Nebrioporus stearinus suavis ( Sharp, 1882 )

Toledo, Mario, 2009, Revision in part of the genus Nebrioporus Régimbart, 1906, with emphasis on the N. laeviventris-group (Coleoptera: Dytiscidae), Zootaxa 2040, pp. 1-111 : 76-78

Nebrioporus stearinus suavis ( Sharp, 1882) stat.n.

Deronectes suavis Sharp (1882: 430) (orig. descr.); Marseul (1882: 72); Branden (1885: 49); Severin (1892: 473).

Deronectes (Potamodytes) suavis Sharp ; Zimmermann (1920: 129) (synonymy of variegatus Aubé ).

Deronectes (Potamonectes) suavis Sharp ; Zimmermann (1933: 176) (synonymy of variegatus Aubé ).

Potamonectes suavis (Sharp) ; J. Balfour-Browne (1944a: 350); Guignot (1958a: 67); Guéorguiev (1965a: 497; 1971: 15; 1981: 410; 1987: 89); Franciscolo (1979: 475); Angelini (1984: 108); Rico et al. (1990: 116).

Nebrioporus (s. str.) suavis (Sharp) ; Nilsson & Angus (1992: 287); Nilsson (2001: 173; 2003: 67); Shaverdo (2003: 36).

Deronectes variegatus (Aubé) ; Seidlitz (1887: 55) (misinterpretation); Csiki (1908: 477); Sahlberg (1913: 42).

Hydroporus (Deronectes) variegatus Aubé ; Apfelbeck (1904: 370); Scholz (1916: 170).

Deronectes (Potamodytes) variegatus (Aubé) ; Zimmermann (1919: 188; 1920: 129).

Deronectes (Potamonectes) variegatus (Aubé) ; Zimmermann (1933: 176).

Potamonectes variegatus (Aubé) ; J. Balfour-Browne (1944a: 350) (synonymy of suavis Sharp ); Zaitzev (1953: 215).

Nebrioporus variegatus (Aubé) ; Nilsson (2001: 173) (synonymy of suavis Sharp ).

Deronectes variegatus var. infuscatus Sahlberg (1913: 42) (orig. descr.); Biström (1987: 35).

Type locality. D. suavis : Greece > Mt. Parnassus; D. variegatus var. infuscatus : Turkey > Smyrne> Jamanal Dagh.

Type material. Deronectes suavis : Lectotype ♂ ( BMNH), here designated "type [ BMNH red bordered, circular label] \ Mt. Parnassus [ Sharp hw.] \ Sharp Coll. 1905-313 \ type 258 ♂ [ Sharp hw.] \ Graecia n.sp. [Sharp hw.] \ Deronectes suavis n. sp. [Sharp hw.]" plus BMNH circular, blue bordered " Lectotype " label, added by me and my red designation label . Paralectotype 1 ♀ ( BMNH) with BMNH circular, yellow bordered " Paratype " label, and same indications of lectotype, except for "Graecia [Sharp hw.]" instead of "Graecia n.sp. " plus BMNH circular, light-blue bordered " Paralectotype " label, added by me and my red designation label .

Deronectes variegatus var. infuscatus : Syntypes (DABUH) not seen.

Additional material examined. Albania: Voskopos , 28.VIII.1935, A. G. H. Alston., B. M. 1935-431 (1 ex. BMNH) .

Bosnia Herzegovina: "Savina [Sasina], Hercegnovi, YU, Bach, 10.IV.84" (9 exs. HF) .

Bulgaria: Madan , 5.VII.1962, leg. V. Gueorguiev, Brit. Mus. 1964-151 (4 exs. BMNH) ; Gozedeltschevsko, S. Osikovo, 14.V.1959, P. Beron (2 exs. MIZIT) ; S. Nadejden, Charman , 6.VII.1959, Rekicka [Rekichka], P. Beron (1 ex. MIZIT) .

France: " H. variegatus Aubé , France, 67-56, D. (Potamonectes) variegatus Aubé J.B. Balfour-Browne det." (1 ex. BMNH) .

