Pinodytes, Horn, 1880

Peck, Stewart B. & Cook, Joyce, 2011, Systematics, distributions and bionomics of the Catopocerini (eyeless soil fungivore beetles) of North America (Coleoptera: Leiodidae: Catopocerinae) 3077, Zootaxa 3077 (1), pp. 1-118 : 15-16

Key to North American species of Pinodytes (males)

Males can be separated from females by the presence of phanerae (apically widened setae) ventrally on at least the protarsomeres.

1 Both protarsomeres and mesotarsomeres bearing phanerae ventrally ( Figs. 46 , 55 , 64 , 72 )............................ 2

– Only protarsomeres bearing phanerae ventrally.............................................................. 5

2 Median lobe of aedeagus short and broad ( Fig. 49 ); parameres broad ( Fig. 49 ); interior highlands region of the United States … newtoni group................................................................................. P. newtoni

– Median lobe of aedeagus elongate ( Figs. 59 , 67 , 75 ); parameres narrow ( Figs.59 , 67 , 75 ); occurs in western North America. 3

3 Larger (total length about 2 mm); longitudinal carina of mesoventrite expanded at middle ( Fig. 61 ); the state of Alaska … borealis group................................................................................ P. borealis

– Smaller (total length 1.5 mm or less); longitudinal carina of mesoventrite not expanded at middle ( Figs. 69 , 78 ); does not occur in Alaska ............................................................................................ 4

4 Median lobe of aedeagus in lateral view with depth at middle about equal to depth of paramere ( Fig. 66 ); longitudinal carina of mesoventrite with a small tooth ( Fig. 69 ); the states of Washington and Idaho … punctatus group............ P. punctatus

– Median lobe of aedeagus in lateral view with depth at middle about twice depth of paramere ( Fig. 74 ); longitudinal carina of mesoventrite lacks teeth ( Fig. 78 ); the state of Colorado … colorado group................................ P. colorado

5 Antennomeres 9 and 10 lack visible sensory vesicles; antennomere 7 clearly larger than 8 ( Fig. 257 ); parameres slender and distinctly shorter than median lobe of aedeagus ( Fig. 262 ); protibia strongly widened apically ( Fig. 258 ); mesoventrite with a distinct, deep excavation posterior to transverse carina ( Fig.282 )........................... cryptophagoides group...6

– Antennomeres 9 and 10 with visible sensory vesicles ( Fig. 79 ); antennomeres 7 and 8 subequal in size ( Fig. 79 ); parameres broad, extending nearly to or beyond apex of median lobe of aedeagus ( Fig. 84 ); protibia not strongly widened apically; mesoventrite without a distinct excavation posterior to transverse carina ( Fig. 86 ) pusio group...........................21

6 Mesoventrite with large patch of dense white setae on each side of longitudinal carina ( Fig.334 )....................... 7

– Mesoventrite without patches of white setae................................................................ 8

7 Median lobe of aedeagus in dorsal view slightly constricted before narrow apex ( Fig. 332 ).................... P. delnorte

– Median lobe of aedeagus in dorsal view broadly rounded apically ( Fig. 373 )................................ P. setosus

8 Apex of median lobe of aedeagus demarked by a ridge ( Figs. 271 , 340 )......................................... 9

– Apex of median lobe of aedeagus not demarked by a ridge................................................... 10

9 Apex of median lobe of aedeagus narrow ( Fig. 340 ) and strongly declivous ( Fig. 339 )...................... P. garibaldi

– Apex of median lobe of aedeagus broad ( Fig. 271 ) and not declivous ( Fig. 270 )........................ P. subterraneus

10 Mesotibia abruptly widened before apex ( Fig. 313 )......................................................... 11

– Mesotibia evenly widened from base to apex.............................................................. 13

11 Median lobe of aedeagus not sinuate in dorsal view; apex of median lobe narrow with a weak, rounded lobe ( Fig. 316 )................................................................................................... P. tibialis

– Median lobe of aedeagus sinuate in dorsal view ( Figs. 366 , 389 ); apex of median lobe broad ( Figs. 366 , 389 )............ 12

12 Total length more than 4.0 mm; longitudinal carina of mesoventrite irregularly serrate ( Fig. 367 ); apical one-third of mesotibia expanded ( Fig. 362 ).............................................................................. P. orca

