Jassa, , Vader & Krapp, 2005

Conlan, Kathleen E., Desiderato, Andrea & Beermann, Jan, 2021, Jassa (Crustacea: Amphipoda): a new morphological and molecular assessment of the genus, Zootaxa 4939 (1), pp. 1-191 : 162-163

Key to the world species of Jassa (both sexes) 6

1 Gnathopod 2, basis, at least the distal part of the anterolateral margin bearing a fringe of setae (at least some setal lengths 20–40% or more of the maximal basis width) (e.g., Figs 15 , 28 , 78 , 88 and 102 ) (fringe alsopresent in juveniles but may be less pronouncedthanin adults...e.g., see Fig. 102 ) ............................................................... 2

- Gnathopod 2, basis, setae short, minute or absent (setal lengths <20% of the maximal basis width) (e.g., Figs 37 , 85 , 95 and 104 )............................................................................................... 13

2 Gnathopod 2, propodus, anteroproximal margin with a row of long setae (setal length Ẑthe maximal basis width). Gnathopod 1, carpuswithoutasetaorclusterofsetaeattheanterodistaljunctionofthepropodus ( Figs 67 and 68 ) ... J. staudei Conlan, 1990

- Gnathopod 2, propodus, setae on the anteroproximal margin short or absent (setal length <65% of the maximal basis width). Gnathopod 1, carpus, seta(e) at the anterodistal junction of the propodus present or absent (may be slightly medial or lateral and aslongasthecarpusorveryshort) (e.g., Figs 15 , 22 , 65 and 88–90 ) ................................................3

3 Gnathopod 2, female, palmsinuous (e.g., Figs 88 , 97 and 102 ).....................................................4

- Gnathopod 2, female, palmconcave (e.g., Figs 15 , 26 , 29 and 65 )..................................................6

4 Gnathopod 1, basis, anterior margin with a row of spine-like setae; carpus, anterodistal margin with a single or cluster of setae at thejunctionof thepropodus ( Figs 88–90 ) ................................................. J. alonsoae Conlan, 1990

- Gnathopod 1, basis, anteriormarginwithoutarowof spine-likesetae ( Figs 85 and 102 ) ................................5

5 Gnathopod 1, carpus, anterodistal margin with a seta or cluster of setae at the anterodistal junction of the propodus (seta(e) ~40% thelengthof the carpus) ( Fig. 85 ) ......................................................... J. myersi Conlan , 19907

- Gnathopod 1, carpus, anterodistal margin without a seta or cluster of setae at the junction of the propodus ( Fig. 102 ).......... .................................................................................... J. gruneri Conlan, 1990

6 Uropod 1, peduncular spinous process that extends ventrally from the peduncle and underlies the rami very short (±10% of the lengthof thelongest ramus) ( Fig. 65 ) ................................................... J. borowskyae Conlan, 1990

- Uropod 1, peduncular spinous process that extends ventrally from the peduncle and underlies the rami at least 25% to 50% the lengthofthelongestramus (e.g., Figs 42 , 88 and 99 ) ............................................................7

7 Gnathopod 1, carpus with a seta or cluster of setae at the anterodistal junction of the propodus (seta or setal cluster may be slightly lateralormedialandshortoraslongasthe carpus) (e.g., Figs 15 , 22 , 29 , 31 , 62 and 81 ) ................................8

- Gnathopod 1, carpuswithoutasetaorclusterofsetaeattheanterodistaljunctionofthepropodus ... J. monodon ( Heller, 1866)

8 Gnathopod 1, carpus, seta or cluster of setae at the anterodistal junction of the propodus short (length <25% of the length of the carpus) andslightlymedialorlateral ( Figs 15 and 31 )............................................................9

- Gnathopod 1, carpus, seta or cluster of setae at the anterodistal junction of the propodus long (length 25–50% of the length of the carpusor longer) andslightlymedialorlateral ( Figs 22 , 29 , 62 and 81 ).............................................10

9 Telson, tip bearing a seta or setae extending between the third uropods (these setae are in addition to the usual upright setae at each lateral cusp and are visible when the uropods are pressed downwards and away from the telson). Antenna 2, thumbed males and adult females of any size without plumose setae on the posterior margin of article 5 and the flagellum (though setae may look finely pectinate). Gnathopod 2, propodus of major form thumbed male, thumb conical, tip acute, spine or spine group on the posteriormarginattheoriginof thethumbabsent (character statesdifferintheminor form) ( Figs 31–33 ) ................... ..................................................................................... J. valida ( Dana, 1853)

- Telson, tip without apical setae (though the usual upright setae at each lateral cusp are present). Antenna 2, large thumbed males and adult females, with dense plumose setae on the posterior margin of article 2 and the flagellum. Gnathopod 2, propodus of major form thumbed male, thumb wide, tip squared, spine or spine group on the posterior margin at the origin of the thumb absent (character statesdifferintheminor form) ( Figs 15–16 )..................................... J. marmorata Holmes, 1905

