Ancylorhynchus cruciger ( Loew, 1858 )

Londt, Jason G. H., 2011, A review of Afrotropical Ancylorhynchus Berthold, 1827 (Diptera: Asilidae: Stenopogoninae), African Invertebrates 52 (2), pp. 471-471 : 479-484

publication ID

https://doi.org/ 10.5733/afin.052.0214

persistent identifier

https://treatment.plazi.org/id/03B75F77-C637-FFB0-FE0E-D8DEFDC2FB22

treatment provided by

Felipe

scientific name

Ancylorhynchus cruciger ( Loew, 1858 )
status

 

Ancylorhynchus cruciger ( Loew, 1858) View in CoL

Figs 2 View Fig , 3 View Figs 3–30 , 31, 32 View Figs 31–40 , 61–66 View Figs 61–74 , 148 View Fig

Xiphocerus cruciger: Loew 1858: 348 View in CoL ; 1860: 154.

Ancylorrhynchus crux Bezzi, 1908: 377 ; Hull 1960: 217; Oldroyd 1974: 33 (figs 16 head (lat.), 22 head (dor.)). Syn. n.

Ancylorrhyncus cruciger: Kertesz 1909: 100 View in CoL (catalogue); Hull 1960: 217 (fig. 1785 ♂ terminalia); Oldroyd 1974: 31.

Ancylorrhynchus apicalis Curran, 1934: 7 ; Hull 1960: 217; Oldroyd 1974: 33.

Ancylorrhynchus splendens Bromley, 1936: 137 ; Hull 1960: 217; Oldroyd 1974: 33.

Ancylorhynchus apicalis: Oldroyd 1980: 360 View in CoL (catalogue).

Ancylorhynchus cruciger: Oldroyd 1980: 360 View in CoL (catalogue).

Ancylorhynchus crux: Oldroyd 1980: 360 View in CoL (catalogue).

Ancylorhynchus splendens: Oldroyd 1980: 360 View in CoL (catalogue).

Redescription:

Female.

Holotype cruciger (Condition: Excellent; although an old specimen, closely mounted to the pin head, only the tip of the right wing shows any sign of damage):

Head: Dark red-brown to black, silver and gold pruinose, yellow-orange and red-brown setose.Antenna ( Fig. 3 View Figs 3–30 ): Entirely orange, scape yellow and orange setose, pedicel with few pale yellow setae dorsally, postpedicel, tipped with obliquely placed pit-enclosed spine-like sensory element (terminal stylus ‘segment’ absent). Segmental ratios: 1:0.2:4.0. Face blackish, strongly gold pruinose except for lateral parts of epistomal margin, mystax shiny yellow occupying ventral ½ of face. Frons and vertex blackish, silver pruinose, yellow setose; ocellar tubercle fine pale yellow setose (no macrosetae). Occiput blackish, uniformly silver and gold pruinose strongly gold pruinose adjacent to eye margins), orange setose. Palpus orange, orange and dark red-brown setose (few dark setae terminally), terminal palpomere swollen. Proboscis orange and red-brown, red-brown setose.

Thorax: Black with orange areas dorsally, fine silver-grey pruinose, mainly pale yellow setose. Prothorax extensively orange (blackish laterally), fine silver pruinose, orange and dark red-brown (laterally) setose. Mesonotum orange, except for blackish medial stripe and dark patches centrally, anterior of transverse suture, fine silver-grey pruinose, fine short pale yellow setose. Mesonotal macrosetae (orange when present): dc weakly developed posterior to transverse suture (some may have been rubbed off), 3 npl, 2 spal, 3 pal. Scutellum orange except for dark red-brown anteromedial part, fine silver pruinose, disc sparsely yellow setose, hind margin with c. 6 orange weakly developed macrosetae accompanied by minor setae. Pleura extensively black (dorsal part of kepst orange-brown), entirely silver-grey pruinose, setae weak dark brown except for yellow setose ktg. Anatg and mes pnot chiefly blackish (orange-brown areas laterally), contrasting with orange scutellum. Legs: cx orange to brownish black, silver pruinose, white to yellowish setose; tro orange (tro 2 with row of short black setae ventrally, Fig. 2 View Fig ); fem, tib and tar uniformly orange, yellow setose except for few dark red-brown setae ventrally on fem 1 and some tarsomeres, claws dark red-brown with orange proximal parts. Wings ( Fig. 31 View Figs 31–40 ): 10.4× 4.3 mm. Membrane extensively microtrichose (some proximal cells partly bare), distal and posterior parts brown (from about r–m and posterior of Cup), other parts yellowish. Veins orange and dark brown (correlating with brown and yellow areas of membrane). Haltere dark red-brown.