– Total length less than 2.5 mm; longitudinal carina of mesoventrite with a single small tooth ( Fig. 391 ); mesotibia expanded before apex ( Fig. 386 ).......................................................................... P. sinuatus

13 Median lobe of aedeagus sharply dorsoventrally angulate before flattened apex ( Figs.323 , 279 ); inner margin of mesotibia concave ( Figs. 321 , 277 ).................................................................................. 14

– Median lobe of aedeagus not as above; inner margin of mesotibia convex or straight.............................. 16

14 Total length about 2 mm; internal sac of aedeagus with a bilobed sclerite ( Fig. 324 )....................... P. angulatus

– Total length greater than 2 mm; internal sac of aedeagus without a bilobed sclerite................................. 15

15 Internal sac of aedeagus with several large sclerotized structures ( Fig. 280 ); the state of Oregon ............. P. imbricatus

– Internal sac of aedeagus with several types of spines ( Fig. 348 ); the islands of Haida Gwaii , British Columbia P. haidagwaii

16 Median lobe of aedeagus sharply dorsoventrally angulate in lateral view ( Fig. 296 ).......................... P. capizzii

– Median lobe of aedeagus variously dorsoventrally curved.................................................... 17

17 Median lobe of aedeagus in dorsal view wider in apical one-fifth than at middle ( Figs. 356 , 381 ); inner margin of mesotibia straight ( Figs. 353 , 378 ); the state of Idaho ................................................................. 18

– Apex of median lobe of aedeagus in dorsal view narrower than middle of median lobe ( Figs. 262 , 288 ); inner margin of mesotibia convex ( Figs. 259 , 285 ); Pacific coast............................................................ 19

18 Total length less than 2.0 mm; median lobe of aedeagus most strongly curved in apical one-half ( Fig. 355 )....... P. isabella

– Total length more than 2.0 mm; median lobe of aedeagus most strongly curved in posterior one-half ( Fig. 380 ).. P. shoshone

19 Apex of median lobe of aedeagus in dorsal view doubly constricted before apex ( Fig. 288 ).................... P. newelli – Apex of median lobe of aedeagus not doubly constricted before apex............................................ 20

20 Apex of median lobe of aedeagus in dorsal view attenuately spinose ( Fig. 262 )....................... P. cryptophagoides

– Apex of median lobe of aedeagus broadly triangular ( Fig. 307 )............................................ P.rothi

21 Median lobe of aedeagus constricted before rounded, lobed apex ( Fig. 158 ); known only from the state of Idaho .... P.idaho

– Median lobe of aedeagus not as above; California or southern Oregon .......................................... 22

22 Outer margin of mesotibia with a distinctive elongate spine near middle that is about two times length of other spines on outer margin ( Figs. 90 , 147 )................................................................................. 23

– Outer margin of mesotibia without a distinctive elongate spine near middle....................................... 29

23 Inverted internal sac of median lobe of aedeagus with a series of dark spines extending beyond base ( Figs. 150 , 230 )...... 24

– Inverted internal sac of median lobe of aedeagus without a series of dark spines extending beyond base................ 27

24 Mesoventrite with longitudinal carina on different plane than transverse carina ( Figs. 152 , 232 )....................... 25

– Mesoventrite with longitudinal carina on about same plane as transverse carina................................... 26

25 Apex of median lobe of aedeagus broadly rounded ( Fig. 150 ); pronotum with reticulate microsculpture.... P. humboldtensis

– Apex of median lobe of aedeagus narrowly rounded ( Fig. 230 ); pronotum with transverse, substriate microsculpture P.shasta

26 Median lobe of aedeagus narrow at apex ( Fig. 246 ); sides of base of median lobe meet at dorsal midline ( Fig.246 ). P.tehama

– Median lobe of aedeagus not narrow at apex ( Fig. 101 ); sides of base of median lobe do not meet at dorsal midline ( Fig. 101 )........................................................................................... P. chandleri

27 Apex of median lobe of aedeagus strongly narrow ( Fig. 93 ); internal sac with one pair of large spines and numerous small spines ( Fig. 93 )................................................................................. P. ovatus