10 Gnathopod 2, basis and propodus, anterior marginal setae abundant and plumose ( Figs 62–63 ). Mandibular palp, article 2, with a fringeof setaeonthedorsalmargin ( Fig. 64 ) ................................................. J. oclairi Conlan, 1990

- Gnathopod 2, basis and propodus, anterior marginal setae sparse and simple ( Figs 22 , 28 and 78 ). Mandibular palp, article 2, withoutafringeofsetaeonthedorsalmargin ( Figs 27 , 30 and 82 ) ................................................11

11 Telson, tip bearing apical seta or setae in addition to the usual upright seta or setae at each lateral cusp ( Fig. 28 ).............. .................................................................................... J. morinoi Conlan, 1990

- Telson, tipwithoutapical setae, onlywiththeusualuprightsetaorsetaeateachlateralcusp ( Figs 22 and 78 ) ..............12

12 Antenna 2, large thumbed male and large adult female, peduncular article 5 and flagellum, posterior margin bearing dense plumose setae ( Figs 22 and 23 ). Gnathopod 2, propodus of major form male, thumb tip acute; spine or spine group on the posterior margin at the origin of the thumb absent (although present in small thumbed males) ( Figs 22 and 23 ). Gnathopod 1, female propodus, palm straight to shallowly concave. Gnathopod 2, female propodus, maximum width about 50% of maximum length, palmar anglenotclosetothedefining spines; spinestightly clustered ( Fig. 26 ) ........................... J. slatteryi Conlan, 1990

- Antenna 2, thumbed male and large adult female, peduncular article 5 and flagellum setae not plumose (though may be microscopically pectinate) ( Fig. 81 ). Gnathopod 2, propodus of major form male, thumb tip angled posteriorly, spine or spine group on the posterior margin at the origin of the thumb present ( Fig. 78 ). Gnathopod 1, female propodus, palm convex. Gnathopod 2, female propodus, maximum width about 65% of maximum length, palmar angle close to the defining spines; spines sequential

6 This key is limited by incomplete knowledge of J. kimi , J. laurieae , J. monodon and J. shawi . Sufficient information for J. kimi , J. monodon and J. shawi allow for their inclusion but Jassa laurieae has been excluded because its characteristics are unknown posterior of pereon segment 4. See Table 11 and Remarks section for J. laurieae for distinguishing character states.

7 J. myersi appearstwiceinthekeybecausethesetalfringeontheanterolateralmarginofthebasisofgnathopod 2 canbe interpreted as being present or absent due to the sparseness of the setae on some individuals

( Fig. 81 ) ............................................................................. J. carltoni Conlan, 1990

13 Uropod 1, peduncular spinous process that extends ventrally from the peduncle and underlies the rami short (<10% of the length of thelongest ramus) (e.g., Figs 65 and 83 ). Onlyknownfromthe Northeastern Pacific................................14

- Uropod 1, peduncular spinous process that extends ventrally from the peduncle and underlies the rami at least 25% to 50% the lengthofthelongestramus) (e.g., Figs 42 , 88 and 99 ). Notknownfromthe Northeastern Pacific ........................15

14 Gnathopod 1, carpus, seta or setal cluster at the anterodistal junction of the propodus about 35% of the length of the carpus. Gnathopod 2, female propodus, palmsinuous ( Fig. 85 ) ............................................ J. myersi Conlan, 1990

- Gnathopod 1, carpus, seta or setal cluster at the anterodistal junction of the propodus minute (<15% of the length of the carpus) and slightly medial. Gnathopod 2, female propodus, palm concave, palmar defining angle acute and close to the defining spines ( Fig. 83 ) ............................................................................... J. shawi Conlan, 1990

15 Pereopods 5–7, anterior margin of the propodus, spines strong and spine row extending half or more of its full length, propodus expandedornotexpanded forgrasping ( Figs 92 , 95 and 99 ). .....................................................16

- Pereopods 5–7, anterior margin of the propodus, spines mostly on the distal half, propodus not expanded for grasping (e.g., Figs 42 , 48 , 93 and 96 )........................................................................................18

16 Gnathopod 1, carpus, seta or setal cluster at the anterodistal junction of the propodus about 1/3 the length of the carpus and slightly lateral and medial. Body length at maturity 15–25 mm (thumbed males and females with setose brood plates) ( Fig. 99 )........ .................................................................................... J. ingens ( Pfeffer, 1888)

- Gnathopod 1, carpus without a seta or setal cluster at the anterodistal junction of the propodus. Body length at maturity 5–9 mm (thumbedmalesandfemaleswithsetosebroodplates) ( Figs 92 and 95 ) ............................................17

17 Pereopods 5–7, anterior margin of the propodus expanded proximally for grasping. Antenna 2, thumbed male, posterior margin of article 5 andflagellumdenselyclothedinplumose setae. Gnathopod 2, female, palmof thepropodussinuous ( Fig. 95 ) ........ .................................................................................... J. fenwicki Conlan, 1990

