Edaphus

Puthz, V., 2014, Review of the New World Edaphus MOTSCHULSKY (Coleoptera, Staphylinidae) 120 th Contribution to the Knowledge of Euaesthetinae, Linzer biologische Beiträge 46 (1), pp. 799-827 : 814-822

Key to the New World Edaphus

1 Pronotum with (additional) puncture-like impressions on disc......................................2

- No (additional) puncture-like impressions on pronotal disc...........................................9

2 Two puncture-like impressions on pronotal disc. unknown. 1.3-1.9 mm (FB: 0.7- 0.8 mm). Paraguay, Venezuela, Panama................................................ biforatus PUTHZ

- More than two puncture-like impressions on pronotal disc............................................3

3 Pronotum with a distinct mbc.........................................................................................4

- Pronotum without a distinct mbc....................................................................................5

4 Larger: 1.4-1.5 mm (FB 0.9 mm), pronotum with more than 16 puncture-like impressions on disc (fig. 11).: A (fig. 48). Ecuador................... ecuadorensis nov.sp.

- Smaller: 0.95 mm (FB 0.55 mm), pronotum with 16 punctures on disc (fig. 8, OROUSSET, 1986). Venezuela............................................................ poulaini OROUSSET

5 Elytra punctate (fig. 7).: A (fig. 60). 1.0 mm (FB 0.6 mm). Peru variipunctus nov.sp.

- Elytra impunctate...........................................................................................................6

6 Smaller: 1.3 mm (FB 0.7 mm), pronotum with many very large puncture-like impressions (fig. 14). A (fig. 44). Peru............................................. foraminosus nov.sp.

- Larger: FB t 0.8 mm, pronotum with less large puncture-like impressions....................7

7 Pronotum beside of larger discal impressions finely punctate (fig. 12).: E (fig. 50). 1.4-1.5mm (FB 0.8-0.9 mm). Peru............................................. foveolicollis nov.sp.

- Pronotum beside of larger discal impressions impunctate..............................................8

8 Bf of pronotum of about equal size (fig. 13). unknown. 1.6 mm (FB 1.1.- 1.2 mm). Ecuador......................................................................................... scupolai nov.sp.

- Bf of pronotum of different size.: A (fig. 5, OROUSSET, 1987). 1.7 mm. Ecuador....... .......................................................................................................... huggerti OROUSSET

9 Antennal club 3-segmented..........................................................................................10

- Antennal club 2-segmented..........................................................................................21

10 Lbc present...................................................................................................................11

- Lbc absent....................................................................................................................14

11 Humeral angle with a distinct carina (fig 6, PUTHZ, 1973a).: A (fig. 10, l. c.). 1.2- 1.3mm (FB 0.75mm). Brazil: São Paulo .......................................... angulipennis PUTHZ

- Humeral angle simple...................................................................................................12

12 Mbc present. A remarkably slender species. unknown. 1.4mm (FB 0.7mm). Paraguay........................................................................................... phloeocharis PUTHZ

- Mbc absent, broader species.........................................................................................14

13 Pronotum and elytra extremely finely, densely, distinctly punctate, pronotum and elytra narrower.: S 8 and A (fig. 16, and as in fig. 4, PUTHZ, 2007. Panama, Costa Rica.......................................................................................................... hidalgo PUTHZ

- Pronotum and elytra at 60 x almost impunctate (setiferous punctures barely noticeable, pronotum broader.: S 8 and A (as in figs 14, 17, PUTHZ, 2007). Panama, Costa Rica............................................................................... caballero PUTHZ

14 Sides of elytra with an elevated narrow boarder and with granules. 2.2-2.5 mm (FB 1.1 mm). Ecuador............................................................................... limbipennis PUTHZ

- Sides of elytra simple...................................................................................................15

15 Brachypterous, PL>SL................................................................................................16

- Macropterous, PL <SL.................................................................................................18

16 Sides of pronotum strongly angular (about as in fig 1, PUTHZ, 1973a). 2.0- 3.2mm (FB 1.0- 1.3mm).: A (fig. 12, PUTHZ 1973a). Colombia, Venezuela............................ ......................................................................................................... columbianus PUTHZ

- Sides of pronotum less angular. Smaller species..........................................................17

17 AlpF impunctate.: S 8 (fig. 20), A (fig. 47). 1.6-2.1mm (FB 0.9-1.1mm) ................... ................................................................................ variipennis nov.sp. (brachypterous)

- AlpF with some punctures anteriorly.: S 8 (fig. 19), A (figs 45, 46). 1.5-2.1mm (FB 0.9-1.0mm).................................................................................... electrinus nov.sp.

