Edessa Fabricius, 1803
|
publication ID |
https://doi.org/ 10.11646/zootaxa.5240.1.1 |
|
publication LSID |
lsid:zoobank.org:pub:2FE467C1-EAC2-4E90-B673-CCE2CCA93C1D |
|
DOI |
https://doi.org/10.5281/zenodo.7871399 |
|
persistent identifier |
https://treatment.plazi.org/id/03B487C0-FFCF-2518-FF47-C6A5FA9AFB84 |
|
treatment provided by |
Plazi |
|
scientific name |
Edessa Fabricius, 1803 |
| status |
|
Subgenus Edessa Fabricius, 1803 View in CoL View at ENA
Edessa View in CoL (part.); Amyot & Serville, 1843: 158; Stål, 1872: 49.
Edessa (Edessa) ; Kirkaldy, 1909: 154; Almeida et al., 2018: 256; Fernandes et al., 2018: 204; Nunes et al., 2020: 402; Fernandes & Silva, 2021: 630.
Type species: Edessa antilope (Fabricius, 1798) (see Silva et al. 2017 for historic notes).
Diagnose. Dorsal surface usually green to brown ( Figs. 24 A,E View FIGURE 24 ). Ventral surface yellow to reddish-brown ( Figs. 24 B,F View FIGURE 24 ), but always lighter than dorsal surface. Pronotum with anterolateral margin concave ( Figs. 24 C View FIGURE 24 , 25 C View FIGURE 25 ). Humeral angles conical ( Figs. 24 A View FIGURE 24 , 26 C View FIGURE 26 , 27 C View FIGURE 27 ); somewhat developed and projected, with different sizes ( Figs. 24 A,B,C View FIGURE 24 , 26 A,C View FIGURE 26 ); apex rounded or slightly acuminated (never acute), shiny, smooth or shagreen and with few punctures; concolorous with pronotum ( Fig. 24 E View FIGURE 24 ), yellow ( Figs. 27 A,C View FIGURE 27 ), reddish-brown ( Fig. 29 A View FIGURE 29 ) or black ( Figs. 24 A,C View FIGURE 24 , 26 A,C View FIGURE 26 ) color extended ventrally ( Figs. 24 D,F View FIGURE 24 , 25 B View FIGURE 25 , 26 B,D View FIGURE 26 , 27 D View FIGURE 27 , 28 F View FIGURE 28 ).
Description. Head: Triangular, wider than long ( Fig. 24 A,C,E View FIGURE 24 ). Mandibular plates sulcate, contiguous ahead of tylus and ventrally curved. Bucculae high, slightly setulose. Rostrum setulose, yellow, segment IV distally black ( Figs. 24 D View FIGURE 24 , 25 B View FIGURE 25 , 29 F View FIGURE 29 ); segments I<II>III<IV. Antennae setulose, usually lighter than body ( Figs. 24 A,B View FIGURE 24 , 27 A,C,E View FIGURE 27 , 28 E View FIGURE 28 ).
Thorax: Pronotum trapezoidal, wider than long, densely punctured ( Figs. 24 A View FIGURE 24 , 26 A View FIGURE 26 , 27 C View FIGURE 27 , 28 C View FIGURE 28 ). Scutellum triangular, densely punctured ( Figs. 24 A View FIGURE 24 , 25 A View FIGURE 25 , 26 A View FIGURE 26 , 27 C View FIGURE 27 , 28 C View FIGURE 28 , 29 C View FIGURE 29 ). Corium densely punctured, normally with at least one vein contrasting with corium color ( Figs. 25 A View FIGURE 25 , 26 A View FIGURE 26 , 27 C View FIGURE 27 , 28 C View FIGURE 28 , 29 A View FIGURE 29 ). Membrane of the hemelytron brown with metallic shine ( Fig. 24 C View FIGURE 24 ). Evaporatorium dull, rugose; peritrema ruga-like ( Figs. 25 F View FIGURE 25 , 26 A View FIGURE 26 , 27 D View FIGURE 27 , 28 D View FIGURE 28 , 29 B View FIGURE 29 ). Metasternal process longer than wide, bifurcated anteriorly; apex of each arm enlarged ( Figs. 24 F View FIGURE 24 , 25 B View FIGURE 25 , 26 D View FIGURE 26 , 27 D View FIGURE 27 , 28 F View FIGURE 28 , 29 B View FIGURE 29 ). Legs setulose ( Fig. 24 B View FIGURE 24 ).
