Okanopteryx jeppesenorum Archibald & Cannings, 2021

Archibald, Bruce, Cannings, Robert A., Erickson, Robert J., Bybee, Seth M. & Mathewes, Rolf W., 2021, The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America, Zootaxa 4934 (1), pp. 1-133 : 79-85

Okanopteryx jeppesenorum Archibald & Cannings , new species

Figs. 44–48 .

Diagnosis. Wings distinct from those of O. macabeensis , O. fraseri by CuA–A space wider, four cells wide at widest (difficult to count number of columns by preservation), between the levels of origins of IR1, RP2 [ O. macabeensis : 5–10 columns of three cells wide; O. fraseri : all two cells wide, but sometimes a single column of three]; further from O. macabeensis , by longer wing, holotype, 22.4 mm nodus to apex; paratype 1 (SR 11-43-09): 23.5 mm [ O. macabeensis holotype: estimated 18.4 mm] (all O. fraseri wings missing apex).

Material. Holotype: SR 13-005-013 ( Fig. 44 ), collected at B4131 by Jackie and Tim Jeppesen, 15.vii.2013, housed in the Stonerose Interpretive Center collections; paratype 1: SR 11-43-09, ( Fig. 45 ), collected at Republic B 4131 by Christin Gustafson, 15.vi.2011, in the Stonerose Interpretive Center collections; paratype 2: SR 18-002- 001 ( Fig. 46 ), collected at Republic B 4131 by Valerie Paquin, 28.iv.2018, in the Stonerose Interpretive Center collections; paratype 3: SR 15-003-001 ( Fig. 47 ), collected at Republic B 4131 by Ethan Fisher, 30.v.2015, in the Stonerose Interpretive Center collections; paratype 4: SR 02-32-23 ( Fig. 48 ), collected at Republic B 4131 by Chris Engle, 6.vi.2002, in the Stonerose Interpretive Center collections; paratype 5: SR 07-22-05AB ( Fig. 47 ), collected at Republic B 4131 by Kathleen Malloch 13.viii.2006, in the Stonerose Interpretive Center collections; paratype 6: SR 19-005-002AB ( Fig. 48 ), collected at Republic B 4131 by Gregg Wilson, 1.v.2019, in the Stonerose Interpretive Center collections.

Description. Holotype, SR 13-005-013, wing ( Fig. 44 ). Mostly complete, with portions of mid-posterior not preserved. Petiolate. Evenly oval distal to nodus, nodus to apex 22.4 mm, maximum width mid-wing 7.7 mm. Hyaline except for broad dark fascia across wing between level of nodus, just over half way to pterostigma. Pterostigma broad, about 2.5 times longer than wide, proximal side nearly vertical; subtends 2.5 cells. Oblique brace vein joins RA at proximal corner of pterostigma. Most crossveins roughly aligned in proximal half of postnodal, postsubnodal spaces. Costal space distal to pterostigma two cells wide. RA meets margin posterior to apex, RA slightly sigmoidal near margin. RA–RP1, RP1–IR1 spaces narrow near margin. RA–RP1 space mostly one cell wide, briefly two near apex. RP1–IR1 space two cells wide four cells distal to origin of IR1. Origin of IR1 presumed four cells distal to origin of RP2, mid-way between arculus, pterostigma. IR1–RP2, RP2–IR2 spaces two cells wide shortly proximal to level of pterostigma. Origin of RP2 about 5–6 cells distal to subnodus. RP2 mostly rather straight, curved slightly posteriorly in distal quarter. IR2: origin at subnodus; mostly rather straight, linear, slightly zigzagged in distal quarter, strongly so near margin. Origin of RP3-4 about 2/3 between arculus, nodus; mostly rather straight. MA linear, straight at least to half way from quadrangle (partially not preserved), strongly zigzagged at least distal third. MP linear, rather straight from quadrangle to terminus. CuA linear until shortly after level of nodus, then increasingly zigzagged. CuA–A space with 4 cells at widest (from just proximal to, distal to level of RP2 origin as preserved).