Abdomen: Predominantly blackish with small orange to orange-brown patches dorsally, fine silver-grey pruinose, pale yellow and white setose. T1 uniformly blackish, T2–4 blackish with orange posterior margins and large silver pruinose areas posterolaterally; T5–7 dark red-brown to black, uniformly silver-grey pruinose; T8–9 orange-brown. Sternites uniformly blackish, silver-grey pruinose, white setose. Terminalia (Nelspruit ♀, Figs 64–66 View Figs 61–74 ).Acanthophorites each with 6 well developed macrosetae. Spermathecae with tightly coiled tubes.

There being a fair degree of variation in the morphology of this species (see below) it is believed that the following redescription of the crux holotype provides useful additional information.

Holotype crux (Condition: Fair; both antennae broken off beyond pedicels; right palp with terminal palpomere missing; left mesothoracic leg broken off beyond trochanter. Some green verdigris associated with the pin is evident):

Head: Black, fine silver and gold pruinose, yellow and black setose.Antenna: Scape and pedicel orange, yellow setose (strongly ventrally), postpedicel missing. Segmental ratios: 1:0.3:–. Face black, extensively strong gold pruinose (except lateral parts of epistomal margin and small area below antennal sockets), mystax shiny yellow occupying entire facial profile. Frons and vertex black, fine silver-grey pruinose except for small lateral areas and anterior part of ocellar tubercle, pale yellow setose; ocellar tubercle fine pale yellow setose. Occiput black, strongly silver and silver-gold pruinose, orange setose except for few black setae mostly ventrally situated. Palpus with basal palpomere red-brown, apical palpomere orange-brown, black setose, swollen, apex with terminal sensory element. Proboscis dark red-brown, black setose.

Thorax: Black with orange areas dorsally, fine silver pruinose, pale yellow and black setose. Prothorax dorsally orange, laterally black, fine silver-grey pruinose, black and yellow setose. Mesonotum orange with black cross (broad medial band extending from anterior to posterior margins and transverse band at midlength that almost reaches lateral margins), fine silver-grey pruinose, fine black and yellow setose. Mesonotal macrosetae (yellow when present – some missing and represented only by their sockets): dc not evident, 2–3 npl, 2–3 spal, 3 pal. Scutellum black anteriorly, orange apically, fine silver pruinose, disc sparsely dark red-brown setose, hind margin with c. 4 yellow macrosetae accompanied by minor setae. Pleura entirely dark red-brown to black, entirely silver-grey pruinose, setae confined mainly to anepst (black), kepst (black) and ktg (yellow). Anatg and mes pnot black. Legs: cx black, silver pruinose (cx 3 strongly so), white setose; tro red-brown (tro 2 with row of short black setae ventrally); fem, tib and tar uniformly orange, yellow setose (except for black setae ventrally on fem 1), claws dark red-brown with orange proximal parts. Wings: 10.8× 4.3 mm. Veins orange and brown. Membrane extensively microtrichose (some proximal cells partly bare) giving wings a brownish colour, distal cells darker brown (from about level of r–m crossvein), proximal cells yellowish, posterior cells somewhat transparent. Haltere orange-brown.

