Miridiba gressitti ( Frey, 1970 ) Gao & Coca-Abia, 2021

Gao, Chuan-bu & Coca-Abia, María Milagro, 2021, Revision of the genus Miridiba Reitter, 1902 (Coleoptera, Scarabaeidae, Melolonthinae): genital morphotypes and new taxonomic data, European Journal of Taxonomy 749, pp. 1-94 : 48

Miridiba gressitti ( Frey, 1970) comb. nov.

Figs 5–7 , 65

Holotrichia gressitti Frey, 1970: 249 , 263 (type loc.: Vientiane, Laos).

Diagnosis

External morphology of adult ( Fig. 65 ). Body size 18.6 mm. Dorsal surface glabrous, shiny appearance. Clypeus shorter than frons, flat, oblique sides, anterior edge strongly emarginate at middle. Frons with carina moderately developed. Pronotal anterior margin glabrous, moderately wavy forward just behind eyes; posterior margin glabrous, finely flanged with a row of punctures except at middle; lateral margins moderately serrated almost smooth with a few short setae; anterior and posterior angles obtuse, anterior angle not projected forward. Prosternal process cone-shaped. Scutellum glabrous, with scattered punctures and sides without punctures. Elytra with punctures bigger than those of pronotum, epipleuron with a few short setae at basal part. Foretibia with dorsal carina moderately developed; insertion of inner spur closer to third outer tooth than second one. Meso- and metatibia with transverse carina interrupted at middle and complete, respectively; inner margin of dorsal surface with few spines. Prepygidium and pygidium with scattered punctures and glabrous, pygidium with apical margin pubescent, widened and moderately raised. Ventrites 3 and 4 with inconspicuous punctures and glabrous. Ventrite 5 not depressed at posterior half, with pubescence of different length, with conspicuous scattered punctures. Ventrite 6 moderately bulging at anterior margin, with long pubescence near posterior margin. Male and female genitalia as described above (genital morphotype II “ Gressitti ”). Male genitalia: parameres ( Fig. 5 ) with all branches of dorsal complex shorter than ventral branches. Endophallus ( Fig. 6 ) with a strong raspulae with numerous setae. Female genitalia ( Fig. 7 ).

Material examined

Paratype LAOS • 1 ♀; Vientiane; 15 Jun. 1965; Ban Van Eue leg.; G. Frey 1969 det.; NHMB .

Remarks

Frey (1970) described Holotrichia gressitti based on three female specimens designated as a holotype and two paratypes, remaining the male unknown. We have studied one paratype (NHMB) which shows external morphology characteristic of Miridiba . Moreover, female genitalia has features not shared with species of other genital morphotypes. Hence, the female genitalia of morphotype II “ Gressitti ” was described based on this species ( Fig. 7 ). Moreover, the male genitalia of M. gressitti is described based on a paratype of Miridiba herteli (see remarks under M. herteli ), which is described in morphotype II “ Gressitti ” ( Figs 5–6 ).

Distribution

Laos (Vientiane).

Figures

Figs 5–7. Morphotype II “Gressitti”. 5–6. Male genitalia of Miridiba herteli (Frey, 1971), paratype (NHMB). 5. Parameres in lateral view. 6. Endophallus and temones in dorsal view. 7. Female genitalia of M. gressitti (Frey, 1970), paratype (NHMB) in dorsal (A) and ventral (B) views. Scale bars = 1.0 mm.

Figs 65–68. 65. Miridiba gressitti (Frey, 1970), paratype, ♀ (MHNB). Habitus in dorsal view. 66– 68. M. herteli (Frey, 1971) holotype, ♂ (SNSD 10629). 66. Habitus in dorsal view. 67. Parameres in dorsal (A), ventral (B) and lateral (C) views, 68. Endophallus and temones in lateral view. Scale bars: 65–66 = 5.0 mm; 67–68 = 1.0 mm.