Coridromius chenopoderis, Tatarnic & Cassis, 2008

Tatarnic, N. J. & Cassis, G., 2008, Revision Of The Plant Bug Genus Coridromius Signoret (Insecta: Heteroptera: Miridae), Bulletin of the American Museum of Natural History 2008 (315), pp. 1-95 : 35-44

Coridromius chenopoderis , new species Figures 2B–C, 4, 6, 7K–M, 8H, map 4

HOLOTYPE: Male: AUSTRALIA: Western Australia: 13.2 km N of jct of Agana Kalbarri Rd and Brand Hiway , Galena River Bridge , 27.82917 ° S 114.6873 ° E, 190 m, 24

Oct 2004, Cassis, Wall, Weirauch, Symonds, Atriplex amnicola Paul G. Wilson (Chenopodiaceae) , det. Perth staff PERTH6988431 ( AMNH _PBI 00015692) ( AM).

DIAGNOSIS: Recognized by the following combination of characters: proepisternum unilobed; metanotum not prominently flared; posterior margin of pygophore biconvex, with prominent fold on left side forming deep U-shaped groove, without small apophysis on right margin of groove. C. chenopoderis , monotocopsis , pilbarensis , and variegatus are all very similar and often difficult to tell apart. Of the four species, chenopoderis exhibits the most phenotypic variation in both size and coloration (e.g., fig. 2B–C) and has the widest variety of known host plants of all Coridromius . It has a somewhat narrower body shape and is typically smaller than the others, though size ranges do overlap. Males can be distinguished from the other species by the relatively shorter left paramere, the much deeper U-shaped fold on the posterior margin of the pygophore, by the lack of a small apical process on the right margin of this fold (found only in pilbarensis ) and by the lack of a small rounded lobe on the posterior margin of the metepimeron (as in fig. 14B) as found in variegatus .

DESCRIPTION: COLORATION (figs. 2B– C, 4, 6): Extremely variable, ranging from light tan or olive to brown or black with brown markings (fig. 2B–C). Head: Mostly brownish yellow to orange-brown with dark brown vittae on frons; lateral margins bordering eyes and posterior margin of vertex yellow; paired tubercles on vertex yellow, bordered posteriorly with dark brown markings, sometimes merging with dark brown spot in middle of vertex; clypeus with irregular median and lateral dark brown markings; mandibular and maxillary plates darkened along ventral margins; buccula pale; labrum light brown; labium yellow, brown apically; in some specimens head mostly dark brown. Antenna: AI yellowbrown to orange-brown with thin brown apical and basal annulations; AII yellowbrown to orange-brown, sometimes slightly brown apically; AIII and AIV orange-brown to brown. Thorax: Pronotal collar yellow with brown anterior and posterior margins, uniformly dark brown in darker specimens; pronotum light tan, olive, brown, or dark brown, often with pale medial stripe, usually with irregular dark brown or black markings on callar region; mesoscutum orange-brown to dark brown; scutellum ranging from mostly dark brown with yellow corners to mostly light brown or tan with darker brown markings; thoracic pleura light brown to dark brown, often with yellow markings; mesopleuron brown to black ventrally. Hemelytra: Light tan or olive to brown or black, sometimes with a brown chevron on corium, sometimes darker along claval commissure; membrane faint brown with darker veins. Legs: Coxae yellow to brown; pro- and mesofemora yellow to dark brown, sometimes with faint, subapical brown markings; hind femur yellow to orange-brown with 8 or 9 dark brown diagonal stripes on outer surface; tibiae yellow to brown; tarsi brown apically. Abdomen: Yellow to dark brown or black, sometimes with lateral brown markings around spiracles, males sometimes with reddish markings on genital capsule. SUR- FACE AND VESTITURE (figs. 4, 6): Head glossy and shallowly punctate; pronotum shallowly punctate; propleuron, metepimeron, scutellum, and hemelytra impunctate; head and dorsum with sparse distribution of short, simple, white, decumbent setae. STRUCTURE (figs. 4, 6): Head: Wide; frons broadly tumescent medially; paired tubercles adjacent to eyes; transverse depressions above tubercles shallow; posterior margin of head slightly rounded medially but otherwise flat. Thorax: Pronotum broad and weakly rounded, margins thinly carinate, anterolateral margins weakly upturned, posterior margin weakly medially cleft, submarginal region of humeral angles weakly excavated; proepisternum unilobed; posterior margin of metepimeron truncate; metanotum not prominent and flared. Hemelytra: Embolium thin and evenly rounded, nearly straight. Legs: Metatibial spines thick. Abdomen: When viewed laterally, posterior margin of abdominal sternite II not distinctly angular. MALE GENITALIA (figs. 7K–M, 8H): Right paramere triangular and elongate (figs. 7L–M); left paramere sickle-shaped, apex acute (figs. 7K, 8H); posterior margin of pygophore biconvex, with distinct fold on left forming a deep U-shaped groove (fig. 8H). FEMALE PARAGENITALIA: No visible external paragenitalia.

ETYMOLOGY: This name was chosen to reflect this species’ preference for chenopodiaceous plants.

HOSTS: This species is known from several plant families but is most commonly collect- ed on chenopods ( Chenopodiaceae ).

DISTRIBUTION: The most widely distribut- ed of the Australian species, chenopoderis is found throughout the country. It has also been collected in New Zealand and Norfolk Island, where it has probably been introduced (map 4). This species has also recently been discovered in North America by Michael D. Schwartz; it has been collected off chenopods in Mexico, Arizona, New Mexico, southern California, Florida, and Texas. Additionally, specimens of Coridromius have been collected from Hawaii and previously identified as variegatus . Examination of these specimens, collected from Chenopodium murale , confirms that they are in fact chenopoderis .

PARATYPES: AUSTRALIA: New South Wales: 2.9 km W of Nyngan, 31.55001 ° S 147.1553 ° E, 202 m, 18 Oct 2001, Cassis, Silveira, Wall, Einadia nutans subsp. nutans (R.Br.) A. J. Scott (Chenopodiaceae) , det.