Paratype 1, SR 11-43-09, wing ( Fig. 45 ). Petiolate. Evenly oval distal to nodus; nodus to apex 23.5 mm: maximum 7.2 mm wide. Hyaline except for broad dark fascia across wing between level of nodus, just over half way to pterostigma. Pterostigma broad, about 2.5 times longer than wide; proximal side with shallow angle to costa. Region of oblique brace vein not preserved. Crossveins in postnodal, postsubnodal spaces mostly rather aligned. Two cells wide distal to pterostigma in costal space. RA–RP1 space one cell wide. RA meets margin posterior to apex. RP1 –IR1 space goes to two cells wide five cells distal to origin of IR1; narrows near margin. Origin of IR1 four cells distal to origin of RP2 , mid-way between arculus, pterostigma. IR1– RP2 , RP2 –IR2 spaces mostly not preserved. RP2 : origin about 3 cells distal to subnodus; proximal half linear, rather straight; distal half not preserved. IR2: origin at subnodus; mostly rather straight, liner, slightly zigzagged in distal quarter, missing by preservation near margin. RP3-4 : origin <2/3 between arculus, nodus; mostly rather straight (not preserved near margin). MA linear, straight at least to half way from quadrangle; slightly zigzagged in small portion preserved distal to this. MP linear, rather straight from quadrangle to terminus in regions preserved. CuA slightly zigzagged from level of quadrangle to level of origin of RP2 , mostly not preserved apical to this. CuA–A space with 4 cells wide shortly proximal to level of origin of RP2 (portion distal to this missing)

Paratype 2, SR 18-002-001, wing ( Fig. 46 ). Petiolate. Evenly oval distal to nodus (missing apical region), nodus to base of pterostigma 16.5 mm, maximum width mid-wing 7.2 mm. Hyaline except for broad dark fascia across wing between level of nodus, to about presumed 2/3 distance to pterostigma. Pterostigma not preserved. Most crossveins roughly aligned in proximal half of postnodal, postsubnodal spaces, not in distal half. RP1 –IR1 space two cells wide eight cells distal to origin of IR1. IR1 origin two cells distal to origin of RP2 . RP2 : origin apparently five cells distal to subnodus; mostly rather straight in region preserved. IR2: origin at subnodus; mostly rather straight, linear, apical quarter missing. RP3-4 : origin missing; mostly rather straight, missing apical quarter. MA linear, straight at least to half way between nodus, presumed level of pterostigma. MP linear, rather straight from quadrangle to terminus. CuA linear until shortly after level of nodus, then slightly zigzagged to distal quarter, then increasingly so. CuA–A space four cells at widest from just proximal to level of origin of RP2 to about level IP2 origin .

Paratype 3, SR 15-003-001, wing ( Fig. 47 ). Proximal two-thirds. Petiolate. Maximum width 7.7 mm. Hyaline proximal to level of nodus, darkly infuscate distal to it. Most crossveins roughly aligned in proximal half preserved of postnodal, postsubnodal spaces. RP1 – IR1 space two cells wide five cells distal to origin of IR1. IR1: origin four cells distal to origin of RP2 . RP2 : origin five cells distal to subnodus; mostly rather straight in region preserved. IR2: origin at subnodus; mostly rather straight, linear, apical half missing. RP3-4 : origin missing; mostly rather straight, missing apical third. MA mostly linear, slightly zigzagged around levels of origin of RP2 to IR1, mostly not preserved distal to that. MP mostly linear, slightly zigzagged in portions, gently curved from quadrangle, near terminus not preserved. CuA rather linear to level of origin of RP2 , then increasingly zigzagged. CuA–A space four cells at widest from just proximal to, to just distal to level of origin of RP2 .