Greece: " Greece, Merlin Coll. 96-275 [bearing hw. pink label with " xanthoporus Steph." the first and " variegatus Aubé " the second], Received with this name from Merlin C.O.W., D. (Potamonectes) variegatus Aubé J.B. Balfour-Browne det. (2 exs. BMNH); H. variegatus (Kraats) [sic!] Greece, D. (Potamonectes) variegatus Aubé J.B. Balfour-Browne det. (1 ex. BMNH); Eubea, between Procopion and Pilion, 12.IX.82, leg. A. Sette, Potamonectes suavis (Sharp) det. Schizzerotto (9 exs. MSNV); Kephallinia, Moczarski, Potamonectes variegatus Aubé det. Hlisnikovskij 1952 (1 ex. MIZIT); Kephallinia, A. Vinkler, ex coll. Fischer (2 exs. MNHN); "Graecia, coll. E. Friv., Balkán, coll. E. Friv." (2 exs. MNHN, HNHM); Rhodos, 24.VI.–4.VII.1958, Palm, ZML 1998 155 (5 exs. MZLU); Rhodos S., 300 m, W Nano, 19.VI.1933 (1 ex. MNHN); Peloponnese, rd. Simopoulon–Lampeia, 4 km fork for Olympia, stream 650 m, 4.VII.1989, leg. M. Toledo (6 exs. MTP); Attica, Reitter, Deronectes variegatus Aubé , coll. Reitter (1 ex. HNHM); Arfa [?Arfarà, Peloponnese], Kanallakion, Back, 25.III.85, (13 exs. HF); Peloponnese, Astros, 14.VI.86, leg. H. Hebauer (7 exs. HF); Thassos [Thàsos], 12.VII.76, Tümpel, (4 ex. HF); idem, Bach (4 exs. HF); Khalkidhiki, Parhtenon, 25.VII.76, Bach (8 exs. HF); Akroriali, Estl. Salonìki, Bach, 11.IV.84 (1 ex. HF); Echinos, Xànthii, Bach, 6.IV.85, (24 exs. HF); Xànthii, Bach, 6.IV.85, (6 exs. HF); Staurupolis, Xànthii, Bach, (3 exs. HF); Westl. Kavalla [Kávala], Bach, 10.IV.87, (1 ex. HF); Lesbos, 6 km W Plomari, 20 m, 500 m N Melinda, leg. M. Jäch (9) (1 ex. HF); Samos, (5) 2 km NE Kalithea, 26.V.79, leg. Malicky (4 exs. GWW); Thassos (Tas 25), Prinos, 780 m, 16.X.80, leg. Malicky (1 ex. GWW); Ikaria, 2 km E Karavostamon, 30.V.79, leg. Malicky (4 exs. GWW); Ikaria, W.v. Mileopon, 2.VI.79 leg., Malicky (2exs. GWW).

Montenegro: "Budva, 22.VII.65" (1 ex. HF) .

Macedonia: Stari Doiran [Stari Dojran], 5.VII.1965 , Makedonija, Pretner (2 exs. MIZIT) ; Yugoslavia, M. Berovo , 17.VI.1957, T. Petkovski leg., Potamonectes variegatus Aubé, V. Guéorguiev det. 1958 (2 exs. MIZIT) ; Pirin [perhaps Pirin mountains, in Bulgaria, close to the actually eastern border of Macedonia], Maced., 5.VI.29, Maran et Táborský lgt. (1 ex. MIZIT) ; Kajmakcalan [?], 12.VII.1937, leg. Dr. J. Fodor (1 ex. HNHM) ; Skoplje , IV–VII.1937, leg. Dr. J. Fodor (2 ex. HNHM) .