- Pereopods 5–7, anterior margin of the propodus not expanded proximally for grasping. Antenna 2, thumbed male, posterior margin of article 5 and flagellum without plumose setae (though setae are minutely barbed). Gnathopod 2, female, palm of the propoduswithadistinctledgedistalof thepalmardefiningspines ( Fig. 92 ) .......................... J. justi Conlan, 1990

18 Gnathopod 1, carpus with a seta or setal cluster at the anterodistal junction of the propodus (length about 25% of the carpus length) ( Fig. 93 )............................................................................ J. thurstoni Conlan, 1990

- Gnathopod 1, carpus without a short seta or setal cluster at the anterodistal junction of the propodus (e.g., Figs 42–43 ), or if seta present, length <15% of thecarpuslength (e.g., Figs 60 and 96 ). ..................................................19

19 Mandibular palp, article 2 withafringeof setaeonthedorsalmargin ( Figs 47 and 53 ) ................................20

- Mandibular palp, article 2 withoutafringeofsetaeonthedorsalmargin ( Figs 41 , 61 and 98 ) ...........................21

20 Uropod 3, inner ramus bearing 1 or 2 spines centrally in addition to the usual spine at the tip ( Fig. 42 ). Antenna 2, large adult male and female, posterior margin of peduncular article 5 and flagellum article 1 bearing dense plumose setae ( Figs 42–43 ) (these absentinsmall juveniles). Gnathopod 2, female propodus, palmarangle bulbous, distantfromthedefiningspines ( Fig. 44 ). .... .................................................................................. J. falcata ( Montagu, 1808)

- Uropod 3, inner ramus without central spines (only the usual spine at the tip) ( Fig. 48 ). Antenna 2, large adult male and female, posterior margin of peduncular article 5 and flagellum article 1 without plumose setae ( Fig. 48 ) (although setae may be minutely barbed) ( Fig. 50 ). Gnathopod 2, female propodus, palmarangle acute, close tothedefining spines ( Figs 51 and 52 ) ........... ................................................................................. J. herdmani ( Walker, 1893)

21 Gnathopods 1 and 2, basis, anteriormarginbearingafringeof setaeorspines ( Fig. 104 ) ......... J. hartmannae Conlan, 1990

- Gnathopods 1 and 2, basis, anterior margin without an obvious fringe of setae or spines (setae minute or restricted distally ( Figs 37 , 60 and 96 ) ..........................................................................................22

22 Gnathopod 2, female, palm sinuous ( Fig. 97 ). Gnathopod 2, propodus of the major form thumbed male, thumb short with acute tip, spineorspinegroupontheposteriormarginattheoriginof thethumb onaledge ( Fig. 96 ). Southern Hemisphere ......... .......................................................................... J. kjetilanna Vaderand Krapp, 2005

- Gnathopod 2, female, palm concave ( Fig. 37 ). Gnathopod 2, propodus of the major form thumbed male, thumb long with rounded or incised tip, spine or spine group on the posterior margin at the origin of the thumb residual or absent ( Figs 39 and 60 ). Northern Hemisphere ............................................................................................23

23 Antenna 2 markedly larger than antenna 1, width up to 2x the width of antenna 1 in the thumbed male. Gnathopod 2, propodus of themajorformthumbedmale, thumbtipnotincised ( Fig. 60 ). North Pacific ............................... J. kimi n. sp.

- Antenna 2 not markedly larger than antenna 1, width up to 1.5x the width of antenna 1 in the thumbed male. Gnathopod 2, propodusof themajorformthumbedmale, thumbtipincised ( Figs 37–39 ). North Atlantic.................. J. pusilla ( Sars, 1894)

Figures

FIGURE 15. Jassa marmorata Holmes, 1905. Neotype, adult male, major form, 6.7 mm, NMCC 1987-1067; adult female, 5.3 mm. Barren Islands Marina, Jamaica Bay Wildlife Refuge, Brooklyn, New York (40°35ʹN, 73°55ʹW), early July 1984, B. Borowsky, coll., among hydroids and Ulva on submerged docks at the marina, IZ 1984-112 (CMN). Lateral views: whole body, pereopod 3 and uropod 3; dorsal view: telson; other views medial. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 22. Jassa slatteryi Conlan, 1990. Holotype, adult male, major form, 5.1 mm, NMCC 1987-1068. Moss Landing Harbor, California (36°48’00.7ʺN, 121°47’13.9ʺW), 4 July 1986, K. E. Conlan, coll., station C9, scraping of the bottom of a row boat just below water level, IZ 1986-057 (CMN). Lateral view: whole body and gnathopod 1; dorsal view: telson; other views medial. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 23. Jassa slatteryi Conlan, 1990. Paratype, juvenile male, 2.7 mm; paratype, subadult male, 4.5 mm; paratype, minor form male, 3.5 mm; holotype, major form male, 5.1 mm. Moss Landing Harbor, California (36°48’00.7ʺN, 121°47’13.9ʺW), 4 July 1986, K. E. Conlan, coll., station C9, scraping of the bottom of a row boat just below water level, IZ 1986-057 (CMN). Setae omitted except for those around the thumb and spines in order to landmark position changes with growth. Lateral view: second gnathopods; medial view: second antennae. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 26. Jassa slatteryi Conlan, 1990. Intersex 1, 4.6 mm; intersex 2, 5.6 mm. Tanabe Bay, Wakayama Prefecture, Japan (33°41ʹN, 135°22ʹE), 2 Apr. 1976, H. Morino, coll., no habitat data, A2020.0036 (CMN). Allotype, adult female, 4.1 mm, NMCC 1987-1069. Moss Landing Harbor, California (36°48’00.7ʺN, 121°47’13.9ʺW), 4 July 1986, K. E. Conlan, coll., sta- tion C9, scraping of the bottom of a row boat just below water level, IZ 1986-057 (CMN). Lateral views: intersex 1 and 2 left gnathopod 2; other views medial. Scale 0.1 mm.