Abdomen: Black with yellow patches dorsally, strongly gold (yellow parts) and silver (black parts) pruinose, pale yellow and white setose. T1 black, T2–7 black with extensive yellow areas posteriorly, T8 orange. Sternites entirely black, silver pruinose, white setose. Terminalia: Epigynium orange, acanthophorites well developed.

Male (based on holotype splendens . Condition: Good; hind margins of wings tatty.).

Agrees closely with female holotype ( crux ) except for the following: Head: Antennae entirely orange, postpedicel without terminal stylus ‘segment’, but with terminal pit-enclosed seta-like sensory element.Segmental ratios:1:0.3:4.5. Thorax: Mesonotal macrosetae: 3 npl, 2–3 spal, 3–4 pal. Wings: 9.3× 3.8 mm. Abdomen: T1 black, T2 black with posterior ⅔ yellow, T3–8 extensively yellow except for small black anterolateral parts. Genitalia ( Figs 61–63 View Figs 61–74 ): Epand bifid, forming pair of relatively short and stout lobes. Lobes closely abut proximally, each having curved appearance in dorsal view, distal parts strongly inwardly curved, opposing tips almost touching medially. Proct simple, dorsally situated cerc appearing fused proximally. Goncx well developed, outer lobe projecting posteriorly beyond level reached by epand; dorsal finger-like process completely absent; inner lobe well developed, distal end sclerotised, medially directed (overlapping opposing lobe). Gonst laterally compressed, dorsally directed, tip curved anteriorly (largely hidden from view in undissected genitalia). Hypd well developed, tapering fairly rapidly to broad truncate distal end; distal end with pair of diverging, rounded protuberances each with group of converging setae (readily seen in dry material). Aed (largely hidden in undissected genitalia) with tiny bifid distal tip. Note: Komatipoort ♂ has goncx with a posteroventral indentation which appears to be an aberration.

Variation: A fairly variable species with a wide distribution. Some of the more obvious variations are as follows. Mystax is usually limited to ventral half of face but may extend into dorsal half; mystax is usually shiny bright yellow but may also be pale yellow or even white (e.g. some specimens from Namibia and Swaziland) or even brownish (e.g. some specimens from Somalia). While the blackish mesonotal cross-shaped marking is fairly uniformly shaped some specimens may have more extensive markings while others may almost lack a cross (a Mozambique female). While the majority of specimens have yellowish wings with a brownish distal end (e.g. those from the eastern parts of Southern Africa, Fig. 32 View Figs 31–40 ) there is considerable variation involving a darkening of the proximal and posterior parts of the wing, leaving the yellowish colour limited to the anterocentral parts of the wing (e.g. specimens from Northern Cape of South Africa and Somalia). Femora usually uniform orange, but may have red-brown parts dorsally (e.g. a Swaziland specimen). Abdomen commonly extensively yellowish and strongly gold pruinose, but may also be extensively black and weakly pruinose (intermediate states are encountered).

Holotype cruciger (examined): ♀ SOUTH AFRICA: ‘53’, ‘244’, ‘ Xiphocerus / crucifer [sic]’ [white typed] ( NHRS). Note: Loew (1858) gives provenance as ‘Caffraria (Wahlb.)’. The specimen being collected by Wahlberg on an expedition through the eastern parts of Southern Africa.

Holotype crux (examined): ♀ DR CONGO: ‘Type’ [orange with black edge], ‘Musée / Du Congo Belge / Congo Belge’, ‘ Ancylorrhynchus / crux / ♀ n. sp.’ [much folded label], ‘R. Det. / M / 243’, ‘Holo / A. crux / MRAC’ ( MRAC).

Holotype apicalis : ♀ ZIMBABWE: ‘Matetsi [mission, 18°05'S: 26°07'E] / S. Rhodesia / Apr. 1933 ’ [black on white], ‘Acc. 3475’ [red on white], ‘ Ancylorrhynchus / apicalis / Curran ♀ / Holotype.’ [black on red] ( AMNH). Note: I have not personally handled this holotype, however Tam Nguyen ( AMNH) kindly sent me excellent digital photographs that clearly support the synonymy of apicalis with crux .

