Kmentia, Stehlík, 2013

Stehlík, Jaroslav L., 2013, Review and reclassification of the Old World genus Physopelta (Hemiptera: Heteroptera: Largidae), Acta Entomologica Musei Nationalis Pragae 53 (2), pp. 505-584 : 566-567

Kmentia gen. nov.

Type species. Lygaeus festivus Fabricius, 1803 , here designated.

Description. Body large (♂♂ 15.50–19.60 mm, ♀♀ 16.00–18.90 mm).Antennae in both sexes long, always longer than pronotum, sometimes even longer than combined length of head and pronotum; antennomere 1 longer than antennomere 2 ( Figs 93−94 ); there is great variability in length of antennae among specimens. As an example, the length of antennomeres of three large and three small males are given here: Large males (mm): 1 – 7.13, 6.59, 6.43; 2 – 6.32, 6.05, 5.56; 3 – 3.35, 3.19, 2.11; 4 – 3.89, 3.62, 3.59; small males (mm): 1 – 4.27, 4.86, 4.8l; 2 – 3.89, 4.32, 4.37; 3 – 2.11, 2.32, 2.38; 4 – 3.19, 3.24, 3.08.

Callar lobe only slightly gibbose, without sexual differences ( Figs 93−94 ). Labium reaching between mesocoxae. Profemora sexually dimorphic; in females more slender, with smaller number of small denticles; in small males profemora similar to those of females ( Figs 93−94 ). Ventral surface of profemora with short longitudinal furrow limited to its apical third; basal two-thirds medially convex with numerous denticles on the entire surface ( Figs 83 , 97 ). Protibiae ventrally unarmed ( Figs 95−96 ). Peritreme of metathoracic scent glands longitudinal, crescent-shaped, projecting both anteriad and posteriad ( Fig. 84 ).

Pygophore ( Figs 85−88 ). Ventral rim slightly concave, rounded; lateral rim more elevated than ventral rim, strongly rounded, descending towards dorsal rim. Lateral rim infolding only slightly sloping into the genital chamber; anal tube wide, long, reaching middle of genital chambre ( Fig. 86 ).

Parameres ( Figs 89−92 ). Parameres horizontal, their apices approaching and standing opposite each other, their apical portions bent dorsally, apex hook-shaped, pointed ( Figs 91−92 ), distinctly surpassing ventral rim of pygophore (see in lateral view − Fig. 88 ).

Female internal genitalia were described by DUVIARD (1975).

Differential diagnosis. Kmentia gen. nov. belongs to Physopeltinae because of the strongly S-shaped intersegmental sulci between the abdominal sternites. It differs in several characters from the other genera of the subfamily, warranting its placement in a tribe of its own – Kmentiini trib. nov.: labium reaching between mesocoxae; callar lobe weakly gibbose in both sexes; ventral surface of profemora in both sexes with only short longitudinal furrow limited to its apical third, basal two-thirds medially convex with numerous denticles on the entire surface; protibia in both sexes unarmed; and structure of the male genitalia (parameres horizontally positioned, their apices approaching and standing opposite each other ( Fig. 85 ), bent dorsally in their apical portion, apex hook-shaped, pointed dorsally ( Figs 85, 87 ), distinctly surpassing ventral rim of pygophore (see in lateral view − Fig. 88 ).

Etymology. This new genus is named in honour of my friend and colleague Petr Kment (Department of Entomology, National Museum, Prague) with thanks for his help. Gender is feminine.

Figures

Figs 93–98. Kmentia festiva (Fabricius, 1803). 93 – female; 94 – male; 95–96 – fore leg (95 – large male, 96 – small male); 97 – ventral surface of profemur. 98 – Physopelta (Neophysopelta) gutta gutta (Burmeister, 1834), ventral surface of profemur. (Photos: L. Dembický).

Figs 81–84.Kmentia festiva (Fabricius, 1803), male.81 – fore coxae and trochanters (magnification 47×); 82 – detail of procoxa and protrochanter (70×); 83 – ventral surface of profemur (32×); 84 – detail of peritreme (150×). Scale bars: 0.2 mm (Fig. 84), 0.5 mm (Figs 81–83). (SEM micrographs: P. Kment).

Figs 85–92. Kmentia festiva (Fabricius, 1803), male. 85–88 – pygophore: 85 – intact, dorso-posterior view (magnification 50×), 86 – dissected, dorsal view (55×), 87 – intact, posterior view (50×), 88 – intact, lateral view (60×). 89–92 – paramere (different orientations; 150×). Scale bars: 0.2 mm (Figs 89–92), 0.5 mm (Figs 85–88). (SEM micrographs: P. Kment).