Neomolgus Oudemans

Hernandes, Fabio A., Skvarla, Michael J., Fisher, Ray, Dowling, Ashley P. G., Ochoa, Ronald, Ueckermann, Edward A. & Bauchan, Gary R., 2016, Catalogue of snout mites (Acariformes: Bdellidae) of the world, Zootaxa 4152 (1), pp. 1-83 : 41-45

Neomolgus Oudemans

Molgus Dujardin, 1842: 316 (nomen nudum); Molgus Trouessart, 1894: 118 ; Thor, 1913: 30; 1931a: 54.

Molgus (Hoplomolgus) Berlese, 1923: 237 .

Neomolgus Oudemans, 1937: 1229 ; Atyeo, 1960a: 389; Soliman, 1975: 48; Soliman & Zaher, 1975: 78; Wallace & Mahon, 1976: 116; Michocka, 1987: 44.

Type-species: Acarus littoralis Linnaeus, 1758 by original designation.

251. Neomolgus aegyptiacus Soliman & Mohamed, 1972b: 90 ; Faculty of Agriculture farm, Giza, Egypt, from clover field associated with Bryobia cristata ( Acari , Tetranychidae ).

Remarks. Male unknown; although the species was formally described in 1975, three years earlier Soliman & Mohamed (1972b) published the biology of this species along with a morphological description of all stages. The authority and date must be therefore attributed to Soliman & Mohamed, 1972b.

Type deposition. Probably at CUE.

252. Neomolgus aequalis ( Schweizer & Bader, 1963) : 238 comb. nov.; Kreuzlingen, Switzerland, ex moss. Original designation: Hoplomolgus aequalis Schweizer & Bader.

Type deposition. NMB (A 211a).

253. Neomolgus berlesei ( Trägårdh, 1902b) : 17 comb. nov.; in humid areas and under stones, Italy. Original designation: Bdella capillata var. berlesei Trägårdh, 1902b: 17 . Other name: Molgus (Hoplomolgus) berlesei, Thor, 1931a: 57 .

a. Molgus (Hoplomolgus) tuberculatus Berlese, 1923: 238 synonymy according to Thor (1931a: 57); Hoplomolgus tuberculatus, Schweizer & Bader, 1963: 242 .

Type deposition. Unknown.

254. Neomolgus capillatus ( Kramer, 1881) : 446; Germany—Willmann, 1939: 431; 1956: 243. Original designation: Bdella capillata Kramer ; Canestrini, 1886: 188; Berlese, 1891; Michael, 1896: 514; Kramer, 1896: 447; Trägårdh, 1902b: 17; Bäbler, 1910: 812; Halbert, 1915: 112; Hull, 1918: 40. Other names: Molgus capillatus, Trouessart, 1888: 619 ; Thor, 1904b: 77; 1909: 7; Oudemans, 1914: 123; Schweizer, 1922: 79; Thor, 1926: 139; Thor, 1930a: 95; Hoplomolgus capillata, Trägårdh, 1931: 49 ; Molgus (Hoplomolgus) capillatus, Vitzthum, 1929: 59 ; Molgus (Molgus) capillatus, Thor, 1931a: 60 .

Distribution. Germany ( Voigts & Oudemans 1906), Ireland ( Hull 1915), Germany, Switzerland, Italy, England, Ireland, Norway, Svalbard Islands ( Thor 1931a), France ( Schweizer 1922), Austria ( Willmann 1951), Iceland ( Atyeo & Tuxen 1962), Georgia ( Gomelauri 1963b), Switzerland ( Schweizer & Bader 1963), Australia (south western W.A.) ( Wallace & Mahon 1976), Ukraine ( Kuznetsov & Livshits 1979a), Poland ( Michocka 1987), Iran ( Ostovan & Kamali 1995; Kamali et al. 2001).

Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976), Michocka (1987).

Type deposition. CRA.

255. Neomolgus clypeatus ( Thor, 1930b) : 17; Sjusörn (Lillehammer), Norway.— Atyeo & Tuxen, 1962: 290. Original designation: Molgus clypeatus Thor. Other name: Molgus (Hoplomolgus) clypeatus, Thor: 1931a : 55. Distribution. Iceland ( Atyeo & Tuxen 1962), Syria ( Lattakia ) ( Soliman & Zaher 1975), Australia ( Wallace & Mahon 1976), Ukraine ( Kuznetsov & Livshits 1979a), Iran ( Ostovan & Kamali 1995; Kamali et al. 2001). Redescriptions. Atyeo & Tuxen (1962), Wallace & Mahon (1976).

Type deposition. Lost.

256. Neomolgus egregia (Koch, 1839) : 23 comb. nov.; in humid meadows, Europe. Original designation: Bdella egregia Koch; Koch, 1842 : 74; Walckenaer & Gervais, 1847: 532; Andersén, 1863: 184; Canestrini & Fanzago, 1877: 101; Oudemans, 1937: 1196. Other name: Molgus egregious, Thor, 1931a: 63 . Bdella vulgaris, Canestrini, 1886: 183 ; Berlese, 1888: 7; 1893: 43; Halbert, 1915: 113; Schweizer & Bader, 1963: 230; synonymy according to Thor (1931a: 63).