18 Elytra finely punctate...................................................................................................19

- Elytra impunctate (aside of minute setiferous punctures)............................................20

19 Larger, punctation of pronotum very fine, distinct.: S 8 and A (figs 14, 17, PUTHZ 2007). 2.2-2.6mm (FB 1.4mm). Ecuador.................................................... kapac PUTHZ

- Less large, 1.6-2.1mm (FB 1.0- 1.2mm), punctation of pronotum barely noticeable.: S 8 and A (figs 13, 15, PUTHZ 2007). 1.8-2.1mm (FB 1.2mm). Peru........ ....................................................................................................................... apo PUTHZ

20 Shoulders rectangular, sides of elytra not dilated posteriad.: S 8 and A (figs 37, 56). 1.0- 1.1mm (FB 0.6-0.65mm). Peru........................................................................... ............................ atahualpa nov.sp. (see also # 17 variipennis nov.sp. (macropterous))

- Shoulders oblique, sides of elytra dilated posteriad. unknown. 1.8-2.2mm (FB 1.0- 1.2mm). Peru.......................................................................................... bufo PUTHZ

21 Lbc absent....................................................................................................................22

- Lbc present...................................................................................................................46

22 Elytra with a long curved sulcus in lateral half. 3.4-4.0 mm (FB 2.0 mm). Venezuela........................................................................................ sulcatipennis PUTHZ

- Elytra otherwise...........................................................................................................23

23 Elytra with a deep transverse humeral and a circular parascutellar impression. unknown. 1.5-1.7mm (FB 0.8mm). Ecuador............................................. chinan PUTHZ

- Elytra otherwise...........................................................................................................24

24 Elytra simple. Small blackish species with two impressions on elytra. Lbc indistinct or absent.: Figs 1-3, OROUSSET, 1986). 1.2-1.3 mm (FB 0.6 mm). Venezuela............ ............................................................................................................. najtae OROUSSET

- Elytra otherwise...........................................................................................................25

25 Brachypterous or elytra shorter, PL>SL......................................................................26

- Macropterous, PL <SL.................................................................................................33

26 Pronotum broader, PW: PL t1.15. 1.2-1.4mm (FB 0.8mm). unknown. Habitus (fig. 8, PUTHZ, 2006a). 1.3-1.5 mm (FB 0.8mm). Venezuela.............. cimicoides PUTHZ

- Pronotum less broad, PW: PL <.115............................................................................27

27 Temples at least half as long as eyes............................................................................28

- Temples much shorter than eyes..................................................................................29

28 Eyes smaller, pronotum with 6 distinct bf; forebody almost impunctate.: A (fig. 6, PUTHZ, 2006a). 1.1-1.5mm (FB 0.6-0.7 mm). Peru...................... longepilosus PUTHZ

- Eyes larger, pronotum with 4 (-6) indistinctly delimited bf; forebody finely punctate.: A (fig. 10, PUTHZ, 2007). 1.3-1.7mm (FB 0.7-0.75 mm). Venezuela.......... ..................................................................................................................... eliab PUTHZ

29 Elytra distinctly punctate..............................................................................................30

- Elytra impunctate (aside of setiferous punctures)........................................................31

30 Larger, 1.6-1.9mm (FB 0.8mm). Head and pronotum reddish brown, elytra and abdomen blackish brown (fig. 7, PUTHZ 2006a); pronotum and abdomen punctate. unknown. 1.6-1.9 mm (FB 0.8mm). Ecuador........................................... inka PUTHZ

- Smaller. 1.0- 1.4mm (FB 0.6-07mm).: S 8 and A (figs 8, 9, PUTHZ, 2007). Venezuela.................................................................................................... david PUTHZ

31 Larger, 1.2-1.3mm (FB 0.9mm). S 8 and A (figs 24, 25, PUTHZ, 2007). 1.2-1.5mm (FB 0.9mm). Costa Rica........................................................................... kethleyi PUTHZ

- Smaller.........................................................................................................................32

32 Habitus (fig. 39, PUTHZ, 2007).: S 8 and A (figs 41, 43, PUTHZ, l.c.). 1.0- 1.2mm (FB 0.7mm). Costa Rica..................................................................... dybasianus PUTHZ

-: S8 and A (figs 42, 43, PUTHZ, l. c.). 1.0- 1.2mm (FB 0.7mm). Costa Rica.................. ......................................................................................................... wagnerianus PUTHZ