Turkey: Asia Minor, Biledjik [Bilecik], V. Bodemeyer, G. C. Champion Coll., B. M. 1927-409 (4 exs. BMNH) ; Asia Minor, Eski Chehir [Eskisehir Prov.], V. Bodemeyer, Sammlung P. Franck, ex coll. P. Franck, Mus. Zool. Polonicum Varszawa 12/45, Potamonectes suavis Sharp det. V. Guéorguiev 1970 (1 ex. MIZIT) ; " Asia Minor, Böttcher , Berlin C. 2, coll. G. Falkenström " (1 ex. MZLU) ; " Asia Mineure , Tokat " (2 exs. MNHN) ; Artvin, 22.VI–4.VII.09, Bausch. [?], W. Baschkow (1 ex. ZIN) ; Asia Minor, Dr. Lendl, Inönü, 906[sic!].VIII.26 (1 ex. MNHN) ; Asia Minor, Dr. Lendl, Karaköi [Karaköse] 906[sic!].VIII.25 (1 ex. MNHN) ; Bodemeyer, Klein–Asien , Umgeb. Konia [Konya] (1 ex. MNHN) ; Kastamonu, 30 Km N. Kastamonu, 25.V.1989, leg. S. Schödl (3 exs. NMW) ; Kastamonu, road. Kastamonu to Akkaya , 25 km Akkaya, 8.VII.1992, leg. M. Toledo (7 exs. MTP) ; Diranaz Pass, road. Tasköprü to Sinop, stream m 1140, 10.VII.1992, leg. M. Toledo (17 exs. MTP) ; Gomushane [Gümüshane], Kösedagi pass (29), 29.V.1989, leg. S. Schödl (12 exs. NMW) ; West. Marmaris, Bach 5.IV.87, leg. Fery (4 exs. HF) ; Manavgat, westl. Gündogmus, Bach 700 m, 4.IV.85, leg. Fery (18 exs. HF) ; Manavgat, 24 km östl. Kisilot, Bach , 8.IV.87, leg. Fery (8 exs. HF) ; idem, 15 km östl. Kisilot, (6 exs. HF) ; idem, 4 km östl. Kisilot, (4 exs. HF) ; Antalya, Bach 300 m, 3.IV.85, leg. Fery (5 exs. HF) ; Artvin, Veliköy , 1600 m, 5.VI.89, leg. Schödl (53) (1 ex. HF) ; Nördl. Toslak, Alanya, Bach 600 m, 4.IV.85, leg. Fery (21 exs. HF) ; N Antalya, Bach ca. 700 m, 4.IV.85, leg. Fery (7 exs. HF) ; Erzurum, road Tortum to Narman , 9.VI.89, leg. Schödl (72) (1 ex. HF) ; Erzurum, road Pazaryolu to Bayburt, 1.VI.89, leg. S. Schödl (37) (1 ex. HF) ; Ordu, s. Harcbeli P., 28.V.89, leg. Schödl (23) (2 exs. HF) ; Nevsehir, Ürgüp , 12.9.91, leg. G. Wewalka, Potamonectes suavis Sharp det. Wewalka 92 (2 exs. GWW) ; Içel Prov. (Mersin), Büyükeçeli ( Ovaçic ), 50-100 m, 14–18.VII.1998, leg. J. Bezdeck (2 ex. JHP) ; Antalya, Beldibi p. Kemer, Bey dagl. 1–15.VII.1998, leg. P. Bulirsch (2 exs. JHP) .

Description. This subspecies is not morphologically different from N. stearinus stearinus , except for the punctuation on the sternites, as described in the key. TL 4.7–6.0 mm; MW 2.4–2.9 mm. The within population variability is the same as for the nominal subspecies except that there are no known almost completely black examples, like some N. stearinus stearinus from Iran.

Distribution ( Fig. 49). An Eastern Mediterranean subpecies, known with certainty from Albania, Bosnia, Bulgaria, Croatia (Dalmatia), Greece and some Greek islands (except Crete), Macedonia, Montenegro, Serbia, western and northern Turkey (where, in some provinces, it is sympatric with N. s. stearinus ). Historical data exists for westerly areas beyond the range stated above (Baleares, Sardinia, Sicily, Spain) which, like the old specimen examined in BMNH labelled " France ", might be the result of misidentification and/or mislabelling.