NSW staff NSW658277, 73 (AMNH_PBI 00015526–AMNH_PBI 00015532), 3♀ (AMNH_ PBI 00012117–AMNH_PBI 00012119) Rhagodia spinescens R.Br. (Chenopodiaceae) , det. NSW staff NSW658274, 13 (AMNH_PBI 00012105), 5♀ (AMNH_PBI 00012106– AMNH_PBI 00012110) (AM). 42 km SW Bourke toward Louth, 30.33334 ° S 144.5833 ° E, 100 m, 27 Oct 1995, Schuh and Cassis, Flindersia maculosa (Lindley) Benth. (Rutaceae) , det. P. H. Weston 1996 NSW395939, 13 (AMNH_PBI 00002613), 5♀ (AMNH_PBI 00002614–AMNH_PBI 00002618) (AMNH). 9.5 km E of Balranald on Sturt Hwy, 34.702 ° S 143.615 ° E, 20 Oct 1996, Schuh and Cassis, Nitraria billardierei DC. (Zygophyllaceae) , det. Perth staff PERTH05095247, 163 (AMNH_PBI 00006354–AMNH_PBI 00006369), 7♀ (AMNH_PBI 00006370– AMNH_PBI 00006376) Nitraria billardierei DC. (Zygophyllaceae) , det. Perth staff PERTH05095247, 13 (AMNH_PBI 00123417) (AM). Atriplex nummularia omissa Aellen (Chenopodiaceae) , det. Perth staff PERTH- 05054680, 23 (AMNH_PBI 00002699, AMNH_ PBI 00002700), 1♀ (AMNH_PBI 00002701) (AMNH). Comerong Island Nature Reserve, nr. Nowra, Seven Mile Beach, on dunes, 34.882 ° S 150.735 ° E, 09 Apr 2006, C. Symonds, ( Chenopodiaceae ), 53 (AMNH_PBI 00012442–AMNH_ PBI 00012446), 3♀ (AMNH_PBI 00012447– AMNH_PBI 00012449) (AM). Myall Lakes NP, 32.57916 ° S 152.29083 ° E, 16 Nov 1996, L. Wilkie, Acacia longifolia (Mimosaceae) , 23 (AMNH_PBI 00012401, AMNH_PBI 00012402) (AM). Warrumbungle National Park, Wambelong Campground, 31.21666 ° S 149.08333 ° E, 550 m, 25 Oct 1995, Schuh and Cassis, Amyema cambagei (Blakely) Danser (Loranthaceae) , det. B.M. Wiecek 1996 NSW395934, 1♀ (AMNH_PBI 00002731) (AMNH). Northern Territory: 143.3 km NW of Bond Springs on Tanami Rd, 23.03333 ° S 132.722 ° E, 573 m, 22 Oct 2001, Cassis, Schuh, Schwartz, Silveira, Wall, Rhagodia spinescens R.Br. (Chenopodiaceae) , det. NSW staff NSW658306, 23 (AMNH_PBI 00012120, AMNH_PBI 00012121), 1♀ (AMNH_PBI 00012122) (AM). Queensland: 45 km N of Quilpie, 26.3423 ° S 144.3078 ° E, 280 m, 02 Nov 1998, Schuh, Cassis, Silveira, Muehlenbeckia florulenta Meissner (Polygonaceae) , det. NSW staff NSW427471, 23 (AMNH_PBI 00002517, AMNH_PBI 00002518), 2♀ (AMNH_PBI 00002523, AMNH_PBI 00002524) (AMNH). 82.6 km NW of Quilpie, 26.3479 ° S 143.6454 ° E, 190 m, 03 Nov 1998, Schuh, Cassis, Silveira, Frankenia sp. (Frankeniaceae) , det. NSW staff NSW427347, 1♀ (AMNH_PBI 00012386) (AM). 9 km WSW of Adavale, 25.91094 ° S 144.5143 ° E, 270 m, 02 Nov 1998, Schuh, Cassis, Silveira, Hakea leucoptera R. Br. (Proteaceae) , det. NSW staff NSW427661, 83 (AMNH_PBI 00006203– AMNH_PBI 00006210), 8♀ (AMNH_PBI 00006211–AMNH_PBI 00006214, AMNH_ PBI 00006280–AMNH_PBI 00006283) (AM). Hakea leucoptera R. Br. (Proteaceae) , det. NSW staff NSW427661, 83 (AMNH_ PBI 00003054–AMNH_PBI 00003061), 11♀ (AMNH_PBI 00002751, AMNH_PBI 00003062– AMNH_PBI 00003071) (AMNH). 9.6 km W of Mitchell, 26.49202 ° S 147.8728 ° E, 430 m, 31 Oct 1998, Schuh, Cassis, Silveira, 23 (AMNH_PBI 00012382, AMNH_PBI 00012383), 1♀ (AMNH_PBI 00012384) (AM). 91 km N of Quilpie, 25.99847 ° S 144.4098 ° E, 300 m, 02 Nov 1998, Schuh, Cassis, Silveira, Amyema quandang var. quandang (Lindley) Tieghem (Loranthaceae) , det. NSW staff NSW427341, 1♀ (AMNH_PBI 00006117) Hakea leucoptera (Proteaceae) , 133 (AMNH_PBI 00006085– AMNH_PBI 00006097), 20♀ (AMNH_PBI 00006098–AMNH_PBI 00006116, AMNH_PBI 00012387) Hakea leucoptera R. Br. (Proteaceae) , det. NSW staff NSW 427661, 13 (AMNH_PBI 00123415), 1♀ (AMNH_PBI 00123420) (AM). Hakea leucoptera R. Br. (Proteaceae) NSW 427661, 133 (AMNH_ PBI 00003175–AMNH_PBI 00003187), 27♀ (AMNH_PBI 00003188–AMNH_PBI 00003214) (AMNH). South Australia: 11.5 km NE of Wooltana Homestead, 30.33726 ° S 139.4934 ° E, 180 m, 06 Nov 1998, Schuh, Cassis, Silveira, 23 (AMNH_PBI 00012388, AMNH_PBI 00012389), 3♀ (AMNH_PBI 00012390–AMNH_PBI 00012392) (AM). 12.5 km E of Martins Well, 31.50001 ° S 139.2276 ° E, 162 m, 09 Nov 2001, Cassis, Schuh, Schwartz, Acacia oswaldii F.Muell. (Fabaceae) , det. NSW staff NSW666370, 33 (AMNH_PBI 00012111– AMNH_PBI 00012113), 5♀ (AMNH_PBI 00012114–AMNH_PBI 00012116) (AM). 15 km S of Bews, 35.48474 ° S 140.4332 ° E, 130 m, 08 Nov 1998, Schuh, Cassis, Silveira, Light Trap, 13 (AMNH_PBI 00123407) (AM). 17 km E of Nepebunna, Gammon Ranges National Park, 30.57312 ° S 139.1273 ° E, 480 m, 07 Nov 1998, Schuh, Cassis, Silveira, Rhagodia parabolica R. Br. (Chenopodiaceae) , 133 (AMNH_PBI 00006284–AMNH_ PBI 00006296), 20♀ (AMNH_PBI 00006297– AMNH_PBI 00006316) Rhagodia parabolica R. Br. (Chenopodiaceae) , 13 (AMNH_PBI 00123416) (AM). 17.6 km S of Innamincka, 27.88068 ° S 140.6712 ° E, 130 m, 06 Nov 1998, Schuh, Cassis, Silveira, Chenopodium auricomum Lindley (Chenopodiaceae) , det. NSW staff, 423 (AMNH_PBI 00006268– AMNH_PBI 00006279, AMNH_PBI 00006460– AMNH_PBI 00006474, AMNH_PBI 00012039– AMNH_PBI 00012053), 34♀ (AMNH_PBI 00006317–AMNH_PBI 00006350) Teucrium racemosum R. Br. (Lamiaceae) , det. NSW staff NSW427991, 13 (AMNH_PBI 00012055) Muehlenbeckia florulenta Meissner (Polygonaceae) , det. NSW staff NSW427467, 33 (AMNH_PBI 00012054, AMNH_PBI 00012374– AMNH_PBI 00012375), 7♀ (AMNH_PBI 00012376–AMNH_PBI 00012381, AMNH_PBI 00012393) (AM). 