Abdomen: Dorsal surface densely and uniformly punctured, small, deep, with concolorous punctures with its surface. Connexivum exposed, punctured, posterolateral angle slightly projected and dark; with ( Figs. 24 A,C,E View FIGURE 24 , 27 C View FIGURE 27 , 28 E View FIGURE 28 ) or without contrasting spots ( Figs. 26 A,C View FIGURE 26 , 29 A View FIGURE 29 ). Segment VII posteriorly projected and acute ( Figs. 24 A View FIGURE 24 , 25 A View FIGURE 25 , 26 C View FIGURE 26 , 29 C View FIGURE 29 ).
Male genitalia: Pygophore trapezoidal and dorsoposteriorly open ( Figs. 1 A View FIGURE 1 , 2 A View FIGURE 2 , 5 A View FIGURE 5 , 20 A View FIGURE 20 ). Superior process of the genital cup dark, texture different from genital cup ( Figs. 1 D,E View FIGURE 1 , 4 D View FIGURE 4 , 12 D View FIGURE 12 ). Parameres yellow usually with dark margins ( Figs. 1 D,E View FIGURE 1 , 3 D,E View FIGURE 3 , 4 D,E View FIGURE 4 , 12 D,E View FIGURE 12 , 17 D,E View FIGURE 17 ); narrow twisted and sulcate at base; apex expanded in different shapes. Proctiger subcylindrical; dorsal surface sulcate posteriorly; lateral face excavated and setose ( Figs. 1 D,C View FIGURE 1 , 3 D,E View FIGURE 3 , 5 D,E View FIGURE 5 , 7 D,E View FIGURE 7 , 15 D,E View FIGURE 15 ). Ventral rim excavated medially; expansions delimiting excavation somewhat swollen, slightly developed not reaching level of posterolateral angle ( Figs. 1 C View FIGURE 1 , 2 C View FIGURE 2 , 3 C View FIGURE 3 , 20 C View FIGURE 20 , 23 C View FIGURE 23 ). Posterolateral angles developed, narrow with apex rounded ( Figs. 1 C,E View FIGURE 1 , 2 C,E View FIGURE 2 , 3 C,E View FIGURE 3 , 11 C,E View FIGURE 11 , 17 C,E View FIGURE 17 ). Ventral surface punctured ( Figs. 1 C View FIGURE 1 , 4 C View FIGURE 4 , 8 C View FIGURE 8 , 18 C View FIGURE 18 , 20 C View FIGURE 20 , 23 C View FIGURE 23 ).
Female genitalia: Valvifer VIII convex, setulose, punctured, subrectangular. Valvifers IX trapezoidal; presence of a lateral constriction that results in a median keel. Laterotergite VIII distally spiniform. Laterotergite IX triangular; excavated at base. Segment X rectangular and sulcated ( Figs. 1 F View FIGURE 1 , 2 F View FIGURE 2 , 7 F View FIGURE 7 , 12 F View FIGURE 12 , 18 F View FIGURE 18 ).
Comments: The nominal subgenus is the largest of Edessa with 79 species ( Table 1 View TABLE 1 ) that can be separated in several groups of species. Among the species mentioned by Stål (1872) as part of the species group Edessa , only Edessa cervus (Fabricius, 1787) , E. antilope (Fabricius, 1798) , E. taurina Stål, 1872 , E. arabs (Linnaeus, 1758) and E. cylindricornis Stål, 1872 were considered here as part of the subgenus Edessa . Three species groups described to Edessa belong to the nominal subgenus: cervus ( Fernandes & van Doesburg 2000b) , metallica ( Fernandes & Campos 2011) and ovina ( Fernandes & Silva 2021) . The nominal subgenus can be separated from most genera of Edessinae and subgenera of Edessa by the shape and color of the humeral angle. A comprehensive diagnose separating them can be found in Almeida et al. (2018) and Nunes et al. (2020). Four genera— Anisoedessa Nunes & Fernandes, 2019 , Calcatedessa Silva & Fernandes, 2021, Graziaedessa Eger, 2021 , and Mediocampus Thomas, 1994 —were never included in any diagnostic discussion between the genera of Edessinae . The subgenus Edessa can be separated from these four genera by the shape of the humeral angle described above. In all these four genera the humeral angles are dentiform or rounded, slightly developed and flat.