– Apex of median lobe moderately narrow ( Figs. 165 , 190 ); internal sac not as above................................ 28

28 Internal sac of aedeagus with two pairs of large spines ( Fig. 165 )................................... P. klamathensis

– Internal sac of aedeagus with an hour-glass shaped sclerite and, when inverted, a double series of small spines ( Fig. 190 )................................................................................................ P.minutus

29 Median lobe of aedeagus broad, asymmetrical at apex (Fig.); internal sac with a single large, strong spine and clusters of smaller spines ( Fig. 238 ).......................................................................... P. spinus

– Aedeagus not as above................................................................................ 30

30 Median lobe of aedeagus, in dorsal view, with lateral notches before apex ( Figs. 84 , 142 , 254 )........................ 31

– Median lobe of aedeagus, in dorsal view, without lateral notches before apex..................................... 35

31 Inner margin of parameres, in dorsal view, with a flange before apex ( Figs. 142 , 254 ).............................. 32

– Inner margin of parameres straight...................................................................... 33

32 Median lobe of aedeagus, in lateral view, with dorsal margin sharply declivous at apical one-third ( Fig. 141 ); apex of median lobe not narrow ( Fig. 142 )....................................................................... P. gibbosus

– Median lobe of aedeagus, in lateral view, not sharply declivous ( Fig. 253 ); apex of median lobe narrow ( Fig. 254 ).... P.tuolumne

33 Larger species, total length 2–3 mm; longitudinal carina of mesoventrite irregularly serrate ( Fig. 223 )............ P. sequoia

– Smaller species, total length less than 1.7 mm; longitudinal carina of mesoventrite with a single tooth ( Figs. 86 , 135 )..... 34

34 Apex of median lobe of aedeagus flat ( Fig. 132 ), with acute tip ( Fig. 133 )................................... P. fresno

– Apex of median lobe of aedeagus not flat ( Fig. 83 ), with rounded tip ( Fig. 84 )................................ P.pusio

35 Width of metafemur 3–4 times width of metatibia.......................................................... 36

– Width of metafemur less than three times width of metatibia.................................................. 37

36 Metatibia with elongate spine near middle of outer margin ( Fig. 196 ); apex of median lobe of aedeagus triangular in dorsal view ( Fig. 198 )............................................................................... P. monterey

– Metatibia without elongate spine near middle of outer margin; median lobe of aedeagus with narrow apex ( Fig. 173 ) … P. losangeles

37 Body dorsally elongate in shape, length/width more than 2.25................................................. 38

– Body dorsally oval in shape, length/width less than 2.00..................................................... 39

38 Metatibia with strong spine near middle of outer margin ( Fig. 107 ); median lobe of aedeagus sharply constricted before triangular apex ( Fig. 109 )........................................................................ P. constrictus

– Metatibia without strong spine near middle of outer margin; median lobe of aedeagus broad with narrow apex ( Fig. 214 ).............................................................................................. P. sanjacinto

39 Apices of parameres contorted ( Fig. 117 )......................................................... P. contortus

– Apices of parameres not contorted....................................................................... 40

40 Median lobe of aedeagus widest at apical one-third ( Fig. 125 ); apex of median lobe acute ( Fig. 125 )............ P.eldorado

– Median lobe of aedeagus widest at basal one-third ( Figs. 181 , 206 ); apex of median lobe narrowly rounded ( Fig. 181 , 206 ). 41

41 Apex of median lobe of aedeagus ogival in shape ( Fig. 181 ); inverted internal sac with one pair of elongate sclerites ( Fig. 181 ).......................................................................................... P.marinensis

– Apex of median lobe of aedeagus rounded ( Fig. 206 ); inverted internal sac with two pairs of elongate sclerites ( Fig. 206 )................................................................................................. P.parvus

Figures

FIGURES 44–51. Pinodytes newtoni. 44. Male right antenna, dorsal. 45. Male right protibia and protarsus. 46. Male right mesotibia and mesotarsus. 47. Male right metafemur and metatibia. 48. Aedeagus, lateral. 49. Aedeagus, dorsal. 50. Spermatheca. 51. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 50), 0.2 mm (Fig. 49), 0.3 mm (Figs. 45–48, 51), 0.4 mm (Fig. 44).