20 km W of Nepabunna, Mt. Serle, 30.55365 ° S 138.8304 ° E, 630 m, 07 Nov 1998, Schuh, Cassis, Silveira, 1♀ (AMNH_PBI 00012385) (AM). 41.5 km NW of Morgan, 33.63335 ° S 139.9167 ° E, 150 m, 01 Nov 1995, Schuh, Cassis, and Gross, Exocarpos aphyllus R. Br. (Santalaceae) , det. B.M. Wiecek 1996 NSW395968, 53 (AMNH_PBI 00002630– AMNH_PBI 00002634), 6♀ (AMNH_PBI 00002635–AMNH_PBI 00002640) (AMNH). 5 km N Yunta toward Arkaroola, 32.53334 ° S 139.55 ° E, 250 m, 29 Oct 1995, Schuh and Cassis, undetermined sp. ( Chenopodiaceae ), det. R.T. Schuh NSW395955, 63 (AMNH_ PBI 00002465–AMNH_PBI 00002470), 9♀ (AMNH_PBI 00002752–AMNH_PBI 00002760) (AMNH). 5.4 km N of Yunta, 32.53804 ° S 139.5598 ° E, 300 m, 08 Nov 1996, Schuh and Cassis, Acacia sp. (Fabaceae) , 43 (AMNH_ PBI 00006183–AMNH_PBI 00006186), 16♀ (AMNH_PBI 00006187–AMNH_PBI 00006202) (AM). 51 km NW of Morgan, 33.58334 ° S 140 ° E, 150 m, 01 Nov 1995, Schuh, Cassis, and Gross, Light Trap, 23 (AMNH_PBI 00012403, AMNH_PBI 00123404), 2♀ (AMNH_PBI 00123405, AMNH_PBI 00123406) (AM). Eremophila mitchellii Benth. (Myoporaceae) , det. J. Everett 1996 NSW395936, 23 (AMNH_ PBI00002713, AMNH_PBI 00002714), 2♀ (AMNH_PBI 00002715, AMNH_PBI 00002716), Light Trap, 43 (AMNH_PBI 00002709– AMNH_PBI 00002712), 2♀ (AMNH_PBI 00002641, AMNH_PBI 00002642) (AMNH). 52 km SW of Yunta, 32.83335 ° S 139.1 ° E, 500 m, 30 Oct 1995, Schuh and Cassis, 243 (AMNH_PBI 00006383–AMNH_PBI 00006406), 15♀ (AMNH_PBI 00006407–AMNH_PBI 00006421) (AM). 583 (AMNH_PBI 00002487– AMNH_PBI 00002499, AMNH_PBI 00003004– AMNH_PBI 00003017, AMNH_PBI 00003072– AMNH_PBI 00003102), 43♀ (AMNH_PBI 00002500–AMNH_PBI 00002516, AMNH_PBI 00003018–AMNH_PBI 00003035, AMNH_PBI 00003103–AMNH_PBI 00003110) (AMNH). 6 km N of Sturt Hiway, Mt. Mary Road, 34.33334 ° S 139.4167 ° E, 250 m, 01 Nov 1995, Schuh, Cassis, and Gross, Nitraria billardierei DC. (Nitrariaceae) , det. R.G. Coveny 1996 NSW395966, 33 (AMNH_PBI 00006171– AMNH_PBI 00006173), 5♀ (AMNH_PBI 00006174–AMNH_PBI 00006178) (AM). Nitraria billardierei DC. (Nitrariaceae) , det. R.G. Coveny 1996 NSW395966, 53 (AMNH_ PBI 00002619–AMNH_PBI 00002623), 6♀ (AMNH_PBI 00002624–AMNH_PBI 00002629) (AMNH). 66–69 km NW of Morgan, Cane Grass, 33.58334 ° S 140.0333 ° E, 150 m, 02 Nov 1995, Schuh, Cassis, and Gross, undetermined sp., 223 (AMNH_PBI 00002658–AMNH_PBI 00002679), 19♀ (AMNH_PBI 00002680– AMNH_PBI 00002698) (AMNH). 75 km NW of Morgan, 5 km N Cane Grass, 33.53334 ° S 140.05 ° E, 100 m, 02 Nov 1995, Schuh, Cassis, and Gross, Light Trap, 1♀ (AMNH_PBI 00123408) (AM). 96 km NW of Morgan, Pine Valley Stn, 33.31667 ° S 140.2 ° E, 150 m, 02 Nov 1995, Schuh, Cassis, and Gross, Exocarpos aphyllus R. Br. (Santalaceae) , det. B.M. Wiecek 1996 NSW395968, 1♀ (AMNH_PBI 00002702) (AMNH). Brookfield Cons. Pk., 34.22 ° S 139.27 ° E, 25 Nov 1992, I. D. Naumann & J. C. Cardale, 23 (AMNH_PBI 00006179, AMNH_PBI 00006180), 2♀ (AMNH_PBI 00006181, AMNH_PBI 00006182) (AM). Nullabor National Park 5 km E of WA border, 31.63716 ° S 129.0346 ° E, 50 m, 22 Oct 1996, Schuh and Cassis, Myoporum insulare? R. Br. (Myoporaceae) , det. Perth staff PERTH05095239, 43 (AMNH_PBI 00012394–AMNH_PBI 00012397), 2♀ (AMNH_PBI 00012398, AMNH_PBI 00012399) (AM). Scorpion Springs Cons. Park, 35.60421 ° S 140.8646 ° E, 125 m, 10 Nov 1998, Schuh, Cassis, Silveira, Melaleuca brevifolia Turcz. (Myrtaceae) , det. NSW staff NSW427362, 1♀ (AMNH_PBI 00123400) (AM). Yunta, 32.58 ° N 139.559 ° E, 12 Nov 1994, C.N. and A.S. Smithers, 1♀ (AMNH_PBI 00123409) (AM). Victoria: Little Desert National Park, 5–6 km W of McDonald Hiway, 36.61668 ° S 141.1667 ° E, 150 m, 03 Nov 1995, Schuh and Cassis, Acacia acinacea Lindl. (Fabaceae) , det. B.J. Conn 1996 NSW 395983, 1♀ (AMNH_PBI 00002643) (AMNH). Western Australia: 115.4 km E of Norseman, 32.05143 ° S 122.9675 ° E, 600 m, 23 Oct 1996, Schuh and Cassis, Eremophila ionantha Diels (Myoporaceae) , det. Perth staff PERTH05056152, 23 (AMNH_PBI 00002542, AMNH_PBI 00002543) (AMNH). 13.2 km N of jct of Agana Kalbarri Rd and Brand Hiway, Galena River Bridge, 27.82917 ° S 114.6873 ° E, 190 m, 24 Oct 2004, Cassis, Wall, Weirauch, Symonds, Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH6988431, 223 (AMNH_PBI 00015693–AMNH_PBI 00015714), 12♀ (AMNH_PBI 00015715– AMNH_PBI 00015726) (AM). 15 km S of Lake Grace, 33.2367 ° S 118.4692 ° E, 400 m, 07 Dec 1997, Schuh, Cassis, Brailovsky, Asquith, 1♀ (AMNH_PBI 00002548) (AMNH). 2 km E of Nungarin on Rt 50, 31.43596 ° S 118.2627 ° E, 330 m, 16 Nov 1999, R.T. Schuh, G. Cassis, & R. Silveira, Rhagodia preissii preissii Moq. (Chenopodiaceae) , det. Perth staff PERTH05670594, 43 (AMNH_PBI 00006450–AMNH_PBI 00006453), 6♀ (AMNH_ PBI 00006454–AMNH_PBI 00006459) (AM). 