E. sexdens group
( Figs. 1–32 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 View FIGURE 5 View FIGURE 6 View FIGURE 7 View FIGURE 8 View FIGURE 9 View FIGURE 10 View FIGURE 11 View FIGURE 12 View FIGURE 13 View FIGURE 14 View FIGURE 15 View FIGURE 16 View FIGURE 17 View FIGURE 18 View FIGURE 19 View FIGURE 20 View FIGURE 21 View FIGURE 22 View FIGURE 23 View FIGURE 24 View FIGURE 25 View FIGURE 26 View FIGURE 27 View FIGURE 28 View FIGURE 29 View FIGURE 30 View FIGURE 31 View FIGURE 32 )
Diagnosis. Specimens large (13.2–25.2 mm). Dorsal surface green to orange ( Figs. 24 C View FIGURE 24 , 27 E View FIGURE 27 ). Underside of body dark yellow to orange with transversal brown, green or black lines on thorax and abdomen ( Figs. 9 B View FIGURE 9 , 24 B,D View FIGURE 24 , 26 D View FIGURE 26 ). Humeral angles as long as wide, with apices black ( Fig. 26 E View FIGURE 26 ), brown ( Fig. 24 A View FIGURE 24 ) or concolorous with surface ( Fig. 29 E View FIGURE 29 ) and slightly curved backwards ( Fig. 24 A View FIGURE 24 ) or, more rarely, straight ( Fig. 24 E View FIGURE 24 ). Corium with all veins concolorous ( Fig. 24 C View FIGURE 24 ) or yellow ( Fig. 25 A View FIGURE 25 ). Connexival segments with ( Fig. 24 C View FIGURE 24 ) or without ( Fig. 26 A View FIGURE 26 ) spots, these spots may ( Fig. 24 D View FIGURE 24 ) or not ( Fig. 26 B View FIGURE 26 ) extend to ventral side; with or without small concavities. Legs reddish brown to brown ( Fig. 24 B,D View FIGURE 24 ). Posterolateral angles of pygophore projected, tumid, medially slightly excavated ( Figs. 2 A,B View FIGURE 2 , 11 A,B View FIGURE 11 ). Superior process of the genital cup laminar, rectangular, usually continuing ventrally in a carina ( Figs. 1 B,E View FIGURE 1 , 6 B View FIGURE 6 ); oblique to paramere ( Figs. 1 B,D,E View FIGURE 1 , 3 E View FIGURE 3 , 5 E View FIGURE 5 , 20 D View FIGURE 20 ). Parameres club-like with three lobes ( Figs. 1 D,E View FIGURE 1 , 2 D,E View FIGURE 2 , 4 D,E View FIGURE 4 , 6 D,E View FIGURE 6 , 11 D, E View FIGURE 11 , 13 D, E View FIGURE 13 ).
Description. Head: Punctured, punctures small, deep, sparsely distributed and concolorous with surface, brown or black. Mandibular plates deeply sulcate medially ( Figs. 24 A,C View FIGURE 24 , 25 A,C View FIGURE 25 , 26 C,E View FIGURE 26 ). Bucculae tall, receiving nearly all first rostral segment ( Figs. 24 B View FIGURE 24 , 25 D View FIGURE 25 , 26 F View FIGURE 26 , 27 F View FIGURE 27 , 29 F View FIGURE 29 ); margin rounded ( Figs. 24 B View FIGURE 24 , 25 B View FIGURE 25 , 26 D View FIGURE 26 ). Rostrum brown ( Fig. 24 B View FIGURE 24 ). Antennomers, II+III shorter than IV; IV subequal in length to V ( Fig. 24 D View FIGURE 24 ).