33 Pronotum distinctly punctate........................................................................................34

- Pronotum impunctate or indistinctly punctate (minute setiferous punctures)..............35

34 Temples distinct. Smaller, FB 0.7mm.: S 8 and A (figs 19, 21, PUTHZ, 2006a). 1.1-1.4mm. Panama.................................................................................... irenae PUTHZ

- Temples indistinct. Larger, FB 0.8mm.: A (resembling fig. 13, PUTHZ, 1973). 1.2-1.5 mm. Mexico: Oaxaca ................................................................... hirsutus PUTHZ

35 Mbct3 absent ................................................................................................................36

- Mbct3 present, can be very short..................................................................................37

36 Bf distinctly delimited. PtfF semicircular.: S 8 and A (figs 45, 46, PUTHZ, l. c.). 1.1- 1.4 mm (FB 0.85mm). Venezuela........................................ subtilipunctatus PUTHZ

- Bf indistinctly delimited. PtfF angular (trapezoid). S 8 and A (figs 51, 52, PUTHZ, 2007). 1.0- 1.4 mm. Ecuador, Peru............................................................... villac PUTHZ

37 Mbct3 nearly as long as the tergite.: A (fig. 58). 1.1-1.2mm (FB 0.75mm). Ecuador................................................................................................. molestus nov.sp.

- Mbct3 shorter...............................................................................................................38

38 AmpF punctate. Pronotum with 4 bf.: S 8 and A (figs 49, 56, PUTHZ, l. c.). 1.0- 1.2mm. Peru............................................................................................... cinche PUTHZ

- AmpF impunctate.........................................................................................................39

39 Mbct 3 t one third of the length of tergite 3.................................................................40

- Mbct3 <one third of the length of tergite 3.................................................................43

40 Larger, FB t 0.9mm ......................................................................................................41

- Smaller, FB d 0.8mm ....................................................................................................42

41 Pronotum broader, PW: PL>1.05.: S 3-6 with medial impressions. S 8 and A (figs 21, 22, PUTHZ, 2007). 1.4-1.7mm (FB 0.9mm). Costa Rica............... bierigi PUTHZ

- Pronotum less broad, PW: PL <1.05.: S 3-6 simple. S 8 and A (figs 4, 5, PUTHZ, 2001). 1.5-1.7mm (FB 1.0mm). Mexico.............................................. andersoni PUTHZ

42 Antennal segment 10 about as broad as long. Pronotum with 4-6 bf.: S 5 with a projecting lobe posteromedially. A ( Fig. 13 , PUTHZ, 1973b). 1.1-1.5 mm. (FB 0.75- 0.8mm). Mexico: Chiapas; Guatemala................................................. mexicanus PUTHZ

- Antennal segment 10 broader than long. Pronotum with 8 bf.: S 5 without a projecting lobe, S 8 and A (figs 19, 20, PUTHZ, 2007). 1.2-1.5 mm. (FB 0.8mm). Panama............................................................................................... panamensis PUTHZ

43 Larger, FB t 0.8mm ......................................................................................................44

- Smaller, FB 0.7mm.: SpP (fig. 64), A (fig. 63). 1.0- 1.1mm. Chile... chilensis nov.sp.

44 Pronotum broader, PW: PL>1.10, with 6-8 small bf.: S 8 and A (figs 53, 54, PUTHZ 2007). 1.2-1.5mm (FB 0.8-0.85mm). Bolivia........................ [ bolivianus PUTHZ ]

- Pronotum less broad, PW: PL <1.05, with 6 bf............................................................45

45 Elytra shorter, trapezoid.: S 8 and A (figs 34, 35, PUTHZ, 2007). 1.3-1.5mm (FB 0.85-0.9mm). Costa Rica............................................................................ amigo PUTHZ

- Elytra longer, subquadrate. S 8 and A (figs 5, 6, PUTHZ, 2007). 1.3-1.7 mm. (FB 0.9mm). Bolivia................................................................................. similifactus PUTHZ

46 Small blackish species with two impressions on elytra. see above see #13 ..................... ............................................................................................................. najtae OROUSSET

- Reddish or reddish brown species with simple elytra..................................................47

47 At least tergite 3 with an atrium-pocket or with a prolongated atrium elevation (figs 1-3, 15-18, 25, PUTHZ 2006b)...........................................................(ventralis -group) 48

- Tergites simple, no atrium-pockets present..................................................................63

48 Tergite 3 with a narrow, oblique atrium-pocket (fig. 10). Base of pronotum with 2 transverse bf.................................................................................................................49

