Mitrager lucida Lin & Lopardo & Uhl, 2022

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 508-510

MITRAGER LUCIDA ( WUNDERLICH, 1974) COMB. NOV.

( FIGS 38M , 39M , 40N , 55 , 56B; SUPPORTING INFORMATION, FIG. S3A )

Oedothorax lucidus Wunderlich, 1974: 180 , figs 28–33 (Dm).

Type material: Holotype: East Nepal: Mt. Chordung at Jiri , Abies-Rhododendron- forest, 2900 m, 1♂ 1.iv.1973 (SMF 28897, examined) . Paratypes: Jiri, 1800–2000 m, 6♂ i.1970 ( SMF 28898, examined).

Non-type material: 1♂ subadult collected with the paratypes ( SMF 28898).

Diagnosis:

Males: This species shares with M. sexoculata the PME located within the transverse groove between the prosomal elevation and the remaining eyes, the highly sclerotized stump-like setae on both upper and lower surfaces within the groove, and one retrolateral trichobothrium on male palpal tibia instead of the common two. This species can be distinguished from M. sexoculata by the smaller TPA and the papillaebearing PT.

Description:

Male (holotype, SMF 28897): Total length: 1.90. Prosoma: 0.94 long, 0.67 wide, region including PME elevated, transverse groove between elevation and other eyes, with highly sclerotized stump-like setae on both upper and lower surfaces within groove ( Figs 38M , 56B ). Eyes: AME-AME: 0.03, AME width: 0.04, AME-ALE: 0.04, ALE width: 0.05, ALE-PLE: 0.01, PLE width: 0.04; PME inside PME-lobe. Clypeus: slightly elevated, with small setae scarcely distributed. Sternum: 0.50 long, 0.47 wide. Chelicerae: stridulatory striae imbricated, rows widely and evenly spaced ( Fig. 39M ). Legs: dorsal proximal macroseta on tibia III 2.93 times diameter of tibia; Tm I: 0.85. Pedipalp: patella prolateral proximal vertical macrosetae absent; tibia with one prolateral, one retrolateral trichobothria;

TPS scaly, straight, wide at base, slightly retrolaterally directed; TPA slightly elevated; TRA distally oriented; PC median-sized, base not visible from dorsal view, distal setae close to distal clasp, distal clasp without striae, distally extended ( Fig. 55A ); T without papillae; PT with long papillae; TS long, thick, with several small papillae at base ( Fig. 55D ); MSA present; DSA not pointed, not turned retrolaterally; EM flat, without papillae, not exceeding ARP ( Fig. 55C ); ARP rounded at tip, striated; LER without striae, extended dorsal to E; VRP absent; TP broad ( Fig. 55D ); E retrolaterally spiral, anterior margin at base round ( Fig. 55B ). Opisthosoma: dorsal pattern see Fig. 40N ; PMS with mAP, AC absent; PLS with triad, AC absent ( Fig. 55E ).

Female: Unknown.

Variation: The measurements are based on examined material.

Males (N = 7, means in parentheses): Total length 1.88 – 2.13 (1.99). Prosoma: 0.94 – 0.98 (0.96) long, 0.67 – 0.74 (0. 71) wide. Legs: dorsal proximal macroseta on tibia I, II, III and IV 0.78 – 1.29 (1.08, N = 5), 0.77 – 1.24 (0.98, N = 6), 0.44 – 0.70 (0.59) and 1.57 – 2.51 (2.01, N = 3) times diameter of tibia, respectively; Tm I: 0.76 – 0.86 (0.82).

Distribution: East Nepal.

Habitat: Forests.

Figures

Figure 3. Continuation of Figure 2, showing Clade 36.

Figure 38. Male prosomal morphology of Clade 55 in Figure 3, lateral.A, Mitrager hirsuta (Wunderlich, 1974) (Wunderlich, 1974, fig. 8). B, M. clypeellum (Tanasevitch, 1998) (traced from photograph). C, M. elongata (Wunderlich, 1974) (Wunderlich, 1974, fig. 13). D, M. cornuta (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 16). E, M. villosa (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 89). F, M. angela (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 14). G, M. coronata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 6). H, M. sexoculorum (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 19). I, M. globiceps (Thaler, 1987) (traced from photograph). J, M.lineata (Wunderlich, 1974) (traced from photograph).K, M. dismodicoides (Wunderlich, 1974) (traced from photograph). L, M. tholusa (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 24). M, M. lucida (Wunderlich, 1974) (Wunderlich, 1974, fig. 30). N, M. sexoculata (Wunderlich, 1974) (traced from photograph). O. M. noordami van Helsdingen, 1985 (van Helsdingen, 1985, fig. 1). P, M. unicolor (Wunderlich, 1974) (traced from photograph). Q, M. rustica (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 75). R, M. assueta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 1). S, M. malearmata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 50). T, M. savigniformis (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 40). U, M. falcifer (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 46). V, M. modesta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 36). W, M. lopchu (Tanasevitch, 2015) (traced from photograph). X, M. falciferoides (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 35).

Figure 39. Male chelicerae, stridulatory striae on lateral side. A, Mitrager hirsuta. B, M. clypeellum. C, M. elongata. D, M. cornuta. E, M. villosa. F, M. angela. G, M. coronata. H, M. sexoculorum. I, M. globiceps. J, M. lineata. K, M. dismodicoides. L, M. tholusa. M, M. lucida. N, M. sexoculata. O, M. noordami. P,M. unicolor. Q, M. rustica. R, M. assueta. S, M. savigniformis. T, M. falcifer. U, M. modesta. V, M. falciferoides. Scale bars 0.05 mm.

Figure 40. Opisthosoma, dorsal view. A–L, N–Y, male; D, female. A, Mitrager hirsuta (Wunderlich, 1974). B, M. clypeellum (Tanasevitch, 1998). C, M. elongata (Wunderlich, 1974). D, M. cornuta (Tanasevitch, 2015). E, M. villosa (Tanasevitch, 2015). F,M. angela (Tanasevitch, 1998). G, M. coronata (Tanasevitch, 1998). H, M. sexoculorum (Tanasevitch, 1998). I, M. globiceps (Thaler, 1987). J, M. lineata (Wunderlich, 1974). K, M. dismodicoides (Wunderlich, 1974). L, M. tholusa (Tanasevitch, 1998), holotype. M, ‘M. tholusa’, paratype (not conspecific with holotype). N, M. lucida (Wunderlich, 1974). O, M. sexoculata (Wunderlich, 1974). P,. M. noordami van Helsdingen, 1985. Q, M. unicolor (Wunderlich, 1974). R, M. rustica (Tanasevitch, 2015). S, M. assueta (Tanasevitch, 1998). T, M. savigniformis (Tanasevitch, 1998), paratype. U, M. savigniformis, holotype. V, M. falcifer (Tanasevitch, 1998). W, M. modesta (Tanasevitch, 1998). X, M. lopchu (Tanasevitch, 2015). Y, M. falciferoides (Tanasevitch, 2015). Scale bars 0.1 mm.

Figure 55. Mitrager lucida (Wunderlich, 1974). A–D, male left palp. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, male spinnerets. Scale bar 0.1 mm.

Figure 56. Setae inside inter-AME-PME groove, dorsal view, samples soaked in Eugenol.A, Mitrager tholusa (Tanasevitch, 1998). B, M. lucida. Scale bars 0.1 mm.