FIGURE 27. Jassa slatteryi Conlan, 1990. Holotype, adult male, major form, 5.1 mm, NMCC 1987-1068. Moss Landing Harbor, California (36°48’00.7ʺN, 121°47’13.9ʺW), 4 July 1986, K. E. Conlan, coll., station C9, scraping of the bottom of a row boat just below water level, IZ 1986-057 (CMN). Mouthparts. Frontal view: upper lip; lateral view: maxilla 1; other views medial. Scale 0.1 mm.

FIGURE 28. Jassa morinoi Conlan, 1990. Holotype, adult male, minor form, 5.6 mm, NSMT-Cr 26015 (C-33-1-1); paratype, minor form male 1, 4.7 mm, NSMT-Cr 26018 (C033-1-4). Bansho-no-hana Point, Tanabe Bay, Wakayama Prefecture, Japan (~33°42ʹN, 135°20ʹE), 17 March 1971, H. Morino, coll., low intertidal zone, on Sargassum microcanthum. Major form male 1, ~5–6 mm, western Crete, date unknown, G. Krapp Schickel, coll., 1 m+ depth, Schwanune and Corallina, NHMW 7622. Major form male 2, 3.75 mm, Stillwater Cove, Carmel Bay, California, June 1985, P. N. Slattery, coll., 35 ft. depth, rock and coralline bottom, IZ1989-024 (CMN). Subadult male, 4.2 mm, with larger thumb showing inside, and juvenile male, 3.5 mm, south end of Long Beach, Vancouver I., British Columbia, 6 July, 1970, E. L. Bousfield, coll., low intertidal zone, exposed coast, bedrock, kelp, Phyllospadix and fucoids, 1970-152, station P702 (CMN). Setae omitted on the gnathopod 2 profiles except for those around the thumb and spines in order to landmark position changes with growth. Lateral views: whole body and all second gnathopods; dorsal view: telson; medial view: uropod 3. Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 29. Jassa morinoi Conlan, 1990.Allotype, adult female, 5.8 mm, NSMT-Cr 26016 (C-33-1-2). Bansho-no-hana Point, Tanabe Bay, Wakayama Prefecture, Japan (~33°42ʹN, 135°20ʹE), 17 March 1971, H. Morino, coll., low intertidal zone, on Sargassum microcanthum. Major form male 2, 3.75 mm, Stillwater Cove, Carmel Bay, California, June 1985, P. N. Slattery, coll., 35 ft. depth, rock and coralline bottom, IZ1989-024 (CMN). All views medial. Scale 0.1 mm.

FIGURE 30. Jassa morinoi Conlan, 1990. Holotype, adult male, major form, 5.6 mm, NSMT-Cr 26015 (C-33-1-1). Banshono-hana Point, Tanabe Bay, Wakayama Prefecture, Japan (~33°42ʹN, 135°20ʹE), 17 March 1971, H. Morino, coll., low intertidal zone, on Sargassum microcanthum. Mouthparts. Frontal view: upper lip; other views medial. Scale 0.1 mm.

FIGURE 31. Jassa valida (Dana, 1853). Neotype, adult male, major form, 5.5 mm, JSIAH327. Babitonga, Brazil (26.239°S, 48.647°W), 5 Sept. 2017, A. Desiderato, coll., from settling plate, 1.5 m depth (MNRJcarcino 029820, accession no. BMFVF063- 19). Lateral views: whole body, pleopod 3 and uropods; dorsal view: telson; medial view: gnathopod 1. Scale 0.1 mm.

FIGURE 37. Jassa pusilla (Sars, 1894). Paralectotype, adult female 1, about 4.5 mm, UiO F.10562 Mp 41. Hammerfest, Norway (71°N, 23.9°E), date unknown, G. O. Sars, coll., drawn from the slide mounted specimen and also illustrated in Sars (1894), plate 213 as Podocerus pusillus Sars, 1894. Adult female 2, body length unknown, UiO F.10564 Mp 41. Locality, date and collector unknown, illustrated in Sars (1894), plate 213 as Podocerus pusillus Sars, 1894. Adult female 3, 4.7 mm, UiO F.13359. Riŝr, Norway (approx. 59°N, 9°E), date unknown, G. O. Sars, coll. Lateral view: whole body, gnathopods 1 and 2 and pereopod 3; dorsal view: telson; medial view: uropod 3. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 39. Jassa pusilla (Sars, 1894). Variation in male gnathopod 2 propodus and dactyl morphology within a single collection made by G. O. Sars at Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll., UiO F. 13359. Body and thumb lengths of each specimen are shown in FIGURE 40. All views medial. Scale 0.1 mm.