- Tergites 1-4 with transverse atrium pockets (fig. 9). Base of pronotum different........52

49 Elytra almost impunctate.: Parameres of A with one apical seta.............................50

- Elytra very finely and densely punctate. unknown. 1.2-1.5mm (FB 0.75mm). Ecuador................................................................................................ asheianus PUTHZ

50 Smaller: 1.0- 1.3mm (FB 0.6-0.7mm)...........................................................................51

- Larger: 1.3-1.5mm (FB 0.8-0.85mm).: A (fig. 12, PUTHZ 2006b), parameres with two apical setae. Ecuador............................................................................. falini PUTHZ

51 Eyes smaller (figs 6, 7, l. c.).: Antennal segment 11 nearly 4x as long as broad. A (fig. 5, l. c.).: Head with short oblique temples (fig. 7, l. c). Costa Rica, Panama, Nicaragua............................................................................................... ventralis PUTHZ - Eyes larger (figs 10, 11, l. c.).: Antennal segment 11 about 1.5x as long as broad. A (fig. 9).: Head without distinct temples (fig. 11, l. c.). Peru, Ecuador...................... ............................................................................................................. ventricula PUTHZ

52 Pronotum distinctly punctate, with two postero-lateral denticles. 1.9-2.2mm (FB 1.3mm). Ecuador.................................................................................. denticollis PUTHZ

- Pronotum impunctate and without lateral denticles......................................................53

53 Mbct3 present...............................................................................................................54

- Mbct3 absent................................................................................................................56

54: Eyes very large with coarse ommatidia (fig. 13, l. c.). Distance between eyes d half HW. A (fig. 30, l. c.). 1.1-1.4mm. French Guiana............................... boops PUTHZ

-: Eyes smaller with smaller ommatidia (fig. 14, l. c.), Distance between eyes> half HW........................................................................................................................55

55 Smaller: FB 0.7-0.8mm.: Apical notch of S 8 less deep (fig. 23, l. c), A (fig. 22, l. c.), apical part of parameres free. 1.3-1.6mm. Suriname, Trinidad, Brazil, Peru,? Ecuador, Paraguay................................................................... rishwani MAKHAN

- Larger: FB 0.85-0.95mm.: Apical notch of S 8 deeper (fig. 24, l. c.), A (figs 20, 21, l. c.), apical part of parameres fused with the median lobe. 1.3-1.8mm. Brazil, French Guiana, Suriname, Guyana....................................................... humeralis PUTHZ

56 Atrium pockets of tergites 1-4 slightly broader than long (fig. 9)................................57

- Atrium pockets of tergites 1-4 much braoder than long (figs 15-18, l. c.)...................58

57 Larger: 0.9-1.1mm, pronotum narrower, mbct3 absent.: S 8 (fig. 27, l. c.), A (fig. 26, l. c.). Costa Rica........................................................................... schatzorum PUTHZ

- Smaller: 0.8mm, pronotum broader, mbct3 present.: S 8 (fig. 28, l. c.), A (fig. 29, l. c.). Panama: Canal Zone .................................................................. liliputanus PUTHZ

58 Lbc distinct and longer than bf.....................................................................................59

- Lbc indistinct or at most as long as bf (species from Cuba).........................................60

59 Larger: FB 0.7-0.8mm, mbf larger, longer than broad.: A (fig. 19, l. c.). 1.1- 1.5mm. Costa Rica, Panama, Guatemala............................................. marsupifer PUTHZ

- Smaller: FB 0.65mm, mbf smaller, almost as long as broad.: A (fig. 31, l. c.). 1.0- 1.2mm. Brazil...................................................................................... hoppi PUTHZ

60 Larger, FB 1.0mm, eyes larger, genae almost missing. unknown. 1.6-2.0mm. Cuba.................................................................................................. ferdinandae PUTHZ

- Smaller, FB less than 1.0mm, eyes smaller, genae distinct..........................................61

61 Pronotum and elytra with both: fine + distinct and very fine + shallow punctuation...62

- Punctation of pronotum and elytra simple.: S 8 (fig. 10, PUTHZ 1997). 1.0- 1.3mm. Cuba............................................................................................. juanae PUTHZ

62 Eyes larger, genae less long.: S 8 (fig. 6, l. c), A (fig. 5, l. c.). 1.1-1.5mm. Cuba........ ................................................................................................................ cubanus PUTHZ