Bdella longitarsa Karsch, 1881: 37 ; Molgus egregius var. longitarsa, Thor, 1931a: 63 . Distribution. Middle Europe ( Germany, France, Italy, Sweden) ( Thor 1931a), Japan ( Karsch 1881), Ireland ( Hull 1915).

Type deposition. Unknown; type of B. longitarisa in ZMB.

257. Neomolgus helveticus ( Schweizer & Bader, 1963) : 239 comb. nov.; Buffalora mountain, National Park, Switzerland, under stone. Original designation: Hoplomolgus helveticus Schweizer & Bader. Type deposition. NMB (NP 1471).

258. Neomolgus iraniensis Eghbalian, Khanjani, Safaralizadeh & Ueckermann, 2016: 296 ; ex soil and litter under wild almond trees, Amygdalus scoparia L. ( Rosaceae ), Palangan village, Kamyaran, Kermanshah Province, Iran.

Type deposition. Holotype and paratypes at University of Bu-Ali Sina, Hamedan, Iran ; paratype female at NCA.

259. Neomolgus lacustris ( Hull, 1915) : 121 comb. nov.; on soil, England. Original designation: Bdella lacustris Hull.—Hull, 1918 : 40. Other name: Molgus (Molgus) lacustris, Thor, 1931a: 62 . a. Molgus (Hoplomolgus) sublimis Berlese, 1923: 238 synonymy according to Thor (1931a: 62). Type deposition. Unknown.

260. Neomolgus littoralis ( Linnaeus, 1745) : 96; unknown locality.— Atyeo, 1960a: 390. Original designation: Acarus littoralis Linnaeus, 1745: 96 ; 1758: 618; Hammer, 1775: 158; Fabricius, 1775: 815; 1780: 225; 1781: 493; 1787: 374; Müller, 1776: 187; Oudemans, 1926b: 117; Johnston, 1845: 229; Oudemans, 1929: 316. Other names: Bdella littoralis, Neuman, 1875: 104 ; Hull, 1918: 40; 1922: 622; Oudemans, 1926a: 116; Trägårdh, 1904: 46; 1912: 419; Bdella vulgaris var. littoralis Moniez, 1890a: 30, 1890b: 196 .

a. Bdella littoralis + capillata var. pallipediformis Trägårdh, 1902b: 18 ; synonymy according to Thor (1931a: 61); Molgus littoralis, Thor, 1904a: 15 ; 1904b: 78; 1913: 30; 1928: 216; 1930a: 96; 1931a: 61; King, 1914: 138; Berlese, 1923: 237; Trägårdh, 1928: 8; 1931: 47. Trägårdh, 1928: 8; 1931: 47.

b. Bdella groenlandica Trägårdh, 1904: 48 ; Hull, 1922: 622; synonymy according to Thor (1930a).

c. Molgus (Molgus) littoralis var. seurati Berlese, 1923: 237 ; synonymy according to Thor (1931a).

d. Acarus petrarum ruber L.; Ström, 1762: 196; Olafsen & Povelsen, 1772: 607; synonymy according to Thor (1904).

e. Acarus basteri Johnston, 1836: 353 ; Bdella basteri ; Michael, 1896: 478; synonymy according to Thor (1904).

f. Acarus longicornis, Johnston, 1845: 227 ; synonymy according to Thor (1904).

g. Bdella podurophila White, 1852: 210 ; synonymy according to Thor (1931a).

h. Bdella arctica Thorell, 1871: 698 ; Kramer, 1897: 79; Trägårdh, 1900: 9; Banks, 1919: 11; Molgus arcticus (Thorell) , Trouessart, 1888: 119; Thor, 1902: 162; synonymy according to Thor (1904).

i. Bdella marina Packard, 1873: 544 ; 1884: 828; Banks, 1894: 220; Michael 1896: 479; Banks 1907: 596; 1908: 5; Thor, 1931a: 39; synonymy according to Thor (1904).

j. Molgus longicornis, Murray, 1877: 143 ; synonymy according to Thor (1931a).

k. Bdella grandis Kramer, 1879: 131 ; synonymy according to Trägårdh (1904: 46).

l. Bdella villosa Kramer & Neuman, 1883: 525 ; Michael 1896: 479; Oudemans & Koenike, 1897: 238; Banks, 1899: 348; 1907: 596; Molgus villosa, Trouessart, 1888: 619 ; synonymy according to Thor (1904).

m. Eupalus sanguineus Trouessart, 1888: 753 ; Bdella sanguinea, Trouessart, 1894: 119 ; Michael 1896: 478; Trägårdh, 1900: 9; Molgus sanguineus, Trouessart, 1894: 117 ; synonymy according to Thor (1904).

n. Bdella villosa + frigida Banks, 1899: 348 ; synonymy according to Thor (1931a).