Paratype 4, SR 02-32-23, wing ( Fig. 48 ). Apical half, missing much of anterior region. Apical portion hyaline from about half way between origin of IR1, pterostigma; darkly infuscate proximal to this. Pterostigma broad, proximal portion not preserved. Postnodal, most of postsubnodal spaces not preserved. Costal space distal to pterostigma two cells wide. RA meets margin posterior to apex. RA–RP1, RP1 –IR1 spaces narrow near margin. RA–RP1 space mostly one cell wide, briefly two near apex. RP1 –IR1 space constantly two cells wide twelve cells distal to origin of IR1. Origin of IR1 presumed three cells distal to origin of RP2 . IR1– RP2 space two cells wide at presumed level of proximal side of pterostigma; IR2– RP3-4 space two cells wide from a few cells proximal to this. RP2 : linear, mostly rather straight, slightly curved posteriorly in apical quarter. IR2 straight, linear to level RP2 –IR2 space becomes two cells wide, then slightly zigzagged, curves posteriorly near terminus. RP3-4 : mostly rather straight, wing damaged near margin. MA preserved portion linear to about level of origin of IR1, then increasingly zigzagged. MP preserved portion linear, slightly curved to terminus. CuA increasingly zigzagged in region preserved, strongly so apically. CuA–A space four cells wide from just proximal to level of RP2 origin to just distal to level of origin of IR1 .

Paratype 5, SR 07-22-05 A, B, wing ( Fig. 47 ). Parts of proximal half. Petiolate. Hyaline proximal to level of nodus, darkly infuscate distal to it. Most crossveins in postnodal, postsubnodal spaces roughly aligned in region preserved. Origin of IR1 three cells distal to origin of RP2 . RP2 : origin 2.5 cells distal to subnodus; portion preserved rather straight, linear. IR2: origin at subnodus; straight, linear. RP3-4 , MA, MP: origins, distal portions missing; preserved portion mostly rather straight, rather linear. CuA–A space 4 cells wide just proximal to level of RP2 origin, not preserved distal to this .

Paratype 6, SR 19-005-002 A, B, wing ( Fig. 48 ). Broken up middle portion from closely distal to nodus to base of pterostigma, posterior region missing or damaged. Proximal portion darkly infuscate, distal portion hyaline. Many crossveins roughly aligned in preserved portion of postnodal, postsubnodal spaces. IR1 origin four cells distal to origin of RP2 . RP2 , IR2, RP3-4 , MA: preserved portions mostly linear, rather straight, except MA zigzagging in distal portion preserved. Small portions of MP reserved. Disarticulated fragment of CuA–A space preserved, four cells wide .

Etymology. The specific epithet is a patronymic formed from the surname of Jackie and Tim Jeppesen, the collectors and donors of the holotype, recognising their contribution. The Latin suffix – orum indicates the genitive case of plural nouns.

Range and age. Tom Thumb Tuff Member of the Klondike Mountain Formation, locality B4131 [and other maybe?] at Republic, Washington, USA; second half of the Ypresian.

Figures

FIGURE 44. Okanopteryx jeppesenorum holotype, SR 13-005-013. A, photograph of part SR 13-005-013A; B, photograph of counterpart SR 13-005-013B; C, drawing from part and counterpart.All to scale, 5 mm

FIGURE 45. Okanopteryx jeppesenorum paratype 1, SR 11-43-09. A, photograph; B, drawing. Both to scale, 5 mm.

FIGURE 46. Okanopteryx jeppesenorum paratype 2, SR 18-002-001. A, photograph; B, drawing. Both to scale, 5 mm.

FIGURE 47. Okanopteryx jeppesenorum paratype 5, SR 07-22-05. A, photograph; B, drawing; paratype 3, SR 15-003-001: C, photograph; D, drawing. All to scale, 5 mm.

FIGURE 48. Okanopteryx jeppesenorum paratype 4, SR 02-32-23. A, photograph; B, drawing; paratype 6, SR 19-005-002: C, photograph; D, drawing. All to scale, 5 mm.