2.5 km W of Brand Hiway (off Red Emperor Rd), Flat Rocks Beach (S of Geraldton), 29.02061 ° S 114.7863 ° E, 22 Oct 2004, Cassis, Wall, Weirauch, Symonds, Rhagodia preissii subsp. obovata (Moq.) Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH6989543, 13 (AMNH_PBI 00012088), 1♀ (AMNH_PBI 00012089) (AM). 22 km S of Watheroo, 30.43161 ° S 116.0438 ° E, 400 m, 02 Nov 1996, Schuh and Cassis, Halosarcia indica bidens (Nees) Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH05236843, 53 (AMNH_PBI 00012100–AMNH_PBI 00012104), 2♀ (AMNH_PBI 00012425, AMNH_PBI 00012426) (AM). Scholtzia capitata Benth. (Myrtaceae) , det. Perth staff PERTH05236622, 83 (AMNH_PBI 00002473– AMNH_PBI 00002477, AMNH_PBI 00002601– AMNH_PBI 00002603), 11♀ (AMNH_PBI 00002478–AMNH_PBI 00002484, AMNH_PBI 00002604–AMNH_PBI 00002607) Halosarcia indica bidens (Nees) Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH05236843, 123 (AMNH_PBI 00002645–AMNH_PBI 00002656), 1♀ (AMNH_PBI 00002657) (AMNH). 27.6 km N of Coral Bay Rd on Cardabia-Ningaloo Rd, 22.90198 ° S 113.8167 ° E, 25 m, 29 Oct 2004, Cassis, Wall, Weirauch, Tatarnic, Symonds, Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Field ID, 113 (AMNH_PBI 00015622–AMNH_ PBI 00015632), 9♀ (AMNH_PBI 00015633– AMNH_PBI 00015641) (AM). 31.3 km N of Ravensthorpe, 33.36519 ° S 119.8792 ° E, 500 m, 05 Nov 1996, Schuh and Cassis, 13 (AMNH_PBI 00002540), 1♀ (AMNH_PBI 00002541) (AMNH). 31.7 km W of Agnew toward Sandstone, 27.96227 ° S 120.4277 ° E, 800 m, 26 Oct 1996, Schuh and Cassis, Rhagodia preissii Moq. (Chenopodiaceae) , det. Perth staff PERTH05054753, 1♀ (AMNH_PBI 00002526) (AMNH). 39 km NE of King River, 34.78937 ° S 118.2759 ° E, 200 m, 06 Nov 1996, Schuh and Cassis, Agonis parviceps Schauer (Myrtaceae) , det. Perth staff PERTH05236827, 13 (AMNH_ PBI 00002522) (AMNH). 40 km N of Ravensthorpe, 33.31524 ° S 119.8151 ° E, 500 m, 05 Nov 1996, Schuh and Cassis, Maireana oppositifolia (F. Muell.) Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH- 05236630, 73 (AMNH_PBI 00006129– AMNH_PBI 00006135), 18♀ (AMNH_PBI 00006136–AMNH_PBI 00006153) (AM). 42.5 km N of Kojonup, 33.46391 ° S 117.0579 ° E, 400 m, 04 Nov 1996, Schuh and Cassis, Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH- 05236835, 23 (AMNH_PBI 00012421, AMNH_PBI 00012422), 13♀ (AMNH_PBI 00015773–AMNH_PBI 00015785) (AM). Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH05236835, 63 (AMNH_PBI 00002581–AMNH_PBI 00002586), 6♀ (AMNH_PBI 00002587– AMNH_PBI 00002592) (AMNH). Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH 05236835, 43 (AMNH_PBI 00191191–AMNH_PBI 00191194), 4♀ (AMNH_PBI 00191195–AMNH_PBI 00191198) (USNM). 43 km N of Norseman, 31.85648 ° S 121.6414 ° E, 300 m, 19 Nov 1999, R.T. Schuh, G. Cassis, & R. Silveira, Olearia muelleri (Sond.) Benth. (Asteraceae) , det. Perth staff PERTH05670691, 83 (AMNH_ PBI 00002748, AMNH_PBI 00003036– AMNH_PBI 00003042), 13♀ (AMNH_PBI 00002749–AMNH_PBI 00002750, AMNH_ PBI 00003043–AMNH_PBI00003053) (AMNH). Atriplex bunburyana F. Muell. (Chenopodiaceae) , det. Perth staff PERTH05671205, 23 (AMNH_PBI 00114226, AMNH_PBI 00114227) (CNC). 46.5 km W of Yalgoo, 28.41302 ° S 116.2151 ° E, 600 m, 27 Oct 1996, Schuh and Cassis, Eremophila forrestii F. Muell. (Myoporaceae) , det. Perth staff PERTH05671205, 1♀ (AMNH_PBI 00012424) (AM). 51 km N of Leeman on Indian Ocean Rd, Beekeepers Nature Reserve, 29.5116 ° S 114.997 ° E, 06 Nov 2004, Cassis, Weirauch, Tatarnic, Symonds, Rhagodia preissii Moq. (Chenopodiaceae) , det. Field ID, 13 (AMNH_PBI 00012096), 1♀ (AMNH_PBI 00012097) (AM). Rhagodia preissii Moq. (Chenopodiaceae) , det. Field ID, 63 (AMNH_PBI 00114217–AMNH_PBI 00114222), 3♀ (AMNH_ PBI 00114223–AMNH_PBI 00114225) (CNC). 53.9 km N of Kalgoorlie, 30.28882 ° S 121.2558 ° E, 600 m, 24 Oct 1996, Schuh and Cassis, 43 (AMNH_PBI 00002717–AMNH_PBI 00002720), 1♀ (AMNH_PBI 00002721) (AMNH). 56.6 km W of Yalgoo, 28.42397 ° S 116.1233 ° E, 600 m, 27 Oct 1996, Schuh and Cassis, Melaleuca uncinata R.Br. (Myrtaceae) , det. Perth staff PERTH05120640, 2♀ (AMNH_PBI 00002520, AMNH_PBI 00002521) Dicrastylis fulva forma fulva Harv. (Chloanthaceae) , det. Perth staff PERTH05120632, 13 (AMNH_PBI 00002519) (AMNH). 