Tonsilla yanlingensis ( Zhang, Yin & Kim, 2000 )

Zhang, Meng, Irfan, Muhammad, Wang, Lu-Yu & Zhang, Zhi-Sheng, 2022, Six new species of Tonsilla Wang & Yin, 1992 (Araneae: Agelenidae) from southern China, with the first description of the male of T. yanlingensis (Zhang, Yin & Kim, 2000), Zootaxa 5091 (2), pp. 357-372 : 367

Tonsilla yanlingensis ( Zhang, Yin & Kim, 2000)

Figures 7A–E , 9K–L , 10

Coelotes yanlingensis Zhang, Yin & Kim, 2000: 81 , figs 4–6 (♀).

Tonsilla yanlingensis: Yin et al. 2012: 1033 , figs 537a–c (♀); Zhu et al. 2017: 550, figs 364A–B (♀).

Tonsilla subyanlingensis Liu & Xu, 2020 in Liu et al. 2020: 38 , figs 4C–D, 6A–D, 7H–I, 8D–E (♀ only, misidentified).

Material examined: 2 ♂, CHINA, Jiangxi Province, Ji’an City, Jinggang Mountain, Dajing , 26.55866667°N, 114.12896667°E, 882 m, 27.X.2016, L.Y. Wang leg. ; 3 ♂ and 3 ♀ females, Dajing , 26.57593333°N, 114.14155°E, 917 m, 27.X.2016, L.Y. Wang leg.

Diagnosis. The male of T. yanlingensis can be distinguished from all congeners by the following characters: apical end of patellar apophysis bifurcated; conductor apex curved anticlockwise with bifurcated tip in retrolateral view; median apophysis arc-shaped. The female of T. yanlingensis can be distinguished from all other congeners by the atrial ridges outline almost round; copulatory ducts long indistinct; long arc-shaped spermathecae heads and spermathecae ball-shaped.

Description. Male ( Fig. 9K ). Total length 14.32. Carapace 7.57 long, 4.84 wide; opisthosoma 6.64 long, 4.21 wide. Eye sizes and interdistances: AME 0.25, ALE 0.33, PME 0.31, PLE 0.24; AME–AME 0.13, AME–ALE 0.12, PME–PME 0.17, PME–PLE 0.32, ALE–PLE 0.12. MOA 0.77 long, front width 0.59, back width 0.74. Clypeus height 0.21. Leg measurements: I 20.05 (5.36, 6.55, 4.95, 3.19); II 17.93 (4.85, 5.60, 4.45, 3.03); III 15.60 (4.37, 4.83, 4.17, 2.23); IV 19.51 (5.60, 6.41, 5.41, 2.09). Spination of legs: femur I 220, II 240, III 130, IV 220; patella I-II 000, III 020, IV 010; tibia I-II 026, III-IV 046; metatarsus I 046, II 066, III-IV 0106.

Palp ( Fig. 7A–C ). Patella short, patellar apophysis two times longer than patella, L-shaped, apical end bifurcated. Retrolateral tibial apophysis and lateral tibial apophysis distinct with a blunt end. Cymbium approximately 3 times longer than wide; cymbial furrow extensively sclerotized, 1/3 of the cymbial length. Median apophysis arc-shaped. Conductor apex curved anticlockwise with bifurcated tip in retrolateral view. Conductor’s dorsal apophysis robust, grooved. Conductor lamella distinct. Embolus filiform, originating at the 7 o’clock position.

Female ( Fig. 9L ). Total length 15.85. Carapace 7.31 long, 4.73 wide; opisthosoma 7.99 long, 5.23 wide. Eye sizes and interdistances: AME 0.18, ALE 0.33, PME 0.28, PLE 0.32; AME–AME 0.18, AME–ALE 0.20, PME– PME 0.20, PME–PLE 0.39, ALE–PLE 0.14. MOA 0.75 long, front width 0.60, back width 0.81. Clypeus height 0.29. Leg measurements: I 16.72 (4.87, 5.66, 3.77, 2.42); II 14.97 (4.30, 5.04, 3.33, 2.30); III 13.12 (3.80, 4.23, 3.04, 2.05); IV 17.60 (4.85, 6.05, 4.56, 2.14). Spination of legs: femur I 220, II 210, III 130, IV 200; patella I-II 000, III 010, IV 000; tibia I-II 026, III 046, IV 036; metatarsus I-II 066, III 0106, IV 0156.

Epigyne ( Fig. 7D, E ). Epigynal plate almost rectangular, teeth situated anteriorly. Atrial ridges smooth, outline almost round. Epigynal hoods indistinct, located ventro-mesally. Copulatory opening posteriorly situated dorsolaterally within atrium. Copulatory ducts long, indistinct. Spermathecal head long arc-shaped. Spermathecae ballshaped, separated by minimal distance.

Distribution. China (Hunan, Jiangxi) ( Fig. 10 ).

Remarks. Based on the genital characters, the male (holotype) of Tonsilla subyanlingensis Liu & Xu, 2020 resembles Draconarius carinatus ( Wang, Yin, Peng & Xie, 1990) ( Wang, 2003: 524, fig. 19C–E). Hence, we transfer the male of T. subyanlingensis to the genus Draconarius , resulting in the new combination D. subyanlingensis ( Liu & Xu, 2020) comb. nov.. The female of T. subyanlingensis is quite similar to T. yanlingensis and females were collected together with males of the latter species from Jinggang Mountain. The female of Tonsilla subyanlingensis is therefore referred to T. yanlingensis in this paper.

Figures

FIGURE 7. Tonsilla yanlingensis (Zhang, Yin & Kim, 2000). A. Male holotype, left palp, prolateral view; B. Same, ventral view; C. Same, retrolateral view; D. Epigyne, ventral view; E. Same, dorsal view. Abbreviations: At, atrium; CD, copulatory duct; CDA, Conductor’s dorsal apophysis; CF, cymbial furrow; CL, conductor lamella; Co, conductor; CO, copulatory opening; EH, epigynal hood; Em, embolus; ET, epigynal teeth; FD, fertilization ducts; LTA, lateral tibial apophysis; MA, median apophysis; PA, patellar apophysis; RTA, retrolateral tibial apophysis; SH, spermathecal heads; Sp, spermathecae.

FIGURE 9. Habitus of Tonsilla spp. A–B, male holotype and female paratype of T. jinyunensis sp. nov.; C–D, male holotype and female paratype of T. jiugongensis sp. nov.; E–F, male holotype and female paratype of T. rutunda sp. nov.; G–H, male holotype and female paratype of T. subrostrum sp. nov.; I–J, male holotype and female paratype of T. subtruculenta sp. nov.; K–L, male and female of T. yanlingensis (Zhang, Yin & Kim, 2000); M, male holotype of T. yueliangensis sp. nov.

FIGURE 10. Distribution of Tonsilla species treated here.