FIGURES 53–61. Pinodytes borealis. 53. Male right antenna, dorsal. 54. Male right protibia and protarsus. 55. Male right mesotibia and mesotarsus. 56. Male right metafemur and metatibia. 57. Aedeagus, lateral. 58. Everted internal sac, lateral. 59. Aedeagus, dorsal. 60. Spermatheca. 61. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 60), 0.25 mm (Fig. 61), 0.3 mm (Figs. 57–59), 0.4 mm (Figs. 53–55), 0.5 mm (Fig. 56).

FIGURES 62–69. Pinodytes punctatus. 62. Male right antenna, dorsal. 63. Male right protibia and protarsus. 64. Male right mesotibia and mesotarsus. 65. Male right metafemur and metatibia. 66. Aedeagus, lateral. 67. Aedeagus, dorsal. 68. Spermatheca. 69. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 68), 0.3 mm (Figs. 62–67, 69).

FIGURES 70–78. Pinodytes colorado. 70. Male right antenna, dorsal. 71. Male right protibia and protarsus. 72. Male right mesotibia and mesotarsus. 73. Male right metafemur and metatibia. 74. Aedeagus, lateral. 75. Aedeagus, dorsal. 76. Everted internal sac. 77. Spermatheca. 78. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 77), 0.2 mm (Figs. 74, 75, 78), 0.3 mm (Figs. 70–73), 0.4 mm (Fig. 76).

FIGURES 257–264. Pinodytes cryptophagoides. 257. Male right antenna, dorsal. 258. Male right protibia and protarsus. 259. Male right mesotibia. 260. Male right metafemur and metatibia. 261. Aedeagus, lateral. 262. Aedeagus, dorsal. 263. Spermatheca. 264. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 263), 0.3 mm (Figs. 262, 264), 0.4 mm (Figs. 257–261).

FIGURES 275–282. Pinodytes imbricatus. 275. Male right antenna, dorsal. 276. Male right protibia and protarsus. 277. Male right mesotibia. 278. Male right metafemur and metatibia. 279. Aedeagus, lateral. 280. Aedeagus, dorsal. 281. Spermatheca. 282. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 281), 0.6 mm (Figs. 279, 280, 282), 0.7 mm (Figs. 275–278).

FIGURES 79–86. Pinodytes pusio. 79. Male right antenna, dorsal. 80. Male right protibia and protarsus. 81. Male right mesotibia. 82. Male right metafemur and metatibia. 83. Aedeagus, lateral. 84. Aedeagus, dorsal. 85. Spermatheca. 86. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 85), 0.3 mm (Figs. 79–84, 86).

FIGURES 327–334. Pinodytes delnorte. 327. Male right antenna, dorsal. 328. Male right protibia and protarsus. 329. Male right mesotibia. 330. Male right metafemur and metatibia. 331. Aedeagus, lateral. 332. Aedeagus, dorsal. 333. Spermatheca. 334. Mesoventrite, lateral; dotted line indicates area of setal patch. Scale bar = 0.1 mm (Fig. 333), 0.3 mm (Figs. 331, 332, 334), 0.4 mm (Figs. 327–330).

FIGURES 368–375. Pinodytes setosus. 368. Male right antenna, dorsal. 369. Male right protibia and protarsus. 370. Male right mesotibia. 371. Male right metafemur and metatibia. 372. Aedeagus, lateral. 373. Aedeagus, dorsal. 374. Spermatheca. 375. Mesoventrite, lateral; dotted line indicates area of setal patch. Scale bar = 0.1 mm (Fig. 374), 0.3 mm (Figs. 372, 373, 375), 0.4 mm (Figs. 368–371).

FIGURES 266–273. Pinodytes subterraneus. 266. Male right antenna, dorsal. 267. Male right protibia and protarsus. 268. Male right mesotibia. 269. Male right metafemur and metatibia. 270. Aedeagus, lateral. 271. Aedeagus, dorsal. 272. Spermatheca. 273. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 272), 0.45 mm (Figs. 270, 271), 0.6 mm (Fig. 273), 0.7 mm (Figs. 266–269).

