Delta campaniforme campaniforme (Fabricius 1775)

Nugroho, Hari, Kojima, Jun-Ichi & Ubaidillah, Rosichon, 2013, Review of potter wasps with a petiolate metasoma excluding so-called " Zethinae " (Hymenoptera: Vespidae: Eumeninae) in the Lesser Sunda Islands of the Indonesian Archipelago, Zootaxa 3608 (1), pp. 1-25 : 9-11

Delta campaniforme campaniforme (Fabricius 1775)

(Figs 18, 20, 22, 24, 26, 28, 30)

Vespa campaniformis Fabricius 1775: 371 , Ƥ, holotype "Nova Hollandia [New Guinea]", [The Natural History Museum, London].

Delta campaniforme gracilior Giordani Soika 1986: 88 , Ƥ, holotype "Luzon: Matnog, Sorsogon", [Natural History Collection at Ibaraki University, Mito]. New synonymy.

This species, represented by many color forms, is widely distributed throughout the Oriental and Papuan (including northern Australia) regions. Currently more than ten forms are formally recognized as subspecies, but several of them are known to occur sympatrically and thus decision of their taxonomic status, whether they are only color variants or diagnosable species, needs further intensive comparative studies.

Based on the examination of the types of D. campaniforme gracilior and material mainly from the Lesser Sunda Islands, we have reached the conclusions, as detailed below, that D. c. gracilior Giordani Soika 1986 is a synonym of D. c. campaniforme (Fabricius 1775) ; D. c. nigriculum Giordani Soika 1986 is a good species and D. c. rendalloide Giordani Soika 1993 is a synonym of D. nigriculum .

Material examined. LOMBOK I.: 1 3 (MZB), 08°29'S 116°40'E, Labuhan Lombor, Lombor, Pringgabaya, J. Kojima, 8 Nov.2000. SUMBAWA I.: 1 Ƥ (MZB), 08°37'33.3"S 117°10'42.6"E, alt. 916 m, Tepal, Batu Lanteh, H. Nugroho & Y.R. Suhardjono, 14 Apr.2010. TIMOR I.: 1 Ƥ (MZB), Camplong Barat Daya, 26 Feb.1982; 1 Ƥ (MZB), Camplong, A. Hana, 23 Feb.1982; 1 3 (MZB), Camplong, A. Hana, 20 Feb.1982; 1 3 (MZB), Benlutu, Sudarman HK, 27 Mar.1982; 1 Ƥ (MZB), Sillu, A. Hana, 27 Oct.1982; 1 3 (MZB), Sillu, A. Hana, 27 Oct.1982; 1 Ƥ (MZB), 10°13'S 123°50'E, alt. 360m, Oekabiti, Amarasi, Kupang, J. Kojima, 2 Feb.2003; 1 Ƥ (IUNH), 10°11'S 123°35'E, Kupang, J. Kojima, 3 Feb.2003; 3 3 (MZB), 09°57'S 124°09'E, Boentuka, Batuputih, Timur Tengah Selatan, J. Kojima, 1 Feb.2003. PHILIPPINES: 1 Ƥ and 1 3 (IUNH; respectively holotype and paratype - allotype of D. c. gracilior ), Matnog, Sorsogon, Luzon I., J. Kojima, 5 Jun.1980.

Remarks. The last three females from Timor have a black longitudinal band dividing the basal ferruginous area of metasomal tergum 2; the lack of such a black longitudinal band is the character distinguishing gracilior from the nominotypical subspecies (Giordani Soika 1986). Our other specimens from the Lesser Sunda Islands including those from Timor agree well with the holotype and paratype (allotype) of D. c. gracilior , with the following variations in marking pattern [states in the holotype and paratype of D. c. gracilior are given in the brackets]: mesoscutum occasionally with paired lateral yellow bands [mesoscutum entirely black]; mesepisternum sometimes nearly entirely covered by yellow markings [yellow spots covering only dorsal half]. We do not find any grounds to consider D. c. gracilior even as a local color form and D. c. gracilior Giordani Soika, 1986 is synonymized under the nominotypical species.

FIGURES 17 – 30. Delta species. 17–20, 23, 24. Ƥ. 21, 22, 25–30. 3. 17, 19, 21, 23, 25, 27, 29. D. nigriculum . 18, 20, 22, 24, 26, 28, 30. D. c. campaniforme . 17, 18. Metasomal segments 1–2, lateral view. 19–22. Clypeus. 23, 24. Metasomal tergum 1, dorsal view. 25, 26. Apical part of antenna. 27, 28. Aedeagus, lateral view. 29, 30. Volsella. Scale 1mm.

Distribution. India, Nepal, Myanmar, Thailand, Malaysia (Peninsular Malaysia), Singapore, Philippines, Indonesia [Kalimantan, Sumatra, Java (including Bawean Island and Kangean Islands), Bali, Sulawesi, Moluccas, Lesser Sunda Islands: Lombok, Sumbawa, Sumba, Timor, Papua], Papua New Guinea, Australia; adventives in USA: Hawaii.

FIGURES 31 – 42. Delta species. 31–33, 40–42. Ƥ. 34–39. 3. 31, 34, 37, 40. D. pyriforme circinale . 32, 35, 38, 41. D. eremnum . 33, 36, 39, 42. D. sciarum . 31–36. Clypeus. 37–39. Apical part of antenna. 40–42. Metasomal tergum 1, dorsal view. Scale 1 mm.