Thorax: Pronotum densely and uniformly punctured ( Figs. 24 A View FIGURE 24 , 25 C View FIGURE 25 , 26 A View FIGURE 26 , 27 C View FIGURE 27 , 28 C View FIGURE 28 , 29 A View FIGURE 29 , 30 A View FIGURE 30 ); punctures small, deep, and brown, black or concolorous with surface; cicatrices and apices of humeral angles not punctured. Anterolateral angle of pronotum with a small tooth concolorous with surface ( Figs. 24 A,B View FIGURE 24 , 26 A,B View FIGURE 26 , 28 E,F View FIGURE 28 , 29 C View FIGURE 29 ). Anterolateral margin grooved, sulci extending ventrally ( Figs. 24 F View FIGURE 24 , 25 E,F View FIGURE 25 , 26 A,B View FIGURE 26 , 29 E,F View FIGURE 29 ). Scutellum with punctures deep, brown, black or concolorous with surface and evenly distributed; apex acute ( Figs. 24 C View FIGURE 24 , 25 C View FIGURE 25 , 26 C View FIGURE 26 , 27 A View FIGURE 27 , 29 A View FIGURE 29 , 30 E View FIGURE 30 ). Corium with punctures deep, brown or concolorous with surface and uniformly distributed ( Figs. 24 A View FIGURE 24 , 25 A View FIGURE 25 , 27 E View FIGURE 27 , 28 C View FIGURE 28 ). Membrane brown ( Fig. 24 E View FIGURE 24 ). Ventral surface of thorax with brown or black stripes ( Figs. 24 B View FIGURE 24 , 25 B View FIGURE 25 ); propleuron, mesopleuron and metapleuron with brown, green, black or concolorous punctures. Evaporatorium rugulose, concolorous or with contrasting stripes with pleura, opaque ( Figs. 24 D View FIGURE 24 , 25 B View FIGURE 25 ). Peritreme reaching 3/4 of the distance from ostiole of scent gland to lateral margin of metapleuron ( Fig. 25 B View FIGURE 25 ). Metasternal process as long as wide, raised anteriorly; anterior apex of each arm truncated and laterally expanded ( Figs. 1 G View FIGURE 1 , 24 B,F View FIGURE 24 , 25 F View FIGURE 25 , 26 B,F View FIGURE 26 , 27 D View FIGURE 27 , 28 F View FIGURE 28 ); anterior face of each arm rounded, flat or excavated.
Abdomen: Dorsally brown with punctures small, deep and evenly distributed. Connexivum ( Figs. 25 C,E View FIGURE 25 ) densely and uniformly punctured only on concavities; or densely and uniformly punctured except the outer margin; punctuation black or concolorous with connexivum. Abdominal segment VII with posterolateral angles developed, with apices acute and black ( Fig. 24 A View FIGURE 24 ), brown ( Fig. 25 A View FIGURE 25 ), green ( Fig. 9 A View FIGURE 9 ) or concolorous with surface ( Fig. 26 A View FIGURE 26 ). Ventrally, abdomen medially raised sometimes with brown to black spots forming or not a complete band ( Fig. 26 B,D,F View FIGURE 26 ). Intersegmental areas of abdomen black ( Fig. 24 D View FIGURE 24 ) or brown ( Fig. 25 D View FIGURE 25 ). Pseudosutures black ( Fig. 24 B View FIGURE 24 ), brown ( Fig. 26 B View FIGURE 26 ) or concolorous with surface ( Fig. 24 C View FIGURE 24 ). Rounded shallow excavation near each spiracle. Spiracles elliptical ( Fig. 24 B View FIGURE 24 ).