Holotype splendens (examined): ♂ SOUTH AFRICA: ‘Barberton [25°47'S: 31°03'E] / De Kaap. B. 30 / 1-5-20 / H.K. Munro’, ‘ Holotype / Ancylorrhynchus / splendens Br’ [red], ‘ Ancylorrhynchus / splendens

Brom. / ♂ / Holotype No. 148’ [red ink] ( NMSA). Allotype ♀ ‘ Bulawayo [20°09'S: 28°35'E] / S Rhodesia / 19.i.1927 / R.H. Stevenson’ [red], ‘ Ancylorrhynchus / splendens ♀ Brom / ♀ / Allotype No. 149’ [red ink] ( NMSA) GoogleMaps .

Other material examined: BOTSWANA: 1♀ ‘Nord-Botswana / Okavango-Delta / Kulturlandschaft / Sepupa [18°44'S: 22°11'E], 3.iii.1994 / le.g. H. Schumann’ ( ZMHB); 1♀ ‘Botswana SE 2226BD / Farmers Brigade 5km / SE of Serowe Hillside / N slope P Forchhammer / Malaise trap 6 / i.88’ ( NMSA); 1♂ ‘Botswana: Serowe #1 / Farmer’s Brigade / 22°25’S 26°44’E 1000m / Date: xi.1989 / Coll: P. Forchhammer / Malaise Trap’ ( NMSA); 1♀ ‘Beretse Pan [23°52'S: 23°24'E] / Bechuanaland / 6.3.1963 / T. Oatley’ ( NMSA). BURUNDI: 1♀ ‘Urundi: Rumonge [03°57'S: 29°27'E] / alt. 790m. 19–20.vi.1948 / F. François’, ‘Sable et broussailles / près rive de Lac / Tanganyika’ ( ISNB) [recorded by Oldroyd (1970)]. DR CONGO: 1♀ ‘ Congo belge: P.N.U. / Mabwe [07°05'S: 22°59'E] (r. E. lac Upemba) / (585m.) 1–11-i-1949 / Mis. G.F.de Witte. 2179a’ ( MRAC). KENYA: 1♀ ‘East Africa / Protectorate: Foot of Kikyugu / Escarpment, / near Naivasha [00°43'S: 36°26'E] / 7.300 feet / 3.iii.1911. / S.A. Neave. / 1912. 155’ ( BMNH); 1♀ ‘Brit. E. Africa / Tsavo River [02°59'S: 38°31'E] / 25.iii.1911 / S. A. Neave’ ( BMNH). MALAWI: 1♀ ‘Nyasaland / Protectorate / Betw. Fort Maguire [ruins, 13°39'S: 34°53'E] / and Fort Johnston, / S-E shore of L. Nyasa / 15.iii.1911. / S. A. Neave / 1912–155’ ( BMNH); 1♀ ‘Nyasaland / Lingadzi, [13°58'S: 33°48'E] 71 / Nr. Domira Bay, / 30.i.1915 1700 ft. / Dr. W.A. Lamborn.’ ( BMNH); 1♀ ‘Nyasaland / Lingadzi, 71 / Nr. Domira Bay, / 7. iii.1915 1700 ft. / Dr.W.A. Lamborn.’ ( BMNH); 1? ‘Nyasaland / Lingadzi, 71 / Nr. Domira Bay, / 2.iii.1915 1700 ft. / Dr. W.A. Lamborn.’ ( BMNH); 1♂ ‘Nyasaland. / S.W. shore L. Nyasa. / Btwn. Ft. Johnston [= Mangochi, 14°28'S: 35°16'E] / & Monkey Bay. 1650 ft. / 25 Feb.–Mch. 4 1910. / S.A. Neave’, ‘Sammlung / F. Hermann’, ‘ Ancylorrh. / cruciger / var crux / Bzzi’, ‘ cruciger crux / Bezzi’, ‘ Ancylorrhynchus / cruciger crux / Bezz. / det. E. O. Engel’, Ancylorrhynchus / crux Bezzi ♂ ’ ( ZSMC); 2♀ ‘Nyasaland / Fort Johnston / Casey’ ( DMSA); 1♂ 1♀ ‘Nyasaland. / S.W shore L Nyasa. / Btwn. Fort Johnston / & Monkey Bay. 1650 ft. / 25 Feb. – Mch 4 1910 / S.A. Neave’ ( BMNH); 1♂ ‘Nyasaland / Lake Nyasa. / Monkey Bay, [14°05'S: 34°55'E] / 1 xi.1915 / Dr.W.A. Lamborn.’ ( BMNH); 1♂ ‘M9085 / Nkapola Hill [14°20'S: 35°10'E] / Malawi / ii.1978 C.D. [Cornell Dudley]’ ( NMSA); 1♂ ‘M9087 / Nkapola Hill / Malawi / ii.1978 C.D.’ ( NMSA); 1♂ ‘M9088 / Nkapola Hill / Malawi / ii.1978 C.D.’ ( NMSA); 1♂ 1♀ ‘G1xG [or GxG1 - in cop.], S.W. of / L. Chilwa [15°12'S: 35°50'E] / Nyasaland / 15.i.1914 / S.A. Neave’ ( BMNH); 1♀ ‘B>B1 / S.W. of / L. Chilwa / Nyasaland / 14.i.1914 / S. A. Neave’ ( BMNH); 1♀ ‘A>A1 / S.W. of / L. Chilwa / Nyasaland / 15.i.1914 / S. A. Neave’ ( BMNH); 1♀ ‘S.W. of / L. Chilwa / Nyasaland / 15.i.1914 / S. A. Neave’ ( BMNH); 1♂ ‘B>B1 / Mlanje [16°02'S: 35°30'E] / Nyasaland / 12.ii.1914 / S.A. Neave’, ‘ Tachinidae / det. 1938 / J. Smart’ ( BMNH); 1♂ ‘24.iii.13 / S.A.N.’, ‘Mlanje / Nyasaland, / 1913. / S. A. Neave’ ( BMNH); 1♀ ‘Mlanje / Nyasaland, / 18.12.1913. / S. A. Neave’ ( BMNH); 1♀ ‘Mlanje / Nyasaland, / 21.iii.1913. / S. A. Neave’ ( BMNH); 1♂ ‘Nyasaland / Mt. Mlanje / 24.ii.1913. / S. A. Neave’ ( BMNH); 1♂ ‘Mt. Mlanje, / Nyasaland / 7.i.1914. / S. A. Neave’ ( BMNH); 1♀ ‘Mlanje / Nyasaland / 14.iv.1913 / S. A. Neave’ ( BMNH); 1♀ ‘Mlanje / Nyasaland / 13.ii.1914 / S. A. Neave’ ( BMNH); 1♀ ‘Mlanje / Nyasaland / 19.iv.1914 / S. A. Neave’ ( BMNH); 1♂ [no locality but is Mlanje] ‘I1<I / 12.iii.13 / S.A. N’, ‘I>I1 / 24.iii.13 / S.A.N.’, ‘ Jassus maculiner - / vis. Stal. / det. In B.M. / W.E. China.’ ( BMNH); 1♂ ‘Portugese / E. Africa [? Mozambique]: / Mlanje Mt. / 6.i.52. / J. Vincent Coll. / B.M. 1933–194.’ ( BMNH); 1♀ ‘Nyasaland / Chiromo [16°33'S: 35°08'E] / 12.v.1910 / Dr. J.E.S. Old’ ( BMNH); 1♀ ‘Nyasaland / Ruo R. [16°33'S: 35°09'E] / 3.iii.1913 / S.A. Neave’ ( BMNH); 1♂ ‘19.iii.13 / S.A.N.’, ‘Sammlung / F. Hermann’, ‘ Ancylorrh. / cruciger / Lw.’, ‘Nyassa [pink]’, ‘ cruciger Lw’, ‘ Ancylorrhynchus / cruciger Lw. / det. E. O. Engel ♂ ’ ( ZSMC). MOZAMBIQUE: 1♂ ‘ Inhambane [23°51'S: 35°29'E] / P.E.Afr / R.F. Lawrence’ ~ ‘Jan. / 1924’ ( SAMC); 1♀ ‘Zambezi / Caia. [25°19'S: 32°58'E] / 29.v.1912 / H. Swale’ ( BMNH); 1♀ ‘Moçambique / Macaneta [25°52'S: 32°14'E] / 23.3.1980 / Coll. H.R. Feijen’ ( NMSA); 3♀ ‘Moçambique / Inhaca [26°01'S: 32°58'E] / 20.4.1980 / Coll. H.R. Feijen’ ( NMSA); 1♀ ‘Moçambique / Goba [26°12'S: 32°08'E] / 12.5.1980 / Coll. H.R. Feijen’ ( NMSA). NAMIBIA: 1♂ ‘Namibia: Epupa / Falls [17°00'S: 13°15'E], Kunene / 20.