- Eyes less larger, genae longer.: S 8 (fig. 1, l. c.), A (fig. 8, l. c.). 1.4-1.9mm. Cuba................................................................................................................................. .................................................................................................................. infidus PUTHZ

63 Elytra with broad epaulets (a distinct line-like humeral impression separates broad and slightly elevated epaulets) and postero-lateral impressions (fig. 1, PUTHZ, 1973a). Large species...................................................................................................64

- Elytra without broad epaulets but can have other humeral characters. Almost smaller species.............................................................................................................66

64 Longer, exceeding 2.2 mm ...........................................................................................65

- Shorter, less than 2.2 mm. Humeral impression at base with 2 small foveae. Postero-lateral outline of head (eyes + temples) acute angled. unknown. 1.7- 1.9mm (FB 1.1mm). Ecuador............................................................... praefectus PUTHZ

65 Mbc present.: S 8 and A (figs 4, 7, PUTHZ, 2007). 2.5-5.1 mm (!) (FB 1.4- 2.5mm). Venezuela, Ecuador.................................................................... goliath PUTHZ - Mbc absent.: A (fig. 11, PUTHZ, 1973a). 2.2-3.3mm. Brazil: São Paulo crassus PUTHZ

66 Base of pronotum with a mbc, extending to base proper..............................................67

- Base of pronotum without a mbc extending to base proper.........................................89

67 Brachypterous or apterous, PL>SL.............................................................................68

- Macropterous, PL <SL.................................................................................................69

68 Longer, 1.5-2.0mm (FB 0.8mm). Frons impunctate, temples distinct. Habitus (fig. 9, PUTHZ, 2006). unknown. 1.5-2.1mm (FB 0.8mm). Ecuador... trapezipennis PUTHZ

- Less long, 1.3mm (FB 0.6mm). Frons punctate, temples indistinct.: A (fig. 3, OROUSSET, 1986). 1.3-1.4mm (FB 0.65-0.7mm). Venezuela........ fuscescens OROUSSET

69 Elytra with an intrahumeral stria (fig. 8).: S 8 and A (figs 31, 51). 1.2-1.4mm (FB 0.8mm). Bolivia....................................................................................... hanleyi nov.sp.

- Elytra with or without an intrahumeral impression, but no stria..................................70

70 Pronotum distinctly punctate.: figs 28, 43 and figs 13-15 ( OROUSSET, 1983). 1.3- 1.7mm (FB 0.8-0.9mm). French Guiana, Guyana, Suriname........ tavakiliani OROUSSET

- Pronotum impunctate (aside of minute setiferous punctures).......................................71

71 Elytra distinctly punctate (at least near suture)............................................................72

- Elytra impunctate (aside of minute setiferous punctures)............................................74

72 Larger. 1.5-1.8mm (FB 0.9mm). Mbct3 about as long as the tergite.: A (about as in fig. 1, PUTHZ, 1974b). Brazil: Guanabara................................... congerminalis PUTHZ

- Smaller, FB <0.7mm. Mbct3 less than half as long as the tergite................................73

73 Elytral punctuation sparse, shallow, and extremely fine (can be more distinct near suture), elytra uniformly coloured.: Apical notch of S 8 half as deep or deeper than half the length of sternite. A (fig. 41, PUTHZ 1974). 1.0- 1.4mm. U. S. A.................. ........................................................................................................... americanus PUTHZ

- Elytra distinctly, moderately strongly punctate (strong, irregular, and moderately sparse on disc), mostly bicoloured: shoulders and posterior margin lighter (yellowish) than the rest of surface.: Apical notch of S 8 less deep than half the length of sternite, A (fig. 40). 1.2-1.5mm. U. S. A....................... nitidus MOTSCHULSKY

74 Pronotum broader, PW: PL t1.20................................................................................75

- Pronotum less broad, PW: PL <1.18............................................................................79

75 Bf distinct and distinctly delimited..............................................................................76

- Bf less distinctly delimited owing to a broad transverse impression on either side of the mbc.........................................................................................................................77

76 AmpF concave, abdomen shiny, setiferous punctures less distinct, less dense.: S 8 and A (figs 24, 61). 1.4-1.7mm (FB 0.9-1.0mm). Costa Rica, Panama......................... ........................................................................................................ inconveniens nov.sp.

- AmpF transversely elevated, abdomen moderately shiny, setiferous punctures more distinct, dense.: S 8, A and SpP (figs 15, 36, 55). 1.0- 1.3mm (FB 0.75mm). Suriname............................................................................................... torifrons nov.sp.