Male genitalia: Pygophore trapezoidal ( Figs. 1 A View FIGURE 1 , 2 A View FIGURE 2 , 3 A View FIGURE 3 ). Dorsal rim sinuous, wrinkled, ending on each side near posterolateral angles ( Figs. 1 A,B View FIGURE 1 , 14 A,E View FIGURE 14 , 15 A,E View FIGURE 15 ). Posterolateral angles developed, inner part brown, strongly punctured ( Figs. 1 A,B View FIGURE 1 , 2 D,E View FIGURE 2 , 6 D,E View FIGURE 6 ); apices rounded ( Figs. 1 A,B View FIGURE 1 , 7 D View FIGURE 7 , 12 D View FIGURE 12 ). Proctiger laterally constricted with dense band of short setae along the edge ( Figs. 1 D,E View FIGURE 1 , 12 D,E View FIGURE 12 , 15 D,E View FIGURE 15 , 20 D,E View FIGURE 20 ) of the subtriangular ( Figs. 1 D,E View FIGURE 1 ) or subelliptical ( Figs. 2 D,E View FIGURE 2 ) posterior face. Ventral rim setulose (except Edessa (E.) bella ) with median excavation U- or V-shaped ( Figs. 1 C View FIGURE 1 , 7 C View FIGURE 7 , 11 C View FIGURE 11 , 12 C View FIGURE 12 ). Ventral surface ( Fig. 1 C View FIGURE 1 ) punctured; punctures large, shallow, sparse, dark ( Fig. 1 C View FIGURE 1 ) or concolorous with surface ( Fig. 2 C View FIGURE 2 ).
Female genitalia: Valvifers VIII setulose and punctured ( Fig. 1 F View FIGURE 1 ); punctures dark ( Fig. 1 F View FIGURE 1 ) or concolorous with surface ( Fig. 7 F View FIGURE 7 ); convex, medial part with a conspicuous protuberance ( Fig. 1 F View FIGURE 1 ); distal margin straight ( Fig. 1 F View FIGURE 1 ). Valvifers IX ( Fig. 1 F View FIGURE 1 ) trapezoidal, large, carinated (except Edessa (E.) miniata ; Edessa (E.) electa ( Fig. 5 F View FIGURE 5 ); Edessa (E.) nigricornis ; Edessa (E.) nigroangulata , Edessa (E.) schirmeri ). Laterotergites VIII uniformly punctured; punctures small, shallow, concolorous with surface; distal margin acutely projected posteriorly, apices black ( Figs. 1 F View FIGURE 1 , 6 F View FIGURE 6 , 8 F View FIGURE 8 ). Laterotergites IX excavated at the base; apices black surpassing sclerite uniting laterotergites VIII, in ventral view ( Figs. 1 F View FIGURE 1 , 5 F View FIGURE 5 ).
Comments: The species belonging to the E. sexdens group have a large morphological variety, but also groups of very similar species. Despite this, the group has a set of characteristics that are not found in other species of the subgenus Edessa : connexival segments with a pair of dark spots ( Fig. 24 A View FIGURE 24 ); superior process of the genital cup laminar dorsally and continued ventrally by a carina ( Fig. 1 B,E View FIGURE 1 ); and oblique to the parameres ( Fig. 1 B,D,E View FIGURE 1 ); parameres with three lobes ( Fig. 1 D,E View FIGURE 1 ) (Silva 2017). Other characteristics are also related to the group, such as: anterior arms of the metasternal process with excavated apex ( Fig. 1 G View FIGURE 1 ); ventral side of the abdomen raised medially ( Fig. 24 B View FIGURE 24 ); rounded shallow excavation near each spiracle ( Fig. 24 B View FIGURE 24 ); union of dorsal rim with posterolateral angles grooved ( Fig. 2 A,B View FIGURE 2 ). However, within the E. sexdens group there are species that do not have the entire set of characteristics mentioned above. A few species do not have the spots on connexival segments ( Fig. 26 A View FIGURE 26 ); and others do not have the ventral carina of the superior process of the genital cup ( Figs. 4 B,E View FIGURE 4 , 7 B View FIGURE 7 , 8 B View FIGURE 8 ).
