Turris pagasa Olivera, 2000

Kilburn, Richard N., Fedosov, Alexander E. & Olivera, Baldomero M., 2012, Revision of the genus Turris Batsch, 1789 (Gastropoda: Conoidea: Turridae) with the description of six new species, Zootaxa 3244 (1), pp. 1-58 : 41-43

Turris pagasa Olivera, 2000

Plate 25 , figs A–I

Turris pagasa Olivera, 2000: 304 , pl. 1, specimen 4, pl. 5; Olivera & Sysoev 2008: pl. 679, fig. 1 a, b. Type locality: Pamilacan Is., Bohol, Philippines, 100– 150 m.

Turris kilburni Vera-Peláez, Vega-Luz, & Lozano-Francisco, 2000: 7, pl. 1, fig. 8, pl. 2, fig. 2, pl. 6, figs 8–12; Robin 2008: p. 2008, fig. 14. Type loc.: Balicasag Is., near Panglao, Bohol Sea, Philippines. Syn. nov.

DESCRIPTION: Shell relatively thick, narrowly fusiform (b/l 0.25–0.30), with subulate, orthoconoid spire (a/l 0.42–0.48), siphonal canal relatively long (equal to rest of aperture) and recurved, in large examples with a fasciole and false umbilicus. Edge of outer lip fluted, lip in side view evenly convex; anal sinus fairly shallow, narrowly Vshaped or linear and slot-like, interior of outer lip with 6–9 spiral ridges in adult.

Whorls moderately convex, initially with 3 spiral ridges, the median one gemmulate; suture narrow but deep. In adults, subsutural cord relatively low, comprising an angular, ledge-like main ridge, but with age developing feeble, converging lamellae on either side. Sulcus approximately as wide as subsutural cord, moderately deep and steep-sided, but filled with scale-like collabral threads. Sinus cord low but forming a slight shoulder, as narrow as other cords, sometimes distinctly undulating—almost gemmulate - in lateral as well as apical view; crest of sinus cord, typically bearing 2 thin spiral threads. Characteristically the interspace on either side of sinus cord is 2–3 times its width. Base of spire whorls with 2–3 ridges, the uppermost peripheral, equal in width to their intervals, which bear collabral threads and occasionally faint spiral threads. Base of last whorl with 19–22 spiral ridges, upper ones widely-spaced, with a weaker spiral thread in each interval, ridges finer and closer on rostrum. All whorls with strong, lamellar collabral threads interstitially.

Ground colour white, usually tinged with pale fawn on lower part of spire whorls; spiral ridges patterned with dark reddish-brown quadrangular dots and spots, subsutural cord with dark brown blotches, interval between sinus cord and peripheral cord white (and wider than others).

Protoconch conical, of 2.8–3.3 whorls, nucleus small, smooth, other whorls with opisthocline axial riblets; maximum breadth ca 0.75 mm.

Attains 98 mm.

DISTRIBUTION: Central and southern Philippines to Japan and New Caledonia; 10–55 m [18–101 m] also recorded from 80– 300 m.

TYPES: T. pagasa : Holotype (pl. 26, figs A–B) and two paratypes in Olivera colln, holotype to be deposited in PNM . T. kilburni : Holotype MNCN 15.05/39977 (pl. 26, figs H–I) , paratypes in private hands .

OTHER MATERIAL EXAMINED: PHILIPPINES: Masbate Is., 10–20 m; Zamboanga, Laminusa Is., from fishermen; Balut Is., tangle nets in ca 150 m (all G. Poppe colln); Aliguay Is., Zamboanga, 25–40 fath. [46–73 m] NMSA: BO); Pamilcan Is, Bohol, 100–150 m (holotype T. pagasa ); Pamilacan Is, Bohol, 80–120 fath. [146–220 m]; off Dipolog, northern Mindanao, 80–150 m (Paratype 1, T. pagasa ), Bogo Is., 50–100 m (Paratype 2, T. pagasa ); Olango Is. and Leyte (all BO colln). NEW CALEDONIA: 19°05’S, 163°29’E, 230-240 m, MNHN; 22°42’S, 167°10’E, 315 m, MNHN; 24°45’S, 168°09’E, 230 m, MNHN .

REMARKS: Turris pagasa is characterised by its relatively low spiral cords with regular (usually small) brown spots overall, except for a relatively wide white or pale buff zone on either side of sinus cord (the anterior one typically the wider), and by all cords (including the unusually narrow sinus cord) being weakly crenulated. In life, the colour pattern is muted by a persistent, mainly interstitial, dull olive-brown periostracum (although this is usually cleaned off by the collector). The siphonal canal in adults is often longer and more strongly recurved than in other Turris species, and the shell thicker.

Figures

PLATE 25. Turris pagasa Olivera, 2000: A, B—Holotype, Pamilacan Is., Philippines, PNMM, 63.3 x 16.1 mm; C, D—Paratype 2, Bogo, Cebu Is., Philippines, BO colln, 97.9 x 23.9 mm; E—Pamilacan Is., Philippines, BO colln, 85.6 x 21.6 mm; F, G –Tanabe Bay, Japan, ANSP 86.8 x 23.1 mm; H, I—Holotype of Turris kilburni Vera-Peláez, Vega-Luz & Lozano-Francisco, 2000: Balicasag Is., Philippines, MNCN-15.05/39977, 78.6 x 21.2 mm.