Hemipenthes jaennickeana (Osten Sacken)

Ávalos-Hernández, Omar, 2009, A review of the North American species of Hemipenthes Loew, 1869 (Diptera: Bombyliidae), Zootaxa 2074, pp. 1-49 : 26-28

12. Hemipenthes jaennickeana (Osten Sacken)

(Figs. 13, 55–56)

Isopenthes jaennickeana Osten Sacken, 1886a: 97 .

Anthrax jaennickeanus (Osten Sacken) : Kertész, 1909: 42

Villa (Hemipenthes) sinuosa jaennickeana (Osten Sacken) : Painter & Painter, 1962: 113 Hemipenthes sinuosa jaennickeana (Osten Sacken) : Hull, 1973: 385. Hemipenthes jaennickeana (Osten Sacken) : Evenhuis & Greathead, 1999: 445.

Lectotype male designated by Painter & Painter (1962: 113) in BMNH.

Diagnosis: Presence of a crossvein between R4 and R2+3; cell r1 entirely infuscated except for a subapical hyaline area; cell r2+3 cell with basal half infuscated with a spot at tip; r5 cell infuscated at half.

Descriptions: Male. Body length: 5–9 mm; wing length: 5–9 mm. Head: Eyes separated by one and a half width of ocellar triangle. Front black pilose, yellowish tomentose. Face brown, rounded, with black hairs and yellowish tomentum. Scape fulvous, swollen on inner apical margin, with black hairs, twice as long as pedicel; pedicel fulvous, twice as wide as long, short black hairs on the base; flagellomere brown, longer than scape and pedicel combined; base subconical, tapering to styliform apical two-thirds; stylus minute, terminal. Proboscis short, not projecting beyond oral margin. Palpi cinereous with black hairs. Occiput with short black and yellow hairs and yellowish scales.

Thorax: Mesonotum anterior margin pale yellow pilose; lateral margin black and whitish pilose; tomentum on disc entirely yellowish, long, hairlike, not dense; bristles black. Mesopleuron black and yellow pilose; tomentum on katepisternum pale yellow. Proepimeron yellowish pilose with some black hairs present. Mid coxa with black hairs, tomentum on all coxae hairlike, pale yellow. Legs fulvous, tarsi black, femora black pilose, yellow tomentose, some black scales may be present; bristles black. Halter steam brown, knob yellow. Scutellum brown, not pilose, and yellow tomentose, a spot of black tomentum in middle at base; bristles black. Black setulae on basicosta. Cells c, sc, br, bm and r1entirely infuscated, cell r1 with a hyaline spot near tip (Fig. 13); cells a and cup entirely infuscated except tip; cells r5, cua1 and dm with two basal thirds infuscated; cells r2+3 with basal half infuscated, with a spot at tip; cells m2 infuscated just at base; cell dm infuscated beyond r-m crossvein; r-m crossvein behind middle of cell dm; crossvein between R4 and R2+3 present; cell r5 slightly narrowed at wing margin; first section of vein CuA1 twice the long of r-m crossvein, second section one and a half the long of r-m crossvein, third section one and a half the long of first two sections combined; cell a slightly wider then cell cup; alula poorly developed.

Abdomen: Abdominal dorsum white pilose on first to fourth tergites, black hairs on fifth to seventh tergites; black tomentum overall; some pale yellow scales scattered on sides of abdomen; sides of abdomen with first and basal half of second tergites yellowish or white pilose, mixed abundant black with some yellowish hairs on rest. Venter white pilose, whitish tomentose. Genitalia brown with black hairs. Epandrium in lateral view, rectangular, lower margin concave in middle, basal corner narrowed; gonocoxite narrow, basal half enlarged; gonostylus small, hooked apically; epiphallus in lateral view narrow ( Fig. 55 ) slightly curved, cap-shaped, apex swollen, rounded; with a ventral extension broad at base with apex acuminate, portion of the epiphallus behind ventral extension longer than aedeagus; epiphallus in ventral view broad ( Fig. 56 ), lateral margins narrowed at both sides at middle, with scattered spines all along; aedeagus broad at base narrowed at apex, not swollen dorsally; gonopore terminal.

Female. Nearly identical to male. Eyes separated by twice width of ocellar triangle.

Distribution: Mexico (Aguascalientes, Chiapas, Coahuila, Durango, Guanajuato, Guerrero, Hidalgo, Mexico, Morelos, Nuevo Leon, Oaxaca, Puebla, Sonora), USA (Alabama, Arkansas, Arizona, California, Colorado, Florida, Idaho, Illinois, Kansas, Massachusetts, Michigan, Montana, Nebraska, Nevada, New Jersey, New Mexico, New York, Ohio, Oregon, Texas, Utah, Wyoming).

Specimens examined: Mexico. Morelos: Quilamula, 24-IV-2004 (1 female; CNIN), 25-V-2004 (1 female; CNIN), 10-VIII-2003 (1 male; MZFC), 24-II-2004 (1 male; MZFC), O. Ávalos and M. López. USA. Colorado: Jackson Co., 11-VIII-1959, N. Marston (1 male; USNM). Florida: Lake Worth, 11-XI-1925 (1 female; USNM). Kansas: Riley Co., 24-V-1964, N. Marston (1 male genitalia; USNM); Riley Co., 30-V- 1949, James B. King (1 female; USNM); Manhattan, 19-VI-1959, N. Marston (1 male; USNM); Manhattan, Riley Co., 19-VI-1962, N. Marston (1 male; USNM).

Remarks: Hemipenthes jaennickeana is similar to H. sinuosa and H. blanchardiana but can be distinguished from the first by the presence of a crossvein between R4 and R2+3 and from the second by the pigmentation of the r5 cell, infuscated just at half. The genitalia of H. jaennickeana ( Fig. 55 ) has a wider epiphallus in lateral view than H. sinuosa ( Figs. 71 ) and H. blanchardiana ( Fig. 31 ). This species has a widespread distribution in North America, as does the related species H. sinuosa .

Figures

FIGURES 55 – 56. Hemipenthes jaennickeana male terminalia. 55, lateral view; 56, ventral view.

FIGURES 71 – 72. Hemipenthes sinuosa male terminalia. 71, lateral view; 72, ventral view.

FIGURES 31 – 32. Hemipenthes blanchardiana male terminalia. 31, lateral view; 32, ventral view.