Distribution. England ( Hull 1918), Canada (Nunavut) ( Summerhayes & Elton 1928), Alaska, Hudson Bay Area, Greenland, Iceland, Spitsbergen, Bering Island, Russian Lapland (Kola-Hafvon), Novaya Zemlya (Matochkin Strait), Finland ( Atyeo 1960a), Japan ( Ehara 1960, 1961, Shiba 1971), Iceland ( Atyeo & Tuxen 1962).

Redescriptions. Atyeo (1960a), Atyeo & Tuxen (1962).

Remarks. Atyeo (1960a: 391) reported specimens from Matochkin Strait lacking trichobothria on tibia IV, and having a long tactile seta in the position normally occupied by the trichobothria.

Type deposition. Unknown.

261. Neomolgus longipalpis ( Karpelles, 1893) : 106 comb. nov.; Maramures, Hungary, at Pop Ivan mountain. Original designation: Bdella longipalpis Karpelles. Other name: Molgus longipalpis, Thor, 1931a: 63 . Type deposition. Unknown.

262. Neomolgus longipalpus Kuznetsov, 1984: 774 ; park of the Nikita botanical garden, Yalta, Crimea, ex litter. Remarks. Male unknown. According to ICZN (article 57.6), the one-letter difference in spelling in relation to the previous species is enough to prevent the homonymy.

Type deposition. NBG.

263. Neomolgus lumarius Atyeo, 1962: 292 in Atyeo & Tuxen ; Háitindur, Southeast Iceland.

Type deposition. Holotype and paratypes at ZMUC; paratypes at USNM, BMNH and SAM .

264. Neomolgus maculatus ( Karpelles, 1893) : 26 comb. nov.; Sălaj, Hungary. Original designation: Bdella maculata Karpelles. Other names: Molgus maculatus, Thor, 1931a: 64 ; Molgus maculatus var. pilosa ; Thor, 1931a: 64.

Type deposition. Unknown.

265. Neomolgus monticola Willmann, 1951: 162 ; Hohe Tauern, Switzerland. Other name: Hoplomolgus monticola, Schweizer & Bader, 1963: 240 .

Type deposition. ZSM.

266. Neomolgus mutabilis Atyeo, 1960a: 392 ; 2 miles south of Galena , Cherokee Co., Kansas, USA, under board. Distribution . United States (Kansas, Texas) , Mexico (Distrito Federal) ( Atyeo 1960a).

Type deposition. Holotype and paratypes at SEMC, paratypes at USNM, BMNH and SAM .

267. Neomolgus obsoletus ( Berlese, 1923) : 238 comb. nov.; in Germany and Norway. Original designation: Molgus (Hoplomolgus) obsoletus, Thor, 1931a: 58 ; Hoplomolgus obsoletus, Schweizer & Bader, 1963: 242 . Type deposition. CRA.

268. Neomolgus ontakensis Shiba, 1971: 99 ; Mt. Ontaké , Nagano pref., Japan.

Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan.

269. Neomolgus pallipes ( Koch, 1879) : 131; Siberia, Russia. Original designation: Bdella pallipes Koch ; Hull, 1918: 41; Hull, 1922: 622; Neomolgus pallipes ; Mihelčič, 1958a; Willmann, 1952: 166.

a. Bdella hirta Kramer, 1896: 447 ; synonymy according to Thor (1931a: 58).

b. Bdella basteri Johnston forma pallipes, Trägårdh, 1901: 61 ; synonymy according to Thor (1931a: 58).

c. Bdella capillata var. pallipes, Trägårdh, 1902b: 16 ; 1904: 47; synonymy according to Thor (1931a: 59).

d. Molgus (Molgus) pallipes, Thor, 1931a: 58 .

e. Molgus capillatus, Thor, 1927: 139 ; 1930: 95; synonymy according to Thor (1931a: 59); Molgus capillatus var. pallipes, Trägårdh, 1928: 8 ; synonymy according to Thor (1931a: 59).

Distribution. Canada ( Summerhayes & Elton 1928), Northern Europe and Asia (Siberia, Novaja, Semlja, Sweden, Norway, Svalbard, England, Borkum Island) ( Thor 1931a), Germany (Wangerooge Island) ( Willmann 1952), Spain ( Mihelčič 1958a).

Type deposition. BMNH.

270. Neomolgus paracapillatus Michocka, 1987: 45 ; Warsaw, Poland, ex decayed grass. Type deposition. In the author’s collection.

271. Neomolgus pratensis Shiba, 1971: 93 ; Namekawa, Kawauchi-chô, Ehime pref., Japan.

Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan.

272. Neomolgus pygmaeus Shiba, 1969a: 74 ; Shiga Heights, Japan.

Distribution. Japan ( Shiba 1969a), China ( Lin et al. 2006).

Redescription. Shiba (1971).

Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan.

273. Neomolgus raeticus ( Schweizer & Bader, 1963) : 238 comb. nov.; NationalPark, Mt. Mezdi, Switzerland. Original