–22.ii.1994 / le.g. F.Koch’ ( ZMHB); 1♀ ‘Namibia: Kunene / 9km W Ruacana / 17°26’S / 14°09’E / 24.–26.ii.1994 / le.g. H. Schumann’ ( ZMHB); 1♀ ‘Namibia / Popa / Falls [18°07'S: 21°33'E] / 28.ii.–6.iii.1993 / le.g. F. Koch’ ( ZMHB); 1♀ ‘Namibia: Kavango / Camp Popa / 18°07’S / 21°34’E / 28.ii.–5.iii.1994 / le.g. H. Schumann’ ( ZMHB); 1♀ ‘Kaross [19°30'S: 14°20'E] / S.W.A.’ ~ ‘Mus. Exped. / Feb. 1925 ’ ( SAMC); 1♀ ‘Namibia / 20km n of Usakos / Ameib [21°48'S: 15°38'E] / 9 May 1978 / V. B.Whitehead’ ( SAMC). SOMALIA: 2♂ 1♀ ‘Coll.A. Mochi / 17–31.v.35 / Iscia Baidoa [03°04'N: 43°48'E] / Somalia’ ( MCSN); 4♂ 3♀ ‘Iscia Baidoa / Somalia / v.35’ ( MCSN); 1♂ ‘Iscia / Baidoa / V.35 Somalia’ ( MCSN); 1♀ ‘Brit. Somaliland / 1950 / Agric. & Vet. Services / Ex Cotton plants’ ( BMNH). SOUTH AFRICA: 1♂ ‘South Africa, Tvl. / Mogol Nature Reserve / Ellisras Dist. 23.58S / 27.45E. 27–29.ii.1984 / C.D. Eardley’ ( SANC); 1♂ ‘So.Africa: Transvaal / Kruger National Park / Skukuza 2431DC / 1–3.iii.1982 R. Miller / & P. Stabbins Malaise trap’ ( NMSA); 1♂ ‘Moorddrift [24°17'S: 28°57'E] / Oct 1909 / C.J. Swierstra’ ( NMSA); 1♀ ‘GAH Bedford / Pretoria [25°44'S: 28°11'E] / 8.ii.1917 ’ ( BMNH); 1♀ ‘GAH Bedford / Pretoria / 22.ii.1917 ’ ( BMNH); 1♀ ‘Nylstroom [24°42'S: 28°24'E] / 16–31/12/21 / G.P.F. v. Dam’ ( NMSA); 1♂ ‘South Africa, Tvl. / Kruger Nat. Park / Skukuza, 24.59S / 31.55E. 20.i.1984 / M.W. Mansell’ ( SANC); 1♀ ‘Kruger N. Park, Onder Sabie [Rest-camp, 25°07'S: 31°55'E]. / Tvl. 22/5/1969 / L.C. Starke’ ( SANC); 1♀ ‘Platriver [25°10'S: 28°05'E] / Wtb [Waterberg] Distr. / 1–11 [1 xi] 1903 / R.v.Jutrencha’ ( NMSA); 1♂ ‘Komatipoort [25°26'S: 31°57'E], / Tvl. 3.6.69 / M.W. Strydom’ ( SANC); 1♀ ‘Nelspruit [25°28'S: 30°58'E] / 2.1915 / A. Roberts’ ( NMSA); 1♀ ‘Sth Africa Cape Prov / ca 5km W Hotazel / 2722BB 23.iii.1982 / J. Londt & L. Schoeman / Acacias/Grass/Shrubs’ ( NMSA); 1? ‘Lake Sibaya [27°20'S: 32°42'E] / Zululand / 13.4.1968 / T. Schofield’ ( NMSA); 1♂ ‘Magude [Magudu 27°32'S: 31°39'E] 10. 1918 / C.J. Swierstra’ ( NMSA); 1♂ ‘South Africa: N Cape / Witsand Nature Reserve / 28°33.615’S 022°29.105’E / 1160m J Londt & T Dikow / 31.i.