Hemipenthes edwardsii (Coquillett)

Ávalos-Hernández, Omar, 2009, A review of the North American species of Hemipenthes Loew, 1869 (Diptera: Bombyliidae), Zootaxa 2074, pp. 1-49 : 22-23

9. Hemipenthes edwardsii (Coquillett)

(Figs. 10, 49–50)

Anthrax edwardsii Coquillett, 1894b: 102 .

Villa (Hemipenthes) edwardsii (Coquillett) : Painter & Painter, 1965: 435. Hemipenthes edwardsii (Coquillett) : Hull, 1973: 385.

4 syntypes in AMNH, USA; 1 syntype in USNM.

Diagnosis: Tomentum on coxae white; cell a hyaline or infuscated just at base; first abdominal tergite with a stripe of white or yellow scales; second, third and fourth abdominal tergites with scattered, shinning fulvous and white scales.

Description: Male. Body length: 6–10 mm; wing length: 5–9 mm. Head: Eyes separated by a little more than width of ocellar triangle. Front black pilose, fulvous tomentose. Face brown, rounded, with black hairs and fulvous tomentum, denser on inner margin of eyes beneath the antenna. Scape black, swollen on inner apical margin, with black hairs, twice as long as pedicel; pedicel black, twice as wide as long, with short black hairs; flagellomere black, longer than scape and pedicel combined; base subconical, tapering to styliform apical two-thirds; stylus minute, terminal. Proboscis short, not projecting beyond oral margin. Palpi black with black hairs. Occiput with short black hairs and shining fulvous scales.

Thorax: Mesonotum anterior margin yellowish pilose; lateral margin entirely yellow pilose; tomentum on disc shining fulvous, forming three longitudinal stripes, more dense near scutellum; bristles black. Mesopleuron black pilose with yellowish hairs mixed in on proepisternum and anepisternum, katatergite with yellow hairs; tomentum on katepisternum yellow. Proepimeron with mixed black and yellowish hairs. Mid coxa with black hairs, tomentum on all coxae hairlike, yellow. Legs fulvous, femora black pilose and yellowish tomentose; bristles black. Halter steam and knob brown. Scutellum brown, black pilose, shining fulvous and brown tomentose; bristles black. Black setulae on basicosta. Cells c, sc, br, and bm entirely infuscated (Fig. 10); cell r1 with two basal thirds infuscated; cell cup with basal half infuscated; cell dm with basal third infuscated; cells r5, r2+3, a and cua1 infuscated just at base; cell dm infuscated beyond r-m crossvein; r-m crossvein at or slightly behind middle of cell dm; no crossvein between R4 and R2+3; cell r5 slightly narrowed at wing margin; first section of vein CuA1 two and a half the long of r-m crossvein, second section as long as r-m crossvein, third section twice the long of first two sections combined; cell a one and a half wider than cell cup; alula slightly developed.

Abdomen: Abdominal dorsum with whitish pile on tergite one, rest black pilose; white, shining fulvous and white-violet tomentum mixed in on tergites two to seven, darker toward apex, a crossband of white tomentum on first tergite, scales on second to fourth tergites not reaching tergite center; sides of abdomen with first and basal half of second tergites whitish pilose, mixed abundant black with some fulvous hairs on rest. Venter black pilose, shining fulvous tomentose. Genitalia black with black and yellowish hairs. Epandrium in lateral view, rectangular, lower margin straight, basal corner narrowed neck-like; gonocoxite narrow, basal half not enlarged; gonostylus small, hooked apically; epiphallus in lateral view narrow ( Fig. 49 ) slightly curved, cap-shaped, apex swollen, rounded; with a ventral extension broad at base with acuminate apex, portion of the epiphallus behind ventral extension longer than aedeagus; epiphallus in ventral view broad ( Fig. 50 ), lateral margins narrowed at both sides at base and before apex, with scattered spines in the middle and dense spines at apex; aedeagus broad at base narrowed at apex, not swollen dorsally; gonopore terminal.

Female. Nearly identical to male. Eyes separated by two and a half width of ocellar triangle. Tomentum on front and face yellowish. Tomentum on disc yellowish. Tomentum on abdominal dorsum white, fulvous and yellow, not white-violet.

Distribution: Canada (British Columbia), USA (Arizona, California, Colorado, Washington). Specimens examined: USA. California: San Francisco, 24-VII-1930, R. H. Painter (1 male, 1 genitalia; USNM); Seaside Monterrey Co., 23-V-1959, D. D. Linsdale (1 female; USNM); Pt. Reyes Marin Co., 11-IV- 1959, G. I. Stage (1 male; USNM); Pt. Reyes Marin Co., 20-IV-1962, J. Poorbaugh (2 males; USNM). Remarks: Hemipenthes edwardsii is the only species in the genus with shinning fulvous scales. This species has been collected only in western Canada and USA.

Figures

FIGURES 49 – 50. Hemipenthes edwardsii male terminalia. 49, lateral view; 50, ventral view.