe of aedeagus ( Figs. 339, 340 ) elongate, broad, dorsoventrally curved near middle; asymmetrically, sharply declivous before short, narrow apex. Inverted internal sac ( Fig. 340 ) with three curved spines. Parameres ( Figs. 339, 340 ) slender, about two-thirds length of median lobe; each paramere bearing one apical and one subapical seta. Spermatheca. Tubular ( Fig. 341 ), angulate near middle.

Etymology. The name garibaldi , a noun in apposition, refers to the type locality of this species.

Figures

FIGURE 310. Distributions of species in the Pinodytes cryptophagoides species group in northwestern United States. P.rothi, open circles in northwestern Oregon down Coast Mountains to northern coastal California. P. tibialis, closed circles in southwestern Oregon. P. angulatus, open triangles in northwestern Oregon. P. delnorte, closed squares in northwestern California. P. garibaldi, closed triangle in northwestern Oregon.

FIGURES 37–43. Pinodytes habitus photos. 37. P.garibaldi, total length 2.80 mm. 38. P. haidagwaii, total length 3.04 mm. 39. P. isabella, total length 2.18 mm. 40. P.orca, total length 4.20 mm. 41. P.setosus, total length 1.96 mm. 42. P. shoshone, total length 2.18 mm. 43. P. sinuatus, total length 2.28 mm.

FIGURES 335–342. Pinodytes garibaldi. 335. Male right antenna, dorsal. 336. Male right protibia and protarsus. 337. Male right mesotibia. 338. Male right metafemur and metatibia. 339. Aedeagus, lateral. 340. Aedeagus, dorsal. 341. Spermatheca. 342. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 341), 0.45 mm (Figs. 339, 340), 0.6 mm (Fig. 342), 0.7 mm (Figs. 335– 338).