77 Temples distinct, antennal segment 10 longer than broad.: S 8 and A (figs 12, 13, PUTHZ 2006a). 1.3-1.6 mm (FB 0.8mm). Peru........................................ lescheni PUTHZ

- Distinct temples absent, antennal segment 10 broader than long.................................78

78 Mbct3 longer, about as long as the tergite, alpF strongly elevated.: S 8 and A (figs 9, 12, OROUSSET, 1983), habitus (fig. 2, l. c.). 1.3-1.8 mm (FB 0.7-0.8mm). French Guiana, Guyana, Suriname, Brazil: Para............................... remilleti OROUSSET

- Mbct3 shorter, about one half as long as the tergite, alpF slightly elevated.: S 8 and A (figs 5, 6, PUTHZ, 2002). 1.3mm (FB 0.7mm). Mexico: San Luis Potosi ............... ................................................................................................................... izcoatl PUTHZ

79 Temples distinct...........................................................................................................80

- Temples indistinct/absent.............................................................................................84

80 Larger, FB> 0.8mm ......................................................................................................83 - Smaller, FB d 0.7mm ....................................................................................................81

81 Mbf about three times as long as broad.: A (fig. 42). 0.9-1.1mm (FB 0.6-0.7mm). Holarctis, New Zealand (introduced) ................................................. lederi EPPELSHEIM

- Mbf about as long as broad..........................................................................................82

82 Tergites finely and densely punctate, mbct3 longer than half the tergite.: A (fig. 2, PUTHZ, 1974). 1.2-1.3mm (FB 0.7-0.8mm). U.S. A .................. politus MOTSCHULSKY

- Tergites indistinctly, very sparsely punctate, mbct3 nearly half as long as the tergite.: S 8 (fig. 26), A (figs16-18, 39). U. S. A.: Arkansas ...... inexspectatus nov.sp.

83 AmpF distinctly broader than each of the alpF, mbf about as broad as long.: S 8 and A (figs 10, 11, PUTHZ, 2006a). 1.3-1.7 mm (0.85-0.9mm). Venezuela...................... .......................................................................................................... venezolanus PUTHZ

- AmpF slightly less broad than each of the alpF, mbf slightly broader than long. unknown. 1.4-1.5mm FB 0.8-0.9mm). Mexico: Chiapas .............................. tizoc PUTHZ

84 Pronotum with 6 bf.......................................................................................................85

- Pronotum with 4 bf.......................................................................................................86

85 PtfF semicircular, ampF about as broad as each of the alpF.: Figs 2, 3 ( PUTHZ, 1990). 1.0- 1.2mm (FB 0.7mm). Ecuador............................................ quisquilius PUTHZ

- PtfF trapezoid, ampF nearly twice as broad as each of the alpF.: figs 2, 3 ( PUTHZ, 2001). 1.2-1.3mm (FB 0.7mm). Mexico: Chiapas, Nicaragua..................... ashei PUTHZ

86 Pronotum broader, PW: PL>1.10................................................................................87

- Pronotum less broad, PW: PL <1.10............................................................................88

87 Mbf twice as broad as long, ampF distinctly broader than each of the moderately elevated alpF. unknown. 1.0- 1.1mm (FB 0.7mm). Peru.................... peruanus PUTHZ

- Mbf about as broad as lbf, ampF about as broad as each of the strongly elevated alpF.: S 8 and A (figs 48, 50, PUTHZ, 2007). 0.8-1.1 (1.2?)mm. (FB 0.55-0.6 (0.7?)mm). Panama, Costa Rica (?), Honduras (?), Guatemala (?) .......... newtoni PUTHZ

88 Mbf twice as broad as lbf.: A (fig. 2, PUTHZ, 1978). 0.8-1.1 mm (FB 0.5-0.6mm). Brazil: Sta Catharina, Peru, Bolivia, Cuba? ....................................... plaumanni PUTHZ

- Mbf about as broad as lbf.: S 8 and A (figs 14, 15, PUTHZ, 2006). 1.0- 1.2mm (FB 0.65mm). Panama...................................................................... quadrifoveolatus PUTHZ

89 Shoulders with a minute carina/denticle.......................................................................90

- Shoulders simple..........................................................................................................95

90 Brachypterous, PL>SL................................................................................................91

- Macropterous, PL <SL.................................................................................................92

91 Mbct3 absent.: A (fig. 14, l. c.). 1.1-1.3mm (FB 0.6-0.65mm). Brazil: Alexandra, Parana ................................................................................................... depressus PUTHZ

- Mbct3 present.: S 8 and A (figs 32, 33, PUTHZ, 2007). 1.0- 1.2mm (FB 0.7mm). Costa Rica (short winged form of) .......................................................... wagneri PUTHZ

92 PW: PL>1.20.: S 8 and A (figs 27, 57). 1.1-1.2mm (FB 0.7mm). Venezuela............. ............................................................................................................. mediocris nov.sp.