–1.ii.2004 Acacia / savanna & white dune area’ ( NMSA); 1♂ ‘South Africa: Natal / Richards Bay Mineral / 2838S 3215E iii.83 / P Atkinson 33m / at light’ ( NMSA); 1♀ ‘Mfongosi [28°42'S: 30°48'E] / Zulu L. / WE Jones’ ~ ‘1914’ ( SAMC); 1♀ ‘Mfongosi / Zulu L. / WE Jones’ ~ ‘Jan. / 1917’ ( SAMC); 1♂ ‘Mfongosi / Zulu L. / WE Jones’ ( BMNH); 1♂ ‘Natal. / Weenen [28°51'S: 30°05'E] / iii.1924 / H.P. Thomasset’ ( BMNH); 1♀ ‘Natal / Estcourt [29°00'S: 29°53'E]’ ~ ‘Haviland 18’ ( NMSA); 1♂ ‘Estcourt / 12/96’ ( NMSA); 1♂ ‘Durban [29°51'S: 31°01'E] / 12/4/32’ ( NMSA); 1? ‘Marley / Stella B [Stella Bush, Durban, 29°51'S: 31°01'E] / 3-1915’ ( SAMC); 1♀ ‘Congella [29°52'S: 31°00'E] / Marley / 3-15’ ( NMSA); 1♀ ‘Metlapitsi [?] / Jan. 1911 / H.G. Breijer’ ( NMSA); 1♀ ‘Kafferal. [Kafferland] / Drège S’, ‘501’, ‘Zool. Mus. / Berlin’ ( ZMHB). SWAZILAND: 1♂ ‘Swaziland #45 / 13km N. of Ngogolo / 26°19’S 31°38’E 300m / Date: 22–24.iv.1991 / J Londt & L Schoeman / Panata Ranch / Bushveld’ ( NMSA). TANZANIA: 1♂ ‘Tanzania, Mkomazi / Game Reserve, Ibaya / Camp, 3.58S 37.48E. / 11 Mar – 5 Apr 1996 ’, ‘S. Van Noort / Malaise trap. / Acacia / Commiphora / / Combretum bushland’ ( SAMC); 1♀ ‘ Zanzibar / Jambiani [06°19'S: 39°33'E] / 23.iii.1993 / K. Guichard’ ( BMNH); 1♂ ‘Tanganyika T. / Kilossa [06°50'S: 36°59'E] / 3.iii.1921 / A. Loveridge’ ( BMNH); 1♂ ‘Tanganyika Ty. / Morogoro [5 places with this name – district 06°50'S: 37°45'E] 6/5/22 / coll: A.H. Ritchie’ ( BMNH); ZAMBIA: 1♂ ‘N. Rhodesia: / 39 mi SW. of / Ft. Jameson [= Chipata, 13°28'S: 32°46'E] / iii.1.58, 1150m’, ‘E.S. Ross & / R.E. Leech / collectors’ ( CASC). ZIMBABWE: 1♀ ‘T.R.S. / 23.i.59 / N.J.M’, ‘Trelawny [17°32'S: 30°27'E] / S. Rhodesia’ ( NMSA); 1♀ ‘A. Cuthbertson / 1929-328’, ‘Salisbury [Harare, 17°50'S: 31°03'E] / S. Rhodesia / Dept.Agric. / ii.1912 ’ ( BMNH); 1♀ ‘Hillside [17°50'S: 31°05'E], S. Rhod. / 2.2.1923 / Swinburne & Stevenson’ ( NMSA); 1♂ ‘Hillside, S. Rhod. / 4.4.1923 / Swinburne & Stevenson’ ( NMSA); 1♂ ‘Umtali [18°58'S: 32°40'E] District / S Rhodesia / 25.i.1931 / P.A. Sheppard’ ( NMSA); 1♂ ‘14.2.47 / Wankie [19°10'S: 26°30'E] / S.R. / D. Lovemoore’ ( AMGS); 1♂ ‘N. Vumba [Mts, 19°53'S: 31°22'E] / S. Rhodesia / 21.2.1966 / D. Cookson’ ( NMSA); 1♀ ‘Vumba / Umtali Dist / Feb. 1932 / PAS [PA Sheppard]’ ( NMSA); 1♂ ‘ Bulawayo [20°09'S: 28°35'E] / 28.iii.1923 / R. Stevenson’ ( NMSA); 1♀ ‘ Bulawayo / 17.iii.1923 / R. Stevenson’ ( NMSA); 1♂ ‘Hope Fountain [mission, 20°16'S: 28°39'E] / S. Rhodesia / 2.i.1923 / Neville Jones’ ( BMNH).