82.2 km W of Agnew toward Sandstone, 28.01057 ° S 119.9455 ° E, 650 m, 26 Oct 1996, Schuh and Cassis, Alyogyne pinoniana (Gaudich.) Fryxell (Malvaceae) , det. Perth staff PERTH- 05056101, 13 (AMNH_PBI 00123418), 1♀ (AMNH_PBI 00123419) (AM). Blowholes Rd NW of North West Coastal Hiway, Beagle Hill Area, 24.49068 ° S 113.4626 ° E, 20 m, 27 Oct 2004, Cassis, Wall, Weirauch, Tatarnic, Symonds, Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Field ID, 13 (AMNH_PBI 00012086), 1♀ (AMNH_ PBI 00012087) Enchylaena tomentosa var. tomentosa R.Br. (Chenopodiaceae) , det. Perth staff PERTH6988903, 33 (AMNH_PBI 00015490–AMNH_PBI 00015492) (AM). Brand Hiway 55.9 km S of Dongarra Road, 29.62934 ° S 115.2187 ° E, 100 m, 31 Oct 1996, Schuh and Cassis, Scholtzia drummondii Benth. (Myrtaceae) , det. Perth staff PERTH- 05120209, 23 (AMNH_PBI 00002544, AMNH_ PBI 00002545) (AMNH). Burrup Conservation Area, near Karratha, 20.57805 ° S 116.79472 ° E, 17 m, 03 Jun 2004, N. Tatarnic and S. Lassau, Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH6988431, 33 (AMNH_PBI 00015727–AMNH_PBI 00015729), 6♀ (AMNH_PBI 00015730– AMNH_PBI 00015735) (AM). Cape Arid National Park, Thomas River Campground, 33.8539 ° S 123.0126 ° E, 20 m, 24 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Pimelea ferruginea Labill. (Thymelaeaceae) , det. Perth staff05672341, 13 (AMNH_PBI 00002546) Atriplex isatidea Moq. (Chenopodiaceae) , det. Perth staff PERTH05672317, 33 (AMNH_PBI 00002740–AMNH_PBI 00002742), 1♀ (AMNH_PBI 00002744) (AMNH). Cape Naturaliste National Park, 33.54034 ° S 115.0123 ° E, 50 m, 14 Dec 1997, Schuh, Cassis, Brailovsky, Rhagodia baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH05055318, 33 (AMNH_PBI 00006422–AMNH_PBI 00006424), 10♀ (AMNH_ PBI 00006422–AMNH_PBI 00006431) (AM). Rhagodia baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH05055318, 283 (AMNH_PBI 00003124–AMNH_PBI 00003151), 23♀ (AMNH_PBI 00003152– AMNH_PBI 00003174) (AMNH). Carnarvon, Small Boat Harbour, 24.90044 ° S 113.6477 ° E, 26 Oct 2004, Cassis, Wall, Weirauch, Tatarnic, Symonds, Chenopodium murale L. ( Chenopodiaceae ), det. Perth staff PERTH6989047, 13 (AMNH_PBI 00012090), 1♀ (AMNH_PBI 00012091) (AM). Conspicuous Beach, Walpole-Nornalup National Park, 10 km E of Nornalup, 35.03725 ° S 116.8443 ° E, 30 m, 17 Dec 1997, Schuh, Cassis, Brailovsky, Rhagodia baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH- 05095271, 93 (AMNH_PBI 00006154– AMNH_PBI 00006162), 5♀ (AMNH_PBI 00006163–AMNH_PBI 00006167) (AM). 2♀ (AMNH_PBI 00002725, AMNH_PBI 00002726) Pimelea rosea R.Br. (Thymelaea- ceae), det. Perth staff PERTH05099781, 1♀ (AMNH_PBI 00002727) Olearia axillaris (DC.) Benth. (Asteraceae) , det. Perth staff PERTH05095328, 23 (AMNH_PBI 00002722, AMNH_PBI 00002723), 1♀ (AMNH_PBI 00002724) (AMNH). Cosy Corner Beach East, Torbay Sound, W of Albany, 35.06033 ° S 117.6446 ° E, 2 m, 01 Dec 1999, R.T. Schuh, G. Cassis, and R. Silveira, Allocasuarina humilis (Otto and F. Dietr.) L.A.S. Johnson (Casuarinaceae) , det. Perth staff PERTH05671345, 1♀ (AMNH_PBI 00002547) (AMNH). Fitzgerald River National Park, East Mylies Beach, 33.93293 ° S 120.0031 ° E, 30 m, 06 Dec 1997, Schuh, Cassis, Brailovsky, Asquith, Pimelea ferruginea Labill. (Thymelaeaceae) , det. Perth staff PERTH05056403, 1♀ (AMNH_PBI 00002525) (AMNH). Fowlers Camp (at end of Fowlers Camp Rd), Shark Bay World Heritage Area, 26.10801 ° S 113.6167 ° E, 24 Oct 2004, Cassis, Wall, Weirauch, Symonds, Chenopodium murale L. ( Chenopodiaceae ), det. Perth staff PERTH- 6988016, 703 (AMNH_PBI 00015796– AMNH_PBI 00015865), 35♀ (AMNH_PBI 00015866–AMNH_PBI 00015900) Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Perth staff PERTH6988024, 13 (AMNH_PBI 00012093), 1♀ (AMNH_PBI 00012094) (AM). Gladstone, 25.95428 ° S 114.2464 ° E, 04 Nov 2004, Cassis, Weirauch, Tatarnic, Symonds, Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Field ID, 173 (AMNH_PBI 00015593–AMNH_PBI 00015609), 12♀ (AMNH_PBI 00015610– AMNH_PBI 00015621) (AM). Greenough River Mouth, 28.86304 ° S 114.6343 ° E, 05 Nov 2004, Cassis, Weirauch, Tatarnic, Symonds, Rhagodia preissii subsp. obovata (Moq.) Paul G. Wilson (Chenopodiaceae) , det. Perth staff PERTH6988377, 73 (AMNH_PBI 00015493– AMNH_PBI 00015499), 11♀ (AMNH_PBI 00015500–AMNH_PBI 00015510), 4 juveniles (AMNH_PBI 00015511–AMNH_PBI 00015514) (AM). Hammersley Inlet Campground, 33.95788 ° S 119.9146 ° E, 2 m, 28 Nov 1999, R.T. Schuh and G. Cassis, Suaeda australis (R.Br.) Moq. (Chenopodiaceae) , det. Perth staff PERTH05671213, 73 (AMNH_PBI 00006432–AMNH_PBI 00006438), 11♀ (AMNH_ PBI 00006439–AMNH_PBI 00006449) (AM). Leeuwin Naturaliste National Park, Canal Rocks, 33.66513 ° S 115.0165 ° E, 50 m, 15 Dec 1997, Schuh, Cassis, Brailovsky, Asquith, 13 (AMNH_PBI 00002485), 1♀ (AMNH_PBI 00002486) (AMNH). Mosman Park, Perth, 32.0209 ° S 115.7687 ° E, 20 m, 29 Nov 1998, G. Cassis, Rhagodia baccata (Chenopodiaceae) , 43 (AMNH_PBI 00006118–AMNH_PBI 00006121), 3♀ (AMNH_PBI 00006122– AMNH_PBI 00006124) (AM); 05 Nov 1998, G. Cassis, Rhagodia baccata baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH 05227283, 123 (AMNH_PBI 00015902–AMNH_PBI 00015913), 10♀ (AMNH_ PBI 00015914–AMNH_PBI 00015923) (AM). Peak Charles National Park Campground, 32.88335 ° S 121.1703 ° E, 300 m, 20 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Rhagodia preissii preissii Moq. (Chenopodiaceae) , det. Perth staff PERTH05670713, 163 (AMNH_PBI 00012056– AMNH_PBI 00012071), 14♀ (AMNH_ PBI00012072–AMNH_PBI 00012085) Melaleuca uncinata R.Br. (Myrtaceae) , det. Perth staff PERTH 05670721, 13 (AMNH_PBI 