Mitrager assueta, (TANASEVITCH, 1998)

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 491

MITRAGER ASSUETA (TANASEVITCH, 1998) COMB. NOV.

( FIGS 38R , 39R , 40S , 42 , 43B; SUPPORTING INFORMATION, FIG. S4E )

Oedothorax assuetus Tanasevitch, 1998a: 431 , figs 1–5 (Dm).

Type material: Holotype: Nepal: Kathmandu , Godawari, foot of Phulchoki Mt., 1700 m, ♂ 19.iii.1980, leg. Martens & A. Ausobsky ( SMF 38848, examined) . Paratypes: Nepal: same locality, together with holotype, 1♂, leg. J. Martens & A. Ausobsky (body: SMF 38852, palp: SMF 38841, examined) .

Diagnosis:

Males: A depression between protegulum and tegulum is unique for this species. Further distinguished from most Mitrager species by a prosoma with a hump comprising the PME, and an ocular region bearing dense setae. Distinguished from M. falcifer , M. falciferoides and ‘ Oe. ’ meghalaya incertae sedis by the smaller hump elevation; from M. malearmata by the longer spike prolateral to the palpal tibia prolateral apophysis; and from M. modesta , M. lopchu and M. rustica by the smaller body size.

Description:

Male (holotype): Total length: 2.40. Prosoma: 1.01 long, 0.77 wide, PME- and postocular region slightly elevated, with one strong seta at peak of elevation pointing forwards, interocular region with strong setae pointing upwards ( Figs 38R , 43B ). Eyes: AME- AME: 0.04, AME width: 0.04, AME-ALE: 0.03, ALE width: 0.08, ALE-PLE: 0, PLE width: 0.07, PLE-PME: 0.07, PME width: 0.07, PME-PME: 0.11. Clypeus: not hirsute. Sternum: 0.59 long, 0.57 wide. Chelicerae: stridulatory striae imbricated, rows widely and evenly spaced ( Fig. 39R ). Legs: Tm I: 0.56. Pedipalp: patella prolateral proximal vertical macrosetae present; tibia with one prolateral, two retrolateral trichobothria; TPS scaly, retrolaterally pointed; TPA absent; TRA bent retrolaterally; PC short, base not visible from dorsal view, distal setae close to distal clasp, distal clasp without striae, directed retrolaterally ( Fig. 42A ); T without papillae; PT with long papillae, a depression between T and PT creates a discontinuous appearance from lateral view ( Fig. 42D ); MSA present ( Fig. 42D ); DSA not pointed, not retrolaterally curved ( Fig. 42A ); EM flat, anterior margin without obvious papillae, exceeds ARP ( Fig. 42D ); ARP pointed, angled at tip; LER with striae on distal margin, extended dorsal to E; VRP absent; TP round at tip, ventro-posterior area with round extension ( Fig. 42B ); E retrolaterally spiral, anterior margin at base slightly wavy ( Fig. 42B ). Opisthosoma: dorsal pattern see Fig. 40S ; PMS with mAP, AC absent; PLS with triad, one AC ( Fig. 42E ).

Male (paratype): Total length: 2.39. Prosoma: 1.05 long, 0.86 wide. Legs: Tm I: 0.59.

Figures

Figure 4. Character transformation of selected prosomal characters reconstructed on the first among the three most- parsimonious trees generated from the analysis of Matrix II.

Figure 38. Male prosomal morphology of Clade 55 in Figure 3, lateral.A, Mitrager hirsuta (Wunderlich, 1974) (Wunderlich, 1974, fig. 8). B, M. clypeellum (Tanasevitch, 1998) (traced from photograph). C, M. elongata (Wunderlich, 1974) (Wunderlich, 1974, fig. 13). D, M. cornuta (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 16). E, M. villosa (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 89). F, M. angela (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 14). G, M. coronata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 6). H, M. sexoculorum (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 19). I, M. globiceps (Thaler, 1987) (traced from photograph). J, M.lineata (Wunderlich, 1974) (traced from photograph).K, M. dismodicoides (Wunderlich, 1974) (traced from photograph). L, M. tholusa (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 24). M, M. lucida (Wunderlich, 1974) (Wunderlich, 1974, fig. 30). N, M. sexoculata (Wunderlich, 1974) (traced from photograph). O. M. noordami van Helsdingen, 1985 (van Helsdingen, 1985, fig. 1). P, M. unicolor (Wunderlich, 1974) (traced from photograph). Q, M. rustica (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 75). R, M. assueta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 1). S, M. malearmata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 50). T, M. savigniformis (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 40). U, M. falcifer (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 46). V, M. modesta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 36). W, M. lopchu (Tanasevitch, 2015) (traced from photograph). X, M. falciferoides (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 35).

Figure 39. Male chelicerae, stridulatory striae on lateral side. A, Mitrager hirsuta. B, M. clypeellum. C, M. elongata. D, M. cornuta. E, M. villosa. F, M. angela. G, M. coronata. H, M. sexoculorum. I, M. globiceps. J, M. lineata. K, M. dismodicoides. L, M. tholusa. M, M. lucida. N, M. sexoculata. O, M. noordami. P,M. unicolor. Q, M. rustica. R, M. assueta. S, M. savigniformis. T, M. falcifer. U, M. modesta. V, M. falciferoides. Scale bars 0.05 mm.

Figure 40. Opisthosoma, dorsal view. A–L, N–Y, male; D, female. A, Mitrager hirsuta (Wunderlich, 1974). B, M. clypeellum (Tanasevitch, 1998). C, M. elongata (Wunderlich, 1974). D, M. cornuta (Tanasevitch, 2015). E, M. villosa (Tanasevitch, 2015). F,M. angela (Tanasevitch, 1998). G, M. coronata (Tanasevitch, 1998). H, M. sexoculorum (Tanasevitch, 1998). I, M. globiceps (Thaler, 1987). J, M. lineata (Wunderlich, 1974). K, M. dismodicoides (Wunderlich, 1974). L, M. tholusa (Tanasevitch, 1998), holotype. M, ‘M. tholusa’, paratype (not conspecific with holotype). N, M. lucida (Wunderlich, 1974). O, M. sexoculata (Wunderlich, 1974). P,. M. noordami van Helsdingen, 1985. Q, M. unicolor (Wunderlich, 1974). R, M. rustica (Tanasevitch, 2015). S, M. assueta (Tanasevitch, 1998). T, M. savigniformis (Tanasevitch, 1998), paratype. U, M. savigniformis, holotype. V, M. falcifer (Tanasevitch, 1998). W, M. modesta (Tanasevitch, 1998). X, M. lopchu (Tanasevitch, 2015). Y, M. falciferoides (Tanasevitch, 2015). Scale bars 0.1 mm.

Figure 42. Mitrager assueta (Tanasevitch, 1998). A–D, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, male spinnerets. Scale bars 0.1 mm.

Figure 43. Interocular strong setae, lateral view. A, ‘Oedothorax’ meghalaya (Tanasevitch, 2015). B, Mitrager assueta (Tanasevitch, 1998). Scale bars 0.1 mm.