FIGURES 335–342. Pinodytes garibaldi. 335. Male right antenna, dorsal. 336. Male right protibia and protarsus. 337. Male right mesotibia. 338. Male right metafemur and metatibia. 339. Aedeagus, lateral. 340. Aedeagus, dorsal. 341. Spermatheca. 342. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 341), 0.45 mm (Figs. 339, 340), 0.6 mm (Fig. 342), 0.7 mm (Figs. 335– 338).

FIGURES 311–318. Pinodytes tibialis. 311. Male right antenna, dorsal. 312. Male right protibia and protarsus. 313. Male right mesotibia. 314. Male right metafemur and metatibia. 315. Aedeagus, lateral. 316. Aedeagus, dorsal. 317. Spermatheca. 318. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 317), 0.3 mm (Figs. 315, 316, 318), 0.4 mm (Figs. 311–314).

FIGURES 360–367. Pinodytes orca. 360. Male right antenna, dorsal. 361. Male right protibia and protarsus, dorsal. 362. Male right protarsus, ventral. 363. Male right mesotibia. 364. Male right metafemur and metatibia. 365. Aedeagus, lateral. 366. Aedeagus, dorsal. 367. Mesoventrite, lateral. Scale bar = 0.4 mm (Fig. 362), 0.5 mm (Fig. 367), 0.7 mm (Figs. 361, 363–366), 1.0 mm (Fig. 360).

FIGURES 384–391. Pinodytes sinuatus. 384. Male right antenna, dorsal. 385. Male right protibia and protarsus. 386. Male right mesotibia. 387. Male right metafemur and metatibia. 388. Aedeagus, lateral. 389. Aedeagus, dorsal. 390. Spermatheca. 391. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 390), 0.3 mm (Fig. 391), 0.4 mm (Figs. 384–389).

FIGURES 319–326. Pinodytes angulatus. 319. Male right antenna, dorsal. 320. Male right protibia and protarsus. 321. Male right mesotibia. 322. Male right metafemur and metatibia. 323. Aedeagus, lateral. 324. Aedeagus, dorsal. 325. Spermatheca. 326. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 325), 0.3 mm (Fig. 326), 0.4 mm (Figs. 319–324).

FIGURES 343–350. Pinodytes haidagwaii. 343. Male right antenna, dorsal. 344. Male right protibia and protarsus. 345. Male right mesotibia. 346. Male right metafemur and metatibia. 347. Aedeagus, lateral. 348. Aedeagus, dorsal. 349. Spermatheca. 350. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 349), 0.45 mm (Fig. 348), 0.5 mm (Figs. 343–347, 350).

FIGURES 292–301. Pinodytes capizzii. 292. Male right antenna, dorsal. 293. Male right protibia and protarsus. 294. Male right mesotibia. 295. Male right metafemur and metatibia. 296. Aedeagus, lateral. 297. Aedeagus, dorsal. 298. Variation in apex of aedeagus, dorsal (Lane Co., Oregon). 299. Variation in apex of aedeagus, dorsal (Columbia Co., Oregon). 300. Spermatheca. 301. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 300), 0.3 mm (Figs. 292–299, 301).

FIGURES 351–358. Pinodytes isabella. 351. Male right antenna, dorsal. 352. Male right protibia and protarsus. 353. Male right mesotibia. 354. Male right metafemur and metatibia. 355. Aedeagus, lateral. 356. Aedeagus, dorsal. 357. Spermatheca. 358. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 357), 0.3 mm (Figs. 355, 356, 358), 0.4 mm (Figs. 351–354).

FIGURES 376–383. Pinodytes shoshone. 376. Male right antenna, dorsal. 377. Male right protibia and protarsus. 378. Male right mesotibia. 379. Male right metafemur and metatibia. 380. Aedeagus, lateral. 381. Aedeagus, dorsal. 382. Spermatheca. 383. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 382), 0.3 mm (Figs. 381, 383), 0.4 mm (Figs. 377–380), 0.45 mm (Fig. 376).

FIGURES 283–291. Pinodytes newelli. 283. Male right antenna, dorsal. 284. Male right protibia and protarsus. 285. Male right mesotibia. 286. Male right metafemur and metatibia. 287. Aedeagus, lateral. 288. Aedeagus, dorsal. 289. Variation in apex of aedeagus, dorsal (Tillamook Co., Oregon). 290. Spermatheca. 291. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 290), 0.3 mm (Figs. 287–289, 291), 0.4 mm (Figs. 283–286).