00012351) Rhagodia preissii preissii Moq. (Chenopodiaceae) , det. Perth staff PERTH- 05670713, 63 (AMNH_PBI 00015761–AMNH_ PBI 00015766), 4♀ (AMNH_PBI 00015769– AMNH_PBI 00015772) (AM). Pilbara Region: Hamersley Station, Railway Rd. just past Cooks Bore to Ridge paddock, 22.30166 ° S 117.69305 ° E, 30 Sep 2004 – 05 Oct 2004, C.V.M. volunteers, 23 (AMNH_PBI 00123410, AMNH_PBI 00123411), 3♀ (AMNH_PBI 00123412– AMNH_PBI 00123414) (AM). Poison Creek Beach, Cape Arid National Park, 33.90416 ° S 123.3519 ° E, 2 m, 25 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Atriplex isatidea Moq. (Chenopodiaceae) PERTH05672317, 173 (AMNH_PBI 00015648–AMNH_PBI 00015664), 12♀ (AMNH_PBI 00015665– AMNH_PBI 00015676) Atriplex isatidea subsp. tragus Moq. (Chenopodiaceae) PERTH05672317, 173 (AMNH_PBI 00015648– AMNH_PBI 00015664), 12♀ (AMNH_PBI 00015665–AMNH_PBI 00015676) (AM). Atriplex isatidea Moq. (Chenopodiaceae) , det. Perth staff PERTH05672317, 33 (AMNH_ PBI 00002732–AMNH_PBI 00002734), 4♀ (AMNH_PBI 00002705–AMNH_PBI 00002708) (AMNH). Rossiter Bay, Cape Le Grande National Park, 33.96726 ° S 122.2674 ° E, 3 m, 23 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Atriplex isatidea Moq. (Chenopodiaceae) , det. Perth staff PERTH05672317, 13 (AMNH_PBI 00002736), 2♀ (AMNH_PBI 00002737, AMNH_PBI 00002738) (AMNH). Stokes Inlet, Stokes National Park, 33.8179 ° S 121.1506 ° E, 2 m, 26 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Rhagodia baccata baccata (Labill.)Moq. (Chenopodiaceae) , det. Perth staff PERTH05671922, 193 (AMNH_ PBI 00015543–AMNH_PBI 00015561) (AM). Rhagodia baccata baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH- 05671922, 63 (AMNH_PBI 00003111– AMNH_PBI 00003116), 7♀ (AMNH_PBI 00003117–AMNH_PBI 00003123) (AMNH). Useless Loop Rd ca. 20 km W of jct with Shark Bay Rd, 26.56331 ° S 113.9338 ° E, 30 m, 25 Oct 2004, Cassis, Wall, Weirauch, Symonds, Rhagodia latifolia subsp. latifolia (Benth.) Paul G. Wilson (Chenopodiaceae) , det. Perth staff PERTH6987605, 33 (AMNH_ PBI 00015515–AMNH_PBI 00015517), 8♀ (AMNH_PBI 00015518–AMNH_PBI 00015525) (AM). Yalgorup National Park, near Martins Tank Campground, 32.84222 ° S 115.66222 ° E, 15 m, 14 Dec 1997, Schuh, Cassis, Brailovsky, Rhagodia sp. (Chenopodiaceae) , det. Perth staff PERTH05054990, 23 (AMNH_PBI 00006125, AMNH_PBI 00006126), 2♀ (AMNH_ PBI 00006127, AMNH_PBI 00006128) (AM). Rhagodia sp. (Chenopodiaceae) , det. Perth staff PERTH05054990, 23 (AMNH_PBI 00002728, AMNH_PBI 00002729), 1♀ (AMNH_PBI 00002730) (AMNH).

OTHER SPECIMENS EXAMINED: AUSTRA- LIA: Australian Capital Territory: Weise’s Orchard, Pialligo, 35.317 ° S 149.183 ° E, 18 Oct 1968, D. Morgan, Malus sp. (Rosaceae) , 53 (AMNH_PBI 00033216–AMNH_PBI 00033220), 11♀ (AMNH_PBI 00033221– AMNH_PBI 00033231) (ANIC). New South Wales: 32.2 km SSW of Bourke on banks of Darling River, 35.3 ° S 147.9 ° E, 26 Dec 1973, G. F. Gross, Light Trap, 73 (AMNH_PBI 00038546–AMNH_PBI 00038551, AMNH_ PBI 00038576), 6♀ (AMNH_PBI 00038552– AMNH_PBI 00038556, AMNH_PBI 00038577) (SAMA). 9.5 km E of Balranald on Sturt Hwy, 34.702 ° S 143.615 ° E, 20 Oct 1996, Schuh and Cassis, Nitraria billardierei DC. (Zygophyllaceae) , det. Perth staff PERTH05095247, 6 juveniles (AMNH_PBI 00006377–AMNH_ PBI 00006382) (AM). Cowra, 33.819 ° S 148.658 ° E, 25 Mar 1960, M. I. Nikitin, Light Trap, 13 (AMNH_PBI 00192756), 1♀ (AMNH_PBI 00192757) (BMNH). Leeton, 34.549 ° S 146.401 ° E, 27 Sep 1968, E. L. Jones, 4♀ (AMNH_PBI 00038584, AMNH_PBI 00038585) (SAMA). Murwillumbah, 28.33 ° S 153.383 ° E, 08 Apr 1919, F. Muir, 13 (AMNH_PBI 00042182) (BPBM). The Rock Nature Reserve, 35.265 ° S 147.068 ° E, 14 Sep 1968, Key, Upton and Balderson, 1♀ (AMNH_PBI 00033243) (ANIC). Northern Territory: 22 miles S of Alice Springs, 24.3 ° S 134.6 ° E, 15 Feb 1996, J. A. Grant, 33 (AMNH_PBI 00192758–AMNH_PBI 00192760), 2♀ (AMNH_PBI 00192761, AMNH_PBI 00192762) (BMNH). Alice Springs, 23.699 ° S 133.881 ° E, 16 Feb 1966, J. A. Grant, 163 (AMNH_PBI 00015737–AMNH_PBI 00015752), 8♀ (AMNH_PBI 00015753–AMNH_PBI 00015760) (AM); 17 Feb 1996, J. A. Grant, 33 (AMNH_PBI 00192763–AMNH_PBI 00192765) (BMNH); 14 Feb 1996, J. A. Grant, 1♀ (AMNH_PBI 00192766) (BMNH). Queensland: 91 km N of Quilpie, 25.99847 ° S 144.4098 ° E, 300 m, 02 Nov 1998, Schuh, Cassis, Silveira, Hakea leucoptera R. Br. (Proteaceae) , det. NSW staff NSW427661, 1 adult (AMNH_PBI 00002703) (AMNH). Dalby, 27.17994 ° S 151.27176 ° E, F. H. Hobler, 23 (AMNH_PBI 00038578, AMNH_PBI 00038579) (SAMA). Sandgate, 27.317 ° S 153.064 ° E, 09 Nov 1919, F. Muir, 1♀ (AMNH_PBI 00002746) (AMNH). South Australia: 17.6 km S of Innamincka, 27.88068 ° S 140.6712 ° E, 130 m, 06 Nov 1998, Schuh, Cassis, Silveira, Chenopodium auricomum Lindley (Chenopodiaceae) , det. NSW staff, 3 juveniles (AMNH_PBI 00006351– AMNH_PBI 00006353) (AM). 