Distribution ( Figs. 30 View FIGURE 30 , 31 View FIGURE 31 , 32 View FIGURE 32 ): MEXICO: Chihuahua, Jalisco, Veracruz, Morelos, Guerrero, Quintana Roo, Chiapas, Mexico City; JAMAICA; GUATEMALA: El Petén, Izabal; HONDURAS; NICARAGUA: Léon; COSTA RICA: Guanacaste, Alajuela, Heredia, Limón Puntarenas, San Jose; PANAMA: Bocas Del Toro, Colón, Panama, West Panama; COLOMBIA: Bogotá, Cauca; VENEZUELA: Zulia, Carabobo, Aragua, Caracas, Monagas, Bolívar, Amazonas; TRINIDAD AND TOBAGO; GUYANA; SURINAME: Marowijne, Paramaribo; FRENCH GUYANA: Cayenne; BRAZIL: Pará, Amazonas, Maranh„o, Mato Grosso, Acre, Rondônia, Góias; ECUADOR: Napo; PERU: Satipo; BOLIVIA: La Paz, Santa Cruz, Cochabamba, ARGENTINA: Formosa.
Identification key to the species of the E. sexdens group
1- Body predominantly orange to red with large black areas on pronotum, corium and ventral surface of abdomen ( Fig. 27 E,F View FIGURE 27 )...................................................................... Edessa (E.) miniata Westwood, 1837
- Body predominantly green to brown with dark marks usually present only on humeral angles, connexivum, and stripes on ventral surface of abdomen ( Figs. 24 C,D View FIGURE 24 ; 9 A,B View FIGURE 9 ; 29 E,F View FIGURE 29 )........................................................ 2
2- Ventral surface of the body with alternated yellow and green stripes; humeral angles narrow and laterally directed ( Fig. 9 A,B View FIGURE 9 )................................................................... Edessa (E.) imbecilla Breddin, 1903
- Ventral surface of the body yellow to orange usually with brown lines, sometimes without lines; humeral angles different from above ( Figs. 25 C,D View FIGURE 25 ; 29 A,B View FIGURE 29 )........................................................................... 3
3- Humeral angles long and sloping forward ( Figs. 25 A View FIGURE 25 ; 26 C,E View FIGURE 26 )................................................. 4
- Humeral angles usually short and slightly sloping backward or directed sideward ( Figs. 26 A View FIGURE 26 ; 28 E View FIGURE 28 )................... 8
4- Humeral angles straight, posterior margin slightly crenulated ( Fig. 25 A,B View FIGURE 25 ).......... Edessa (E.) cylindricornis Stål, 1872
- Humeral angles distally curved backward, posterior margin smooth ( Fig. 26 C,E View FIGURE 26 ).................................. 5
5- Humeral angles narrow, about 4 x longer than wide ( Fig. 26 C,D View FIGURE 26 ); ventral surface with contrastingly black wide stripes ( Fig. 26 D View FIGURE 26 )................................................................... Edessa (E.) helicoceras Breddin, 1905
- Humeral angles wide, about twice longer than wide; ventral surface usually with narrow brown to black stripes, if broad, stripes are brown ( Figs. 24 C,D View FIGURE 24 ; 26 E,F View FIGURE 26 ; 27 C,D View FIGURE 27 )................................................................. 6
6- Humeral angles dorsally mostly green to olive green with more than distal third black ( Fig. 26 E,F View FIGURE 26 )............................................................................................. Edessa (E.) helix Erichson, 1848
- Humeral angles dorsally mostly yellow or with apices black ( Figs. 24 C,D View FIGURE 24 ; 27 C,D View FIGURE 27 )................................ 7
7- Humeral angles yellow without black apices ( Fig. 27 C,D View FIGURE 27 ); connexival segments without black marks ( Fig. 27 C View FIGURE 27 ); ventral stripes on abdomen broad ( Fig. 27 D View FIGURE 27 )................................... Edessa (E.) lewisi Fernandes & Silva, 2015