FIGURE 41. Jassa pusilla (Sars, 1894).Adult female 3 from FIGURE 37, 4.7 mm,, UiO F.13359. Riŝr, Norway (~ 59°N, 9°E), date unknown, G. O. Sars, coll. Mouthparts. Frontal view: upper lip; lateral views: maxilla 1 and right and left mandibles; other views medial. Scale 0.1 mm.

FIGURE 42. Jassa falcata (Montagu, 1808), type species for the genus Jassa. Holotype, adult male, minor form, 6.3 mm. Torcross, South Devon, England, date unknown, collector unknown, NHM 603a. Lateral views: whole body and right uropods 1-3; dorsal view: telson; other views medial. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 44. Jassa falcata (Montagu, 1808). Adult male 1, major form, 11.0 mm; juvenile male 2, 7.0 mm; adult male 3, major form, 6.5 mm; juvenile male 4, 4.5 mm; and adult female, 7.0 mm. Stoer Bay, Sutherland, Scotland, 22 October 1937, D. M. Reid, coll., very abundant on the floats of coastal salmon nets (SNM). Developing new cuticle is visible inside the propodus and dactyl of the juvenile male 2 gnathopod 2, not yet at the terminal molt as the new cuticle lacks a thumb. All views medial. Scale 0.1 mm.

FIGURE 47. Jassa falcata (Montagu, 1808). Holotype, adult male, minor form, 6.3 mm. Mouthparts. Frontal view: upper lip; lateral views: maxilla 1; other views medial. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 48. Jassa herdmani (Walker, 1893). Neotype, adult male, minor form, 6.0 mm, NHM 1925.9.8: 1686. Port Erin breakwater, Isle of Man, England, November 1892, A. O. Walker, coll.Adult male 1, major form, 7.9 mm, 1 m below water surface on steel frame used to hold settling plates, Langstone Harbour, Plymouth, England, 26 March 1986, R. J. Trott, coll., IZ1986-041 (CMN). Lateral view: whole body; dorsal view: telson; other views medial. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 50. Jassa herdmani (Walker, 1893). Adaptation of illustrations in Sexton and Reid (1951), with their figure numbers indicated. Antennae 1 (1A) and 2 (1B), gnathopod 1 (1C) pereopods 3–7 (1E, F, G, H, J): Plates 6-9, Specimen 1, Narrow-Form male J.CCLXXIII, from Trinity Buoy, Plymouth Docks, 14 May 1928, E. W. Sexton and D. W. Reid, colls.

FIGURE 51. Jassa herdmani (Walker, 1893). Adaptation of illustrations in Sexton and Reid (1951), with their figure numbers indicated. Adult female gnathopod 2: Plate 10, Specimen 2, Narrow-Form female J.CXLIV (2E), dredging from a buoy at the entrance to Plymouth Docks, 23 August 1928, E. W. Sexton and D. W. Reid, colls. Juvenile male gnathopod 2: Plate 12, Specimen 7, Narrow-Form male, cut-across stage, J.5 (1): F1 of female J.CXLIV (7C). Transformation of non-thumbed (13D) to thumbed (13E) gnathopod 2: Plate 19, Specimen 13 (2), Male K. B. 11, from a scrape of a buoy moored outside the [Plymouth] Breakwater, 3 May 1930, E. W. Sexton and D. W. Reid, colls. Adult male gnathopod 2, major form: Plate 7, Specimen 1, Narrow-Form male J.CCLXXIII, from Trinity Buoy, Plymouth Docks, 14 May 1928, E. W. Sexton and D. W. Reid, colls.

FIGURE 52. Jassa herdmani (Walker, 1893). Adult female, 7.2 mm, 1 m below water surface on steel frame used to hold settling plates, Langstone Harbour, Plymouth, England, 26 March 1986, R. J. Trott, coll., IZ1986-041 (CMN). Lateral view: whole body; dorsal view: telson; other views medial. Scale 0.1 mm.

FIGURE 53. Jassa herdmani (Walker, 1893).Adult male 1, major form, 7.9 mm, 1 m below water surface on steel frame used to hold settling plates, Langstone Harbour, Plymouth, England, 26 March 1986, R. J. Trott, coll., IZ1986-041 (CMN). Mouthparts. Lateral views: maxilla 1 and maxilla 2; frontal view: upper lip; other views medial. Scale 0.1 mm.