5 km SW of Whyalla, 33.05085 ° S 137.5004 ° E, 30 m, 21 Oct 1996, Schuh and Cassis, Exocarpos aphyllus R.Br. (Santalaceae) , det. Perth staff PERTH 05056209, 63 (AMNH_PBI 00012352– AMNH_PBI 00012357) (AM). Eyre Peninsula: 35 km N of Streaky Bay, 32.583 ° S 134.128 ° E, 20 m, 13 Aug 1978, W. C. Gagne, ( Chenopodiaceae ), 53 (AMNH_PBI 00042173– AMNH_PBI 00042177), 3♀ (AMNH_PBI 00042178–AMNH_PBI 00042180) (BPBM). Lyup, near Berri, 34.283 ° S 140.603 ° E, 28 Oct 1968, N. Richardson, Prunus sp. (Rosaceae) , 83 (AMNH_PBI 00033232–AMNH_PBI 00033239), 1♀ (AMNH_PBI 00033240) (ANIC). Prunus sp. (Rosaceae) , 23 (AMNH_ PBI 00038574, AMNH_PBI 00038575) (SAMA). Mount Crawford State Forest, 34.7122 ° S 138.9469 ° E, 06 May 1968, D. Colless and Z. Liepa, 2♀ (AMNH_PBI 00033241, AMNH_PBI 00033242) (ANIC). Mount Lofty, 34.974 ° S 138.709 ° E, unknown, 3♀ (AMNH_PBI 00038580, AMNH_PBI 00038581) (SAMA). Owieandana, Flinders Ranges, 30.446 ° S 138.948 ° E, Nov 1924, Hale & Tindale, 43 (AMNH_PBI 00038570, AMNH_PBI 00038573), 2♀ (AMNH_PBI 00038571, AMNH_PBI 00038572) (SAMA). near Victory Well, Everard Pk. Stn, 27.054 ° S 132.506 ° E, 03 Nov 1970, G. Gross, Light Trap, 13 (AMNH_PBI 00038568), 1♀ (AMNH_PBI 00038569) (SAMA); 29 Oct 1970, E.G. Matthews, Light Trap, 23 (AMNH_PBI 00038563, AMNH_PBI 00038564), 2♀ (AMNH_PBI 00038565, AMNH_PBI 00038566) (SAMA). Victoria: 8 km W of Kaniva, 36.377 ° S 141.241 ° E, 21 Oct 1963, J. Sedlacek, 1♀ (AMNH_PBI 00042181) (BPBM). Western Australia: 11 mi. SW of Cocklebiddy, 32.45 ° S 126.03 ° E, 22 Nov 1969, Britton, Taylor and Upton, Light Trap, 13 (AMNH_PBI 00038557), 6♀ (AMNH_PBI 00038558–AMNH_PBI 00038562, AMNH_ PBI 00038567) (SAMA). 13.2 km N of jct of Agana Kalbarri Rd and Brand Hiway, Galena River Bridge, 27.82917 ° S 114.6873 ° E, 190 m, 24 Oct 2004, Cassis, Wall, Weirauch, Symonds, Atriplex amnicola Paul G.Wilson (Chenopodiaceae) , det. Perth staff PERTH6988431, 1 juvenile (AMNH_PBI 00012350) (AM). 27.6 km N of Coral Bay Rd on Cardabia- Ningaloo Rd, 22.90198 ° S 113.8167 ° E, 25 m, 29 Oct 2004, Cassis, Wall, Weirauch, Tatarnic, Symonds, Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Field ID, 6 juveniles (AMNH_PBI 00015642–AMNH_PBI 00015647) (AM). 43 km N of Norseman, 31.85648 ° S 121.6414 ° E, 300 m, 19 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Olearia muelleri (Sond.) Benth. (Asteraceae) , det. Perth staff PERTH05670691, 1 juvenile (AMNH_PBI 00002612) (AMNH). Cape Arid National Park, Thomas River Campground, 33.8539 ° S 123.0126 ° E, 20 m, 24 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Atriplex isatidea Moq. (Chenopodiaceae) , det. Perth staff PERTH- 05672317, 1 juvenile (AMNH_PBI 00002743) (AMNH). Carnarvon, Small Boat Harbour, 24.90044 ° S 113.6477 ° E, 26 Oct 2004, Cassis, Wall, Weirauch, Tatarnic, Symonds, Cheno- podium murale L. ( Chenopodiaceae ), det. Perth staff PERTH6989047, 1 juvenile (AMNH_PBI 00012092) (AM). Conspicuous Beach, Walpole-Nornalup National Park, 10 km E of Nornalup, 35.03725 ° S 116.8443 ° E, 30 m, 17 Dec 1997, Schuh, Cassis, Brailovsky, Rhagodia baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH05095271, 3 juveniles (AMNH_PBI 00006168–AMNH_ PBI 00006170) (AM). Fowlers Camp (at end of Fowlers Camp Rd), Shark Bay World Heritage Area, 26.10801 ° S 113.6167 ° E, 24 Oct 2004, Cassis, Wall, Weirauch, Symonds, Chenopodium murale L. ( Chenopodiaceae ), det. Perth staff PERTH6988016, 1 juvenile (AMNH_PBI 00015901) Salsola tragus subsp. tragus L. ( Chenopodiaceae ), det. Perth staff PERTH6988024, 1 juvenile (AMNH_PBI 00012095) (AM). Mosman Park, Perth, 32.0209 ° S 115.7687 ° E, 20 m, 05 Nov 1998, G. Cassis, Rhagodia baccata baccata (Labill.) Moq. (Chenopodiaceae) , det. Perth staff PERTH05227283, 1♀ (AMNH_PBI 00015924) (AM). Peak Charles National Park Campground, 32.88335 ° S 121.1703 ° E, 300 m, 20 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Rhagodia preissii preissii Moq. (Chenopodiaceae) , det. Perth staff PERTH05670713, 1 juvenile (AMNH_PBI 00012423) (AM). Poison Creek Beach, Cape Arid National Park, 33.90416 ° S 123.3519 ° E, 2 m, 25 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Atriplex isatidea subsp. tragus Moq. (Chenopodiaceae) PERTH- 05672317, 15 juveniles (AMNH_PBI 00015677– AMNH_PBI 00015691) (AM). Rossiter Bay, Cape Le Grande National Park, 33.96726 ° S 122.2674 ° E, 3 m, 23 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Atriplex isatidea Moq. (Chenopodiaceae) , det. Perth staff PERTH- 05672317, 1 juvenile (AMNH_PBI 00002739) (AMNH). Stokes Inlet, Stokes National Park, 33.8179 ° S 121.1506 ° E, 2 m, 26 Nov 1999, R.T. Schuh, G. Cassis, and R. Silveira, Rhagodia baccata baccata (Labill.) Moq. (Chenopodiaceae) ,det. Perth staff PERTH- 05671922, 5 juveniles (AMNH_PBI 00015562– AMNH_PBI 00015566) (AM). MEXICO: Baja California Norte: Bahia de los Angeles, Llave Island, site 5, 28.99869 ° N 113.52048 ° W, 20 m, 11 Mar 1998, F. Sanchez Pinero, 13 (AMNH_PBI 00114164) (CNC). Bahia de los Angeles, Los Gemelos West Island, 28.95548 ° N 113.48144 ° W, 3 m, 06 Mar