- Humeral angles with apices black ( Fig. 24 C,D View FIGURE 24 ); connexival segments with black spots ( Fig. 24 C View FIGURE 24 ); ventral stripes on abdomen narrow ( Fig. 24 D View FIGURE 24 )................................................... Edessa (E.) bella Fernandes & Silva, 2015
8- Connexival segments without dark spots ( Figs. 26 A View FIGURE 26 ; 28 A View FIGURE 28 )................................................... 9
- Connexival segments with dark spots ( Figs. 24 A,E View FIGURE 24 ; 29 C,E View FIGURE 29 )................................................. 13
9- Humeral angle with black apices ( Figs. 26 A View FIGURE 26 ; 28 A View FIGURE 28 )......................................................... 10
- Humeral angles with yellow to light brown apices ( Figs. 25 E View FIGURE 25 ; 29 A View FIGURE 29 )........................................... 11
10- Humeral angles barely developed, black part longer than wide ( Fig. 26 A,B View FIGURE 26 ); evaporatorium darker than thorax ( Fig. 26 B View FIGURE 26 )......................................................................... Edessa (E.) gentilitia Distant, 1890
- Humeral angles little developed, black part as long as wide in dorsal view ( Fig. 28 A,B View FIGURE 28 ); evaporatorium concolorous with thorax ( Fig. 28 B View FIGURE 28 )........................................................... Edessa (E.) nigriclava Walker, 1868
11- Evaporatorium dark brown ( Fig. 29 B View FIGURE 29 ); ventral surface of abdomen with wide transversal and longitudinal dark brown to black stripes ( Fig. 29 B View FIGURE 29 )...................................................... Edessa (E.) phoenicopus Dallas, 1851
- Evaporatorium concolorous with thorax ( Figs. 25 F View FIGURE 25 ; 27 B View FIGURE 27 ); ventral surface of abdomen with narrow transversal black stripes ( Figs. 25 F View FIGURE 25 ; 27 B View FIGURE 27 ).................................................................................... 12
12- Paramere anterior process truncated ( Fig. 6 D,E View FIGURE 6 ); superior process of genital cup narrow, short and continuing ventrally in high crenulate carina ( Fig. 6 B View FIGURE 6 )....................................................... Edessa (E.) fallenii Stål, 1872
- Paramere anterior process developed and triangular ( Fig. 12 D,E View FIGURE 12 ); superior process of genital cup large, continuing ventrally in a short carina crenulate ( Fig. 12 B View FIGURE 12 )........................................... Edessa (E.) leucocera Dallas, 1851
13- Apices of humeral angles concolorous with dorsal surface, yellow or light brown ( Figs. 24 A,E View FIGURE 24 ; 29 C,E View FIGURE 29 )............... 14
- Apices of humeral angles black or blackish ( Figs. 25 C View FIGURE 25 ; 28 C,E View FIGURE 28 )............................................... 17
14- Humeral angles directed sideward ( Fig. 24 E,F View FIGURE 24 ); spots on connexivum small, rounded and not reaching any of the margins separating segment ( Fig. 24 E View FIGURE 24 )............................................... Edessa (E.) congrua Walker, 1868
- Humeral angles directed backwards ( Figs. 24 A View FIGURE 24 ; 29 C,E View FIGURE 29 ); spots on connexivum large, subrectangular and usually reaching at least one of the margins separating segments ( Figs. 24 A View FIGURE 24 ; 29 C,E View FIGURE 29 )............................................. 15
15- Apices of humeral angles not clearly globose, acuminated and short ( Fig. 29 C,E View FIGURE 29 )................................. 16
- Apices of humeral angles clearly globose ( Fig. 24 A,B View FIGURE 24 )............................. Edessa (E.) alces Erichson, 1848
16- Specimens large (about 20 mm) ( Fig. 29 E,F View FIGURE 29 ); venter uniformly dark yellow or orange ( Fig. 29 F View FIGURE 29 ); apices of anterior bifurcation of metasternal process rounded ( Fig. 23 G View FIGURE 23 )....................................... Edessa (E.) urus Erichson, 1848