Distribution, phenology and biology: Widely distributed in Southern, Central and East Africa ( Table 1, Fig. 148 View Fig ) with a few records north of the equator. Adults have been collected from October–June (Table 2). The biology is poorly known, but the limited label data available suggest that this is a woodland, savanna, scrubland species that can also survive in semi-arid habitats. Two prey records (both for BMNH males) are known to me, i.e. Diptera (Tachinidae) and Hemiptera (Cicadellidae) .

Similar species: A member of the nomadus species-group with close similarities to both hylaeiformis and sokokensis . It is necessary to compare male genital structure to adequately differentiate these species.

NHRS

Swedish Museum of Natural History, Entomology Collections

MRAC

Musée Royal de l’Afrique Centrale

AMNH

American Museum of Natural History

NMSA

KwaZulu-Natal Museum

ZSMC

Zoologische Staatssammlung

DMSA

Durban Museum

SAMC

Iziko Museums of Cape Town

MCSN

Museo Civico di Storia Naturale, Verona

SANC

Agricultural Research Council-Plant Protection Research Institute

AMGS

Albany Museum

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Diptera

Family

Asilidae

Genus

Ancylorhynchus

Loc

Ancylorhynchus cruciger ( Loew, 1858 )

Londt, Jason G. H. 2011
2011
Loc

Ancylorrhynchus splendens

BROMLEY, S. W. 1936: 137
1936
Loc

Ancylorrhynchus apicalis

CURRAN, C. H. 1934: 7
1934
Loc

cruciger:

KERTESZ, C. 1909: 100
1909
Loc

Ancylorrhynchus crux

BEZZI, M. 1908: 377
1908
Loc

Xiphocerus cruciger: Loew 1858: 348

LOEW, H. 1858: 348
1858
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