1998, F. Sanchez Pinero, 543 (AMNH_PBI 00114146–AMNH_PBI 00114163, AMNH_ PBI 00114170–AMNH_PBI 00114177, AMNH_ PBI 00114181–AMNH_PBI 00114186, AMNH_ PBI 00114192–AMNH_PBI 00114206, AMNH_ PBI 00114210–AMNH_PBI 00114216), 18♀ (AMNH_PBI 00114143–AMNH_PBI 00114145, AMNH_PBI 00114165–AMNH_PBI 00114168, AMNH_PBI 00114178–AMNH_PBI 00114180, AMNH_PBI 00114187–AMNH_PBI 00114191, AMNH_PBI 00114207–AMNH_PBI 00114209), 1 juvenile (AMNH_PBI 00114169) (CNC). Chihuahua: Ruinas de Paquime, SW of Nuevo Casas Grande, 30.36888 ° N 107.94777 ° W, 1492 m, 26 Aug 1998, M. D. Schwartz, Chenopodium sp. (Chenopodiaceae) , 313 (AMNH_PBI 00114082–AMNH_PBI 00114085, AMNH_PBI 00114088–AMNH_PBI 00114114), 30♀ (AMNH_PBI 00114086–AMNH_PBI 00114087, AMNH_PBI 00114115–AMNH_ PBI 00114142) (CNC). Sonora: 11 km E of Rosario de Tesopaco on road to La Estrella, 27.83247 ° N 109.2528 ° W, 479 m, 03 May 2005, Schwartz, Weirauch, Cervantes, 13 (AMNH_PBI 00108489), 1♀ (AMNH_PBI 00108490) (AMNH). NEW ZEALAND: Hawke’s Bay: Napier, 39.5043 ° S 176.8924 ° E, 06 Apr 1975, W.J. Knight, 163 (AMNH_PBI 00192767–AMNH_PBI 00192782), 15♀ (AMNH_ PBI 00192783–AMNH_PBI 00192797) (BMNH). Marlborough: Kaikoura Peninsula, 42.4355 ° S 173.7102 ° E, 26 Jan 2000 – 28 Jan 2000, D.K. McAlpine and R. Meier, 23 (AMNH_PBI 00012439, AMNH_PBI 00012440) (AM). North Auckland: Days Bay, 36.8128 ° S 175.1656 ° E, 01 Feb 1951, T. E. Woodward, Lepidium oleraceum Forst. (Cruciferae) , 43 (AMNH_PBI 00192748–AMNH_PBI 00192751), 3♀ (AMNH_PBI 00192752– AMNH_PBI 00192754) (BMNH). NOR- FOLK ISLAND: none, 29.033 ° S 167.949 ° E, Mar 1971, K.L.S. Harley, 1♀ (AMNH_PBI 00033251) (ANIC); A. M. Lea, 1♀ (AMNH_ PBI 00038582) (SAMA). USA: Arizona: Cochise Co. : Chiricahua Mountains, 1 mi N of Rustler Park, 31.91472 ° N 109.26861 ° W, 2536 m, 18 Aug 2000 – 21 Aug 2000, J. B. Woolley, 13 (AMNH_PBI 00184915) (TAMU). Rt. 191, ca. 5 km N of Cochise , 32.16 ° N 109.94166 ° W, 20 Jun 2005, A. G. Wheeler, Jr., Atriplex canescens (Chenopodiaceae) , 83 (AMNH_PBI 00191166–AMNH_ PBI 00191173), 6♀ (AMNH_PBI 00191174– AMNH_PBI 00191179) (USNM). ca. 12 km N of Benson, 32.05833 ° N 110.34166 ° W, 21 Jun 2005, A. G. Wheeler, Jr., Atriplex canescens (Chenopodiaceae) , 13 (AMNH_PBI 00191180), 2♀ (AMNH_PBI 00191189, AMNH_PBI 00191190) (USNM). Navajo Co.: Holbrook, 34.905 ° N 110.185 ° W, 19 Jun 2005, A. G. Wheeler, Jr., 1♀ (AMNH_PBI 00191181) (USNM). California: Los Angeles Co.: Intercepted from Australia: Los Angeles, 33.94517 ° N 118.40266 ° W, 31 m, 06 Oct 1998, Unknown, Delphinium sp. (Ranunculaceae) , 2♀ (AMNH_PBI 00133761, AMNH_ PBI 00133762) (USNM). Orange Co.: Trampas Canyon at San Juan Creek, 33.51555 ° N 117.57888 ° W, 04 Sep 1999, Yanega and Gates, 1♀ (AMNH_PBI 00119981) (UCR). Riverside Co.: Riverside, 33.95333 ° N 117.39528 ° W, 25 Mar 2003, K. W. Cooper, 1♀ (AMNH_PBI 00133750) (USNM); 07 Aug 2001, K. W. Cooper, 1♀ (AMNH_PBI 00133746) (USNM); 10 Aug 2002, K. W. Cooper, 1♀ (AMNH_PBI 00133748) (USNM); 24 Aug 1999, K. W. Cooper, 1♀ (AMNH_PBI 00133749) (USNM); 14 Oct 2002, K. W. Cooper, 13 (AMNH_PBI 00133747) (USNM). San Bernardino Co.: Pinyon Hills, 34.39722 ° N 117.60944 ° W, 1447 m, 31 Jul 2003, G. R. Ballmer and D. Powell, 1♀ (AMNH_PBI 00119982) (UCR). San Diego Co.: Intercepted from Mexico: San Diego, 32.68787 ° N 117.13418 ° W, 2 m, 11 May 2000, unknown, Atriplex sp. (Chenopodiaceae) , 1♀ (AMNH_PBI 00133754) (USNM); 13 Aug 1998, unknown, Limonium sinuatum (L.) P. Mill. ( Plumbaginaceae ), 13 (AMNH_PBI 00133755), 1 juvenile (AMNH_ PBI 00133756) (USNM); 20 Mar 1998, unknown, Limonium sinuatum (L.) P. Mill. ( Plumbaginaceae ), 13 (AMNH_PBI 00133757) (USNM); 19 Mar 1998, unknown, Limonium sinuatum (L.) P. Mill. ( Plumbaginaceae ), 1♀ (AMNH_ PBI 00133758) (USNM); 06 Nov 1996, unknown, Celosia sp. (Amaranthaceae) , 13 (AMNH_PBI 00133759) (USNM); 18 Sep 2001, unknown, Amaranthus caudatus (L.) ( Amaranthaceae ), 1♀ (AMNH_PBI 00133760) (USNM). Florida: Broward Co.: Intercepted from Mexico: Fort Lauderdale [IPRL, USDA], 26.08128 ° N 80.23768 ° W, 3 m, 21 May 1997, unknown, Atriplex sp. (Chenopodiaceae) , 3♀ (AMNH_PBI 00133751– AMNH_PBI 00133753) (USNM). Hawaii: French Frigate Shoals: Tern Island, 23.87 ° N 166.28472 ° W, 25 Sep 2000, G.M. Nishida, 13 (AMNH_PBI 00042602) Chenopodium murale (Chenopodiaceae) , 23 (AMNH_PBI 00042599, AMNH_PBI 00042600), 1♀ (AMNH_PBI 00042601) (BPBM). New Mexico: Otero Co.: Alamogordo, 32.88 ° N 105.96166 ° W, 22 Jun 2005, A. G. Wheeler, Jr., Atriplex canescens (Chenopodiaceae) , 3♀ (AMNH_PBI 00191182–AMNH_PBI 00191184) (USNM). Texas: Brewster Co.: Big Bend National Park, Rio Grande nr Boquillas, 29.19583 ° N 102.93194 ° W, 576 m, 28 Apr 2001, D. E. Baumgardner, 4♀ (AMNH_PBI 00184911–AMNH_PBI 00184914) (TAMU). Cameron Co.: Jct. FM 509 & US 281, 26.052 ° N 97.73583 ° W, 19 Sep 2006, T. J. Henry and R. Coleman, 13 (AMNH_PBI 00191185), 3♀ (AMNH_PBI 00191186– AMNH_PBI 00191188) (USNM).