FIGURE 60. Jassa kimi n. sp. Holotype, adult male, major form, 3.5 mm, DKU-115 (MARBK). Impo Port, Yeosu-Si, The Republic of Korea, 23 June 2011, S.-S. Hong, coll. (MARBK). Lateral view: whole body; dorsal view: telson; medial view: gnathopod 1 and uropod 3. Scale 0.1 mm.

FIGURE 61. Jassa kimi n. sp. Holotype, adult male, major form, 3.5 mm, DKU-115 (MARBK). Impo Port, Yeosu-Si, The Republic of Korea, 23 June 2011, S.-S. Hong, coll. (MARBK). Mouthparts. Frontal view: upper lip; lateral view: maxilla 1 and right mandible; other views medial. Scale 0.1 mm.

FIGURE 62. Jassa oclairi Conlan, 1990. Holotype, adult male, minor form, 7.7 mm, NMCC 1987-1071; allotype, adult female, 7.6 mm, NMCC 1987-1072; paratypes, juvenile male 1, 7.1 mm, NMCC 1987-1073; juvenile male 2, 4.1 mm, NMCC 1987- 1073 (CMN). Square Bay, Amchitka Island, Alaska (51°27.2ʹN, 170°11.5ʹE), 21 September 1969, C. E. O’Clair, coll., station CT 02069, amongst kelp community dominated by Laminaria longipes on exposed rocky shore in the intertidal zone below MLLW, 1976-30 (CMN). Lateral view: whole body; dorsal view: telson; other views medial. Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 64. Jassa oclairi Conlan, 1990. Holotype, adult male, minor form, 7.7 mm, NMCC 1987-1071 (CMN). Mouthparts. Frontal view: upper lip; lateral view: maxilla 1; other views medial. Scale 0.1 mm.

FIGURE 65. Jassa borowskyae Conlan, 1990. Holotype, adult male, minor form, 4.5 mm, NMCC 1987-1064; allotype, adult female, 5.0 mm, NMCC 1987-1065. Brady’s Beach, Barkley Sound, Vancouver Island, British Columbia, 26 June 1976, E. L. Bousfield, coll., station B5, scrapes of boulders embedded in sand, Phyllospadix, kelp, brown algae, at low water, 1976-157 (CMN). Subadult male, 4.3 mm; adult male, major form, 5.0 mm. Point Grenville, Washington, 22 July 1966, E. L. Bousfield, coll., station W22, scrapes of boulders embedded in sand at low water, 1966-211 (CMN). Lateral views: whole body and uropods; dorsal view: telson; other views medial.Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 67. Jassa staudei Conlan, 1990. Holotype, adult male, major form, 6.0 mm, NMCC 1987-1074; allotype, adult female, 5.5 mm, NMCC 1987-1075. North side of Wouwer Island, Broken Group, Barkley Sound, Vancouver Island, British Columbia (48°51.06ʹN, 125°21.08ʹW), 27 June 1976, E. L. Bousfield, coll., station B7, dip net collection at low water, amongst Phyllospadix Hooker (1840), kelp, sponges and under rock, 1976-157 (CMN). Lateral view: whole body; dorsal view: telson; other views medial. Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 68. Jassa staudei Conlan, 1990. Holotype, adult male, major form, 6.0 mm, NMCC 1987-1074; allotype, adult female, 5.5 mm, NMCC 1987-1075; paratype, adult male 1, minor form, 5.1 mm, NMCC 1987-1076. Adult male 2, minor form, 4.2 mm, Edward King Island, British Columbia, 10 July 1976, E. L. Bousfield, coll., station B28, dip net collection at low water, amongst Phyllospadix, kelp, sponges and under rock, 1976-157 (CMN). All views medial. Appendages are of the holotype unless noted otherwise. Scale 0.1 mm.

FIGURE 78. Jassa carltoni Conlan, 1990. Holotype, adult male, major form, 3.0 mm, Corona del Mar, California, J. L. Barnard, coll., 9–11 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide (NHMLA 62-132.1). Paratype, juvenile male, 1.7 mm, La Jolla, California, 11–13 Nov. 1962, J. L. Barnard, coll., station 45-R- 1, no further collection data, 62-135.1 (NHMLA); subadult male, 4.0 mm, Cable Beach, Vancouver Island, British Columbia, 14 July 1972, D. Kittle, coll., station 342, 1973-128 (CMN); paratype, adult male, minor form, 2.8 mm, La Jolla, California, 11–13 Nov. 1962, J. L. Barnard, coll., station 45-W-5, scraping of calcareous sponge under ledge in surge channel at low tide, 62-134.1 (NHMLA); paratype, adult male 2, major form, 2.9 mm, Hazard Canyon Reef, Montana de Oro State Park, St. Louis Obispo County, California, 1–9 Dec. 1961, J. L. Barnard, coll., station 42-T-5, wash of Egregia holdfasts collected at low tide, 61-100.2 (NHMLA); adult male 3, major form, 4.0 mm, Sarita Bay, Vancouver Island, British Columbia, 2 July 1976, E. L. Bousfield, coll., station B13, 1976-157, bedrock, muddy sand, eelgrass, algae and fine gravel subtidally, hand and dip net collection at low tide, ~14 °C, 26 psu (CMN). Lateral view: whole body and gnathopod 2 views; dorsal view: telson; other views medial. Setae omitted from the juvenile male and major male 2 second gnathopods. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 81. Jassa carltoni Conlan, 1990.Allotype, adult female, 2.8 mm, 62-132.2 (NHMLA). Corona del Mar, California, J. L. Barnard, coll., 9–11 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 82. Jassa carltoni Conlan, 1990. Holotype, adult male, major form, 3.0 mm, 62-132.1 (NHMLA). Corona del Mar, California, J. L. Barnard, coll., 9–11 December 1962, station 46-K-1, wash of 1/6 liter of ascidians and soft polychaete tubes collected at low tide. Mouthparts. Frontal view: upper lip; lateral view: maxilla 1; other views medial. Scale 0.1 mm.