- PW: PL d1.10...............................................................................................................93

93 AmpF very finely punctate.: S 8 and A ( Figs 28, 30, 31 , PUTHZ, l. c.). 1-1.3 mm (FB 0.7mm) ........................................................................................ gallinaceus PUTHZ

- AmpF impunctate.........................................................................................................94

94: S 8, SpP and A (figs. 16-18, PUTHZ, 2006). 1.0- 1.2mm (FB 0.7mm). Costa Rica...... ......................................................................................................... costaricensis PUTHZ

-: S 8 and A (figs 32, 33, PUTHZ, 2007). 1.0- 1.2mm. (FB 0.7mm). Costa Rica.............. ................................................................................................................ wagneri PUTHZ

95 Head broader, PW: HW <1.20.....................................................................................96

- Head less broad, PW: HW t1.20................................................................................108 96 Elytra with a very fine granular sculpture.: A (fig. 2, PUTHZ, 1974). 1.5-2.0mm FB 0.9-1.0mm). Brazil: Nova Teutonia, Paraguay...................... brasilianus WENDELER

- Elytra otherwise...........................................................................................................97

97 Anterior angles of pronotum acute...............................................................................98

- Anterior angles of pronotum rounded, no margin visible in dorsal aspect.................101

98 Antennal segment 10 broader than long. Sides of pronotum in anterior half convex. Mbct3 present...............................................................................................................99

- Antennal segment 10 longer than broad. Mbct3 absent. Sides of pronotum in anterior half straight. Mbct3 absent.: S 8 and A (figs 29, 54). 1.6-1.7mm (FB 0.95mm). Venezuela...................................................................... marginicollis nov.sp.

99 Longer, 1.5-1,9mm (FB 0.8-1.1mm)..........................................................................100

- Less long, 1.1-1.2mm (FB 0.7mm).: S 8 and A (figs 34, 62). 1.1-1.2mm (FB 0.7mm) Peru.................................................................................. angulosicollis nov.sp.

100 AmpF broader than each of the alpF.: S 8 and A (figs 25, 49). 1.5-1.7mm (FB 0.8mm). Paraguay.................................................................................. montoya nov.sp.

- AmpF about as broad as each of the alpF. Longer, 1.6-1.9mm (FB 1.1mm). unknown. Brazil: Rio de Janeiro ............................................................... bryanti PUTHZ

101 Mbct3 at least one third as long as the tergite............................................................102

- Mbct3 shorter than one third of the tergite.................................................................105

102 Antennal segment 10 distinctly broader than long.....................................................103

- Antennal segment 10 about as long as broad or slightly longer.................................104

103 Pronotum with 4 (-5) bf.: S 8 and A (figs 36, 37, PUTHZ, l.c.). 1.2-1.4mm. (FB 0.8mm). Panama, Costa Rica........................................................ andersonianus PUTHZ

- Pronotum with 6-8 bf. 1.2-1.6mm (FB 0.8mm). Panama, Costa Rica........ dybasi PUTHZ

104: S 8 and A (figs 38, 59). 1.1-1.4mm (FB 0.7-0.8mm). Venezuela..... brachati nov.sp.

-: S 8 and A (figs 53, 54, PUTHZ 2007). 1.2-1.5mm (FB 0.8-0.85mm). Bolivia............. ............................................................................................................. bolivianus PUTHZ

105 Pronotum broader, PW: PL t1.20..............................................................................106

- Pronotum less broad, PW: PL <1.20..........................................................................107

106 Larger, alpF strongly elevated.: S 8 and A (figs 32, 53). 1.6mm (FB 0.9mm). French Guiana..................................................................................... falinianus nov.sp.