- Specimens small (around 15 mm) ( Fig. 29 C,D View FIGURE 29 ); venter uniformly greenish yellow or ochraceous ( Fig. 29 D View FIGURE 29 ); apices of anterior bifurcation of metasternal process laterally expanded ( Fig. 20 G View FIGURE 20 )................... Edessa (E.) sexdens Fabricius, 1803
17- Body olive brown ( Figs. 28 C,D View FIGURE 28 ); humeral angles robust, globose, almost as long as wide at base ( Fig. 28 C,D View FIGURE 28 ); venter reddish brown ( Fig. 28 C View FIGURE 28 )......................................................... Edessa (E.) nigricornis Stål, 1862
- Body green to olive green ( Figs. 25 C View FIGURE 25 ; 28 E View FIGURE 28 ); humeral angles not as robust or globose as above, wider at base than long ( Figs. 25 C View FIGURE 25 ; 28 E View FIGURE 28 ); venter greenish yellow or orange ( Figs. 25 D View FIGURE 25 ; 28 F View FIGURE 28 ; 10 F,G View FIGURE 10 )........................................ 18
18- Black spots on connexival segments large ( Fig. 28 E View FIGURE 28 ); stripes on intersegmental areas solid black ( Fig. 28 F View FIGURE 28 )........... 19
- Black spots on connexival segments smaller than above ( Fig. 25 C View FIGURE 25 ); stripes on intersegmental areas faded brown ( Fig. 25 D View FIGURE 25 ) .................................................................................................. 21
19- Intersegmental stripes of abdomen complete ( Fig. 22 B View FIGURE 22 )......................... Edessa (E.) subandina Breddin, 1904
- Intersegmental stripes of abdomen incomplete ( Figs. 19 B View FIGURE 19 , 28 F View FIGURE 28 ).............................................. 20
20- Connexival spots large and expanded ventrally ( Fig. 28 E,F View FIGURE 28 ); pseudosutures faded brown ( Fig. 28 F View FIGURE 28 ); intersegmental stripes of abdomen narrow ( Fig. 28 F View FIGURE 28 )................................... Edessa (E.) nigroangulata Fernandes & Silva, 2015
- Connexival spots small and not expanded ventrally ( Fig. 19 A,B View FIGURE 19 ); pseudosutures solid black ( Fig. 19 B View FIGURE 19 ); intersegmental stripes of abdomen wide ( Fig. 19 B View FIGURE 19 )............................................... Edessa (E.) schirmeri Breddin, 1904
21- Connexivum greenish-yellow, dark spots triangular ( Fig. 25 C View FIGURE 25 ); apex of anterior bifurcation of metasternal process straight ( Fig. 5 G View FIGURE 5 ); propleuron with long dark stripe ( Fig. 25 D View FIGURE 25 )............................. Edessa (E.) electa Walker, 1868
- Connexivum orange, dark spots rectangular ( Fig. 10 F View FIGURE 10 ); apex of anterior bifurcation of metasternal process rounded ( Fig. 11 G View FIGURE 11 ); propleuron with short dark stripe ( Fig. 10 B,G View FIGURE 10 )................................... Edessa (E.) lavata Breddin, 1903
Species description
TABLE 1. Species belonging to subgenus Edessa. * Species group E. sexdens
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
|
Kingdom |
|
|
Phylum |
|
|
Class |
|
|
Order |
|
|
Family |
|
|
SubFamily |
Edessinae |
Edessa Fabricius, 1803
| Mendonça, Maria Thayane Da Silva, Silva, Valéria Juliete Da & Fernandes, José Antônio Marin 2023 |
Edessa (Edessa)
| Fernandes, J. A. M. & Silva, V. J. 2021: 630 |
| Nunes, B. M. & Campos, L. D. & Mendonca, M. T. S. & Cunha, E. V. P. & Fernandes, J. A. M. 2020: 402 |
| Almeida, F. R. A. & Nunes, B. M. & Fernandes, J. A. M. 2018: 256 |
| Fernandes, J. A. M. & Nascimento, A. T. S. & Nunes, B. M. 2018: 204 |
| Kirkaldy, G. W. 1909: 154 |
Edessa
| Stal, C. 1872: 49 |
| Amyot, C. J. B. & Serville, A. 1843: 158 |