FIGURE 83. Jassa shawi Conlan, 1990. Holotype, juvenile male, 6.8 mm, NMCC 1987-1133; allotype, adult female, ovigerous, 5.6 mm, NMCC 1987-1134 (CMN). Bamfield Marine Sciences Centre, Barkley Sound, Vancouver Island, British Columbia, 24 June 1976, E. L. Bousfield, coll., station B1, on settling blocks suspended from floating dock at the water’s surface, 1976-157 (CMN). Lateral view: whole body; dorsal views: uropod 1 magnification, telson; other views medial.Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 85. Jassa myersi Conlan, 1990. Holotype, adult male, 5.8 mm, NHMLA 61-125.1; allotype, adult female, 4.4 mm, NHMLA 125.2; paratype, juvenile male, 3.7 mm, NHMLA 125.3. Coal Oil Point, Goleta, California, 2 July 1961, J. L. Barnard, coll., station 39-H-2, on rock surfaces in Phyllospadix zone, low intertidal zone. Adult male 1, 4.5 mm and subadult male, 5.1 mm, Coal Oil Point, Goleta, California, 2 July 1961, J. L Barnard, coll., station 39-K-1, on rock surfaces in Phyllospadix zone, low intertidal zone (NHMLA). Lateral views: whole body and uropods; dorsal view: telson; other views medial. Appendages are of the holotype unless noted otherwise. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 88. Jassa alonsoae Conlan, 1990. Holotype, adult male, major form, 11.1 mm, ZMH K-33619; allotype, female, 9.8 mm, ZMH K-33635; paratype, adult male, minor form, 9.3 mm; paratype, large juvenile male, 12.8 mm; paratype, small juvenile male, 4.6 mm. South Georgia (54°30’58ʺS, 36°0’45ʺW), 16 January 1884, Deutsche Polar Commission, K. von den Steinen, coll., station 7804 (ZMH K-8017A). All views lateral. Setae omitted from the gnathopod 2 profiles except for those around the thumb and spines of the males in order to landmark position changes with growth. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 92. Jassa justi Conlan, 1990. Holotype, adult male, major form, 8.5 mm, AM P34953; allotype, female, 6.9 mm, AM P37920; paratypes, subadult male, 5.4 mm and juvenile male 4.2 mm, AM P37921. Garden Bay, Macquarie Island (54°29ʹS, 158°58ʹE), 20 November 1977, J. K. Lowry and D. S. Horning, colls., station MA-10, among mixed red and green algae on rocky bottom, -3 m (AM). All views lateral. Setae omitted from the gnathopod 2 profiles except for those around the thumb and spines of the males in order to landmark position changes with growth. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 93. Jassa thurstoni Conlan, 1990. Holotype, adult male, major form, 5.9 mm, NHM 1969:761:2. Paal Harbour, Signy Island, South Orkney Islands (60°43ʹS, 45°36ʹW), 5 February 1965, M. H. Thurston, coll., station 44 (1218).Anchor dredge over mud at 20–25 m depth (NHM).Adult male 1, major form, paratype, 6.0 mm, NMCC-1082. Observation Bluff, Outer Islet, Signy Island, South Orkney Islands (60°42ʹ30ʺS, 45°35ʹ10ʺW), 14 September 1965, M. H. Thurston, coll., station 52 (2085), Agassiz trawl over gravel and sand, sand with some rock, algae on rocks, 20 m depth, NHM 1969:763:1 donated to CMN, accession no. IZ 1989-013. Adult female, paratype, 6.5 mm, NHM 1969:762:3. Berntsen Point–Bare Rock, Signy Island, South Orkney Islands (60°42ʹ10ʺS, 45°35ʹ30ʺW), 16 October 1964, M. H. Thurston, coll., station 50 (AT 76). Agassiz trawl over gravel and sand, some rocks with Desmarestia anceps and Phyllogigas grandifolius, 5–10 m depth (NHM). Lateral view: whole body, antenna 1 and gnathopod magnifications; medial view: uropod 3. Setae omitted from the gnathopod 2 profiles except for those around the thumb and spines of the male 1 in order to landmark position changes with growth. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 95. Jassa fenwicki Conlan, 1990. Holotype, adult male, major form, 7.4 mm, AM P34951. Punui Bay, The Snares Islands, New Zealand (48°01′ S, 166°32′ E), 21 January 1977, G. D. Fenwick, coll. (AM).Allotype, subadult female, Cumberland Bay, South Georgia (54°14′ S, 36°28′ W), 9 May 1902, among algae, 1–2 m depth, type no. 3946 from lot no. 3015 (NRM). All views lateral. Setae omitted from the female gnathopod 2 profile. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 96. Jassa kjetilanna Vader and Krapp, 2005. Adult male, major form, 6.5 mm. Choiseul Sound, East Falkland, 29 October 2003, A. and W. Vader, coll., from the lithodid crab Paralomis granulosa (Hombron & Jacquinot, 1846) at 20–30 m depth (TSZ Cr 19054). Lateral view: whole body and uropod 3; dorsal view: telson; other views medial. Scale 0.1 mm.