- Smaller, alpF moderately elevated. unknown. 1.3mm (FB 0.75mm). Mexico: Hidalgo .................................................................................................. axayacatl PUTHZ

107 AmpF punctate.: S 8 and A (figs 1, 2 PUTHZ, 1990). 1.3-1.5mm (FB 0.85- 0.9mm). Brazil: Rio de Janeiro ............................................................ planifrons PUTHZ

- AmpF impunctate.......................................................................................................108

108 Larger, FB t 0.9mm ....................................................................................................109

- Smaller, FB d 0.8mm ..................................................................................................112

109 Pronotum with 4 bf, mbf much broader than lbf. unknown. 1.3-1.5mm (FB 0.9mm). Guatemala.......................................................................... euplectoides SHARP

- Pronotum with 4-6 bf, mbf not or slightly broader than lbf.......................................110

110 Mbct3 as long as the tergite........................................................................................111

- Mbct3 shorter than the tergite. 1.4-1.7mm (FB 0.9mm).: S 8 and A (figs 21, 22, PUTHZ, 2007). Costa Rica........................................................................... bierigi PUTHZ

111 Larger. unknown. 1.6-1.9mm (FB 1.1mm) Brazil: Rio de Janeiro. ...... bryanti PUTHZ

- Smaller.: S 8 and A (figs 26, 36, PUTHZ, 2011). 1.6mm (FB 0.9mm). Brazil: Bahia ......................................................................................................... agostii PUTHZ

112 Antennal segment 10 distinctly broader than long (e. g. fig 2b, PUTHZ, 1974).........113

- Antennal segment 10 about as long as broad.............................................................114 113 Mbct3 nearly as long as the tergite, head much narrower than pronotum.: A (fig. 1, PUTHZ, 1974). 1.3-1.5 (1.9) mm (FB 0.8mm). Brazil: Nova Teutonia, Paraguay........ ............................................................................................................... congener PUTHZ

- Mbct3 at most one half as long as the tergite, head about as broad as pronotum.: S 8, SpP and A (figs. 16-18, PUTHZ, 2006). 1.0- 1.2mm (FB 0.7mm). Costa Rica........... ......................................................................................................... costaricensis PUTHZ

114 Mbct3 about as long tergite 3. 1.3-1.5mm (FB d 0.8mm).: A (fig. 5, PUTHZ, 1985). Paraguay, Brazil: Sta. Catharina ............................................. torresbolloi PUTHZ

- Mbct3 shorter than tergite 3.......................................................................................115

115 Eyes smaller, LG> 1/3 of LE.: S 8 and A (figs 32, 33, PUTHZ, 2007). 1.0- 1.2mm (FB 0.7mm). Costa Rica.......................................................................... wagneri PUTHZ

- Eyes larger, LG as long as or shorter than 1/3 LE.: S 8 and A (figs 44, 47, PUTHZ, l.c.). 1.1-1.3mm (FB 0.7-0.8mm) ......................................................... siphonifer PUTHZ

Figures

Figs 6-18: Habitus (1), fore body (7, 8), tergite 3 (9, 10), pronotum (11-14), sperm pump and vesica seminalis (15), aedeagus and apical portion of aedeagus (16-18) of Edaphus dybasianus PUTHZ (6, PT), E. variipunctus nov.sp. (7, HT), E. hanleyi nov.sp. (8, HT), E. schatzorum PUTHZ (9, PT), E. ventralis PUTHZ (10, PT), E. ecuadorensis nov.sp. (11, HT), E. foveolicollis nov.sp. (12, HT), E. scupolai nov.sp. (13, HT), E. foraminosus nov.sp.(14, PT), E. torifrons nov.sp. (15, HT) and E. inexspectatus nov.sp. (16-18, HT).

Figs 19-44: Sternite 9 of male (19, 20, 23-38), elytra (21, 22) and aedeagus (39-44) of Edaphus electrinus nov.sp. (19, PT), E. variipennis nov.sp. (20-22), E. foveolicollis nov.sp. (23, HT), E. inconveniens nov.sp. (24, PT), E. montoya nov.sp. (25, HT), E. inexspectatus nov.sp. (26, 39), E. mediocris nov.sp. (27, HT), E. tavakiliani OROUSSET (28, 43, Suriname), E. marginicollis nov.sp. (29, HT), E. foraminosus nov.sp. (30, 44, HT), E. hanleyi nov.sp. (31, HT), E. falinianus nov.sp. (32, HT), E. ecuadorensis nov.sp. (33, HT), E. angulosicollis nov.sp. (34, PT), E. molestus nov.sp. (35, HT), E. torifrons nov.sp. (36, HT), E. atahualpa nov.sp. (37, HT), E. brachati nov.sp. (38, HT), E. nitidus MOTSCHULSKY (40, Kentucky), E. americanus PUTHZ (41, HT) and E. lederi Eppelsheim (42, Switzerland).- Scale = 0.1 mm (19= 20, 23= 24-38, 39= 40-44).