FIGURES 302–309. Pinodytes rothi. 302. Male right antenna, dorsal. 303. Male right protibia and protarsus. 304. Male right mesotibia. 305. Male right metafemur and metatibia. 306. Aedeagus, lateral. 307. Aedeagus, dorsal. 308. Spermatheca. 309. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 308), 0.3 mm (Figs. 302–307, 309).

FIGURES 153–159. Pinodytes idaho. 153. Male right antenna, dorsal. 154. Male right protibia and protarsus. 155. Male right mesotibia. 156. Male right metafemur and metatibia. 157. Aedeagus, lateral. 158. Aedeagus, dorsal. 159, Mesoventrite, lateral. Scale bar = 0.3 mm.

FIGURES 88–95. Pinodytes ovatus. 88. Male right antenna, dorsal. 89. Male right protibia and protarsus. 90. Male right mesotibia. 91. Male right metafemur and metatibia. 92. Aedeagus, lateral. 93. Aedeagus, dorsal. 94. Spermatheca. 95. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 94), 0.3 mm (Figs. 88–93, 95).

FIGURES 145–152. Pinodytes humboldtensis. 145. Male right antenna, dorsal. 146. Male right protibia and protarsus. 147. Male right mesotibia. 148. Male right metafemur and metatibia. 149. Aedeagus, lateral. 150. Aedeagus, dorsal. 151. Spermatheca. 152. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 151), 0.3 mm (Figs. 145–150, 152).

FIGURES 225–232. Pinodytes shasta. 225. Male right antenna, dorsal. 226. Male right protibia and protarsus. 227. Male right mesotibia. 228. Male right metafemur and metatibia. 229. Aedeagus, lateral. 230. Aedeagus, dorsal. 231. Spermatheca. 232. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 231), 0.3 mm (Figs. 225–230, 232).

FIGURES 241–248. Pinodytes tehama. 241. Male right antenna, dorsal. 242. Male right protibia and protarsus. 243. Male right mesotibia. 244. Male right metafemur and metatibia. 245. Aedeagus, lateral. 246. Aedeagus, dorsal. 247. Spermatheca. 248. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 247), 0.3 mm (Figs. 241–246, 248).

FIGURES 96–103. Pinodytes chandleri. 96. Male right antenna, dorsal. 97. Male right protibia and protarsus. 98. Male right mesotibia. 99. Male right metafemur and metatibia. 100. Aedeagus, lateral. 101. Aedeagus, dorsal. 102. Spermatheca. 103. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 102), 0.3 mm (Figs. 96–101, 103).

FIGURES 160–167. Pinodytes klamathensis. 160. Male right antenna, dorsal. 161. Male right protibia and protarsus. 162. Male right mesotibia. 163. Male right metafemur and metatibia. 164. Aedeagus, lateral. 165. Aedeagus, dorsal. 166. Spermatheca. 167. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 166), 0.3 mm (Figs. 160–165, 167).

FIGURES 185–192. Pinodytes minutus. 185. Male right antenna, dorsal. 186. Male right protibia and protarsus. 187. Male right mesotibia. 188. Male right metafemur and metatibia. 189. Aedeagus, lateral. 190. Aedeagus, dorsal. 191. Spermatheca. 192. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 191), 0.3 mm (Figs. 185–190, 192).

FIGURES 233–240. Pinodytes spinus. 233. Male right antenna, dorsal. 234. Male right protibia and protarsus. 235. Male right mesotibia. 236. Male right metafemur and metatibia. 237. Aedeagus, lateral. 238. Aedeagus, dorsal. 239. Spermatheca. 240. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 239), 0.3 mm (Figs. 233–238, 240).

FIGURES 137–144. Pinodytes gibbosus. 137. Male right antenna, dorsal. 138. Male right protibia and protarsus. 139. Male right mesotibia. 140. Male right metafemur and metatibia. 141. Aedeagus, lateral. 142. Aedeagus, dorsal. 143. Spermatheca. 144. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 143), 0.4 mm (Figs. 137–142, 144).