FIGURE 97. Jassa kjetilanna Vader and Krapp, 2005. Adult female, 5.0 mm; subadult male, 5.0 mm; juvenile male, 3.5 mm. Choiseul Sound, East Falkland, 29 October 2003, A. and W. Vader, coll., from the lithodid crab Paralomis granulosa (Hombron & Jacquinot, 1846) at 20–30 m depth (TSZ Cr 19054). All views medial. Scale 0.1 mm.

FIGURE 98. Jassa kjetilanna Vader and Krapp, 2005. Adult male, major form, 6.5 mm. Choiseul Sound, East Falkland, 29 October 2003, A. and W. Vader, coll., from the lithodid crab Paralomis granulosa (Hombron & Jacquinot, 1846) at 20–30 m depth (TSZ Cr 19054). Mouthparts. Frontal view: upper lip; lateral view: maxillae 1 and 2; other views medial. Scale 0.1 mm.

FIGURE 99. Jassa ingens (Pfeffer, 1888). Whole body: lectotype, adult male, major form, 22.1 mm, ZMH K-8017. Moltke- Hafen, Royal Bucht, South Georgia (54°30ʹ58ʺS, 36°0ʹ45ʺW), 16 Januarly 1884, Deutsche Polarstation 1882/83, K. Von den Steinen, coll. (ZMH). Adult male 1, major form, 20.3 mm and juvenile male, 10.6 mm, NHM 1969:705:9. Billie Rocks, Signy Island, South Orkney Islands (60°42ʹS, 45°36ʹW), 5 March 1965, M. H. Thurston, coll., station 10 (97). SCUBA collection on vertical creviced rock with Lithothamnia, Desmarestia anceps, Ascoseira mirabilis, Iophon/Phyllophora, Porifera, Polyzoa, 1.9–2.2 m depth, IZ 1989-013 (CMN). Subadult male, 19.9 mm, NHM 1969:713:4. Billie Rocks, Signy Island, South Orkney Islands (60°42ʹS, 45°36ʹW), 27 February 1965, M. H. Thurston, coll., station 20 (512), SCUBA collection on rock inclined at 45–50°with overlying boulders and sand in crevices. Lithothamnia, Desmarestia anceps, Rhodophyceae, Polyzoa, 7.4–7.7 m depth (NHM). All views lateral. Setae omitted from the gnathopod 2 profiles except for those around the thumb and spines of the males in order to landmark position changes with growth. Scale 1 mm. Illustration after Conlan (1990).

FIGURE 102. Jassa gruneri Conlan, 1990. Holotype, juvenile male, 2.1 mm, NMCC 1987-1077; allotype, adult female, 3.2 mm, NMCC 1987-1078. Hobart Harbour, Tasmania (42°50ʹS, 147°20ʹE), 8 November 1978, G. Edgar, coll., SCUBA, algal washes, 3–5 m depth, 1978-287 (CMN). Lateral views. Scale 0.1 mm. Illustration after Conlan (1990).

FIGURE 104. Jassa hartmannae Conlan, 1990. Holotype, adult male, major form, 4.5 mm, AM P34950. Senecio Pool, The Snares, New Zealand (48°07ʹS, 166°38ʹE), 6 January 1977, G. D. Fenwick, coll., from Lessonia variegata holdfast, 1.5 m depth (AM). Allotype, adult female, NMNH 235008; paratypes, adult male, minor form, 3.1 mm and juvenile male, 1.8 mm, NMNH 235009, St. Clair, Dunedin, New Zealand (45°50ʹS, 17°20ʹE), 25 January 1968, J. L. Barnard, coll., station E973 (J. L. Barnard station NZ-8), formalin wash of algae, especially of pterocladians or streblocladians, heavy turf (Gelidium-Bostrychia) and several specimens of Pyura sp. (NMNH). Setae omitted on the gnathopod 2 profiles except for those around the thumb and spines of the males in order to landmark position changes with growth. All views lateral. Scale 0.1 mm. Illustration after Conlan (1990).