Mitrager angela Lin & Lopardo & Uhl, 2022

Lin, Shou-Wang, Lopardo, Lara & Uhl, Gabriele, 2022, Evolution of nuptial-gift-related male prosomal structures: taxonomic revision and cladistic analysis of the genus Oedothorax (Araneae: Linyphiidae: Erigoninae), Zoological Journal of the Linnean Society 195, pp. 417-584 : 485-489

MITRAGER ANGELA (TANASEVITCH, 1998) COMB. NOV.

( FIGS 26A, B , 38F , 39F , 40F , 41 ; SUPPORTING INFORMATION, FIG. S3D )

Oedothorax angelus Tanasevitch, 1998a: 433 , figs 14–18 (Dm).

Type material: Holotype: Nepal: Panchthar District, Dhorpar Kharka , Rhododendron & Lithocarpus forest, 2700 m, ♂ 13.–16.iv.1988, leg. J. Martens & W. Schawaller (SMF 38828, examined) . Paratypes: Nepal: same locality, together with holotype, 4♂ ( SMF 38853, examined) 2♂ ( ZMMU), leg. Martens & W. Schawaller; Panchthar District, Paniporua, 2300 m, mixed broadleaved forest, 4♂ ( SMF 38860, examined) 2♂ ( ZFMK) 1♂ ( ZMMU) 16.–20.iv.1988, leg. J. Martens & W. Schawaller.

Diagnosis:

Males: Distinguished from all other Mitrager species (except from M. cornuta , M. coronata and M. villosa ) by the two pairs of branched setae in the ocular region; distinguished from the aforementioned three species by the absence of postocular groove.

Description:

Male (holotype): Total length: 1.97. Prosoma: 0.91 long, 0.73 wide, PME- and postocular region elevated, posteriorly with setae bifurcated anterior-posteriorly;

inter-PME region with two pairs of branched setae, posterior pair longer than anterior ( Figs 26A, B , 38F ). Eyes: AME-AME: 0.05, AME width: 0.04, AME-ALE: 0.05, ALE width: 0.06, ALE-PLE: 0.01, PLE width: 0.06, PLE-PME: 0.06, PME width: 0.05, PME-PME: 0.13. Clypeus: not hirsute. Sternum: 0.51 long, 0.53 wide. Chelicerae: mastidia absent, stridulatory striae imbricated, rows widely and evenly spaced ( Fig. 39F ). Legs: dorsal proximal macroseta on tibia I and III 0.34 and 0.35 times diameter of tibia, respectively; Tm I: 0.83. Pedipalp: patella prolateral proximal vertical macrosetae present; tibia with one prolateral, two retrolateral trichobothria; TPS without scale, retrolaterally pointed; several parallel slit organs prolateral to TPS; TPA absent; TRA bent retrolaterally ( Fig. 41C ); PC median-sized, base not visible from dorsal view, distal setae close to distal clasp, distal clasp with striae, directed retrolaterally, ( Fig. 41A ); T without papillae, PT short, TS short, without papillae ( Fig. 41D ); MSA present ( Fig. 41A ); DSA pointed, retrolaterally curved ( Fig. 41D ); EM flat, anterior margin with papillae, length exceeds ARP ( Fig. 41B ); ARP pointed, bent distally; LER without striae, extended dorsal to E; VRP absent; TP round at tip ( Fig. 41B ); E retrolaterally spiral, anterior margin at base slightly wavy ( Fig. 41B ). Opisthosoma: dorsal pattern see Fig. 40F ; PMS with mAP, one AC; PLS with triad, one AC ( Fig. 41F ).

Figures

Figure 3. Continuation of Figure 2, showing Clade 36.

Figure 26. Branched setae at interocular region. A–B, Mitrager angela (Tanasevitch, 1998). A, frontal view. B, lateral view. C, M. coronata (Tanasevitch, 1998), frontal view. D, Callitrichia sellafrontis Scharff, 1990. E–F, Ca. gloriosa (Jocqué, 1984). E, overview. F, closer view.

Figure 38. Male prosomal morphology of Clade 55 in Figure 3, lateral.A, Mitrager hirsuta (Wunderlich, 1974) (Wunderlich, 1974, fig. 8). B, M. clypeellum (Tanasevitch, 1998) (traced from photograph). C, M. elongata (Wunderlich, 1974) (Wunderlich, 1974, fig. 13). D, M. cornuta (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 16). E, M. villosa (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 89). F, M. angela (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 14). G, M. coronata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 6). H, M. sexoculorum (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 19). I, M. globiceps (Thaler, 1987) (traced from photograph). J, M.lineata (Wunderlich, 1974) (traced from photograph).K, M. dismodicoides (Wunderlich, 1974) (traced from photograph). L, M. tholusa (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 24). M, M. lucida (Wunderlich, 1974) (Wunderlich, 1974, fig. 30). N, M. sexoculata (Wunderlich, 1974) (traced from photograph). O. M. noordami van Helsdingen, 1985 (van Helsdingen, 1985, fig. 1). P, M. unicolor (Wunderlich, 1974) (traced from photograph). Q, M. rustica (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 75). R, M. assueta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 1). S, M. malearmata (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 50). T, M. savigniformis (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 40). U, M. falcifer (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 46). V, M. modesta (Tanasevitch, 1998) (Tanasevitch, 1998a, fig. 36). W, M. lopchu (Tanasevitch, 2015) (traced from photograph). X, M. falciferoides (Tanasevitch, 2015) (Tanasevitch, 2015, fig. 35).

Figure 39. Male chelicerae, stridulatory striae on lateral side. A, Mitrager hirsuta. B, M. clypeellum. C, M. elongata. D, M. cornuta. E, M. villosa. F, M. angela. G, M. coronata. H, M. sexoculorum. I, M. globiceps. J, M. lineata. K, M. dismodicoides. L, M. tholusa. M, M. lucida. N, M. sexoculata. O, M. noordami. P,M. unicolor. Q, M. rustica. R, M. assueta. S, M. savigniformis. T, M. falcifer. U, M. modesta. V, M. falciferoides. Scale bars 0.05 mm.

Figure 40. Opisthosoma, dorsal view. A–L, N–Y, male; D, female. A, Mitrager hirsuta (Wunderlich, 1974). B, M. clypeellum (Tanasevitch, 1998). C, M. elongata (Wunderlich, 1974). D, M. cornuta (Tanasevitch, 2015). E, M. villosa (Tanasevitch, 2015). F,M. angela (Tanasevitch, 1998). G, M. coronata (Tanasevitch, 1998). H, M. sexoculorum (Tanasevitch, 1998). I, M. globiceps (Thaler, 1987). J, M. lineata (Wunderlich, 1974). K, M. dismodicoides (Wunderlich, 1974). L, M. tholusa (Tanasevitch, 1998), holotype. M, ‘M. tholusa’, paratype (not conspecific with holotype). N, M. lucida (Wunderlich, 1974). O, M. sexoculata (Wunderlich, 1974). P,. M. noordami van Helsdingen, 1985. Q, M. unicolor (Wunderlich, 1974). R, M. rustica (Tanasevitch, 2015). S, M. assueta (Tanasevitch, 1998). T, M. savigniformis (Tanasevitch, 1998), paratype. U, M. savigniformis, holotype. V, M. falcifer (Tanasevitch, 1998). W, M. modesta (Tanasevitch, 1998). X, M. lopchu (Tanasevitch, 2015). Y, M. falciferoides (Tanasevitch, 2015). Scale bars 0.1 mm.

Figure 41. Mitrager angela (Tanasevitch, 1998). A–E, male right palp, images flipped horizontally. A, retrolateral view. B, prolateral view. C, dorsal view. D, ventral view. E, apical view. F, male spinnerets. Scale bar 0.1 mm.