FIGURES 249–256. Pinodytes tuolumne. 249. Male right antenna, dorsal. 250. Male right protibia and protarsus. 251. Male right mesotibia. 252. Male right metafemur and metatibia. 253. Aedeagus, lateral. 254. Aedeagus, dorsal. 255. Spermatheca. 256. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 255), 0.3 mm (Figs. 249, 253, 254, 256), 0.4 mm (Figs. 250–252).

FIGURES 216–223. Pinodytes sequoia. 216. Male right antenna, dorsal. 217. Male right protibia and protarsus. 218. Male right mesotibia. 219. Male right metafemur and metatibia. 220. Aedeagus, lateral. 221. Aedeagus, dorsal. 222. Spermatheca. 223. Mesoventrite, lateral. Scale bar = 0.2 mm (Fig. 222), 0.3 mm (Fig. 223), 0.5 mm (Figs. 216–221).

FIGURES 128–135. Pinodytes fresno. 128. Male right antenna, dorsal. 129. Male right protibia and protarsus. 130. Male right mesotibia. 131. Male right metafemur and metatibia. 132. Aedeagus, lateral. 133. Aedeagus, dorsal. 134. Spermatheca. 135. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 134), 0.3 mm (Figs. 128–133, 135).

FIGURES 193–200. Pinodytes monterey. 193. Male right antenna, dorsal. 194. Male right protibia and protarsus. 195. Male right mesotibia. 196. Male right metafemur and metatibia. 197. Aedeagus, lateral. 198. Aedeagus, dorsal. 199. Spermatheca. 200. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 199), 0.3 mm (Figs. 193–198, 200).

FIGURES 168–175. Pinodytes losangeles. 168. Male right antenna, dorsal. 169. Male right protibia and protarsus. 170. Male right mesotibia. 171. Male right metafemur and metatibia. 172. Aedeagus, lateral. 173. Aedeagus, dorsal. 174. Spermatheca. 175. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 174), 0.4 mm (Figs. 168–173, 175).

FIGURES 104–111. Pinodytes constrictus. 104. Male right antenna, dorsal. 105. Male right protibia and protarsus. 106. Male right mesotibia. 107. Male right metafemur and metatibia. 108. Aedeagus, lateral. 109. Aedeagus, dorsal. 110. Spermatheca. 111. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 110), 0.3 mm (Figs. 104–109, 111).

FIGURES 209–215. Pinodytes sanjacinto. 209. Male right antenna, dorsal. 210. Male right protibia and protarsus. 211. Male right mesotibia. 212. Male right metafemur and metatibia. 213. Aedeagus, lateral. 214. Aedeagus, dorsal. 215. Mesoventrite, lateral. Scale bar = 0.3 mm.

FIGURES 112–119. Pinodytes contortus. 112. Male right antenna, dorsal. 113. Male right protibia and protarsus. 114. Male right mesotibia. 115. Male right metafemur and metatibia. 116. Aedeagus, lateral. 117. Aedeagus, dorsal. 118. Spermatheca. 119. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 118), 0.3 mm (Figs. 112–117, 119).

FIGURES 120–127. Pinodytes eldorado. 120. Male right antenna, dorsal. 121. Male right protibia and protarsus. 122. Male right mesotibia. 123. Male right metafemur and metatibia. 124. Aedeagus, lateral. 125. Aedeagus, dorsal. 126. Spermatheca. 127. Mesoventrite, lateral. Scale bar = 0.1 mm (Fig. 126), 0.3 mm (Figs. 120–125, 127).

FIGURES 176–183. Pinodytes marinensis. 176. Male right antenna, dorsal. 177. Male right protibia and protarsus. 178. Male right mesotibia. 179. Male right metafemur and metatibia. 180. Aedeagus, lateral. 181. Aedeagus, dorsal. 182. Spermatheca. 183. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 182), 0.3 mm (Figs. 176–181, 183).

FIGURES 201–208. Pinodytes parvus. 201. Male right antenna, dorsal. 202. Male right protibia and protarsus. 203. Male right mesotibia. 204. Male right metafemur and metatibia. 205. Aedeagus, lateral. 206. Aedeagus, dorsal. 207. Spermatheca. 208. Mesoventrite, lateral. Scale bar = 0.15 mm (Fig. 207), 0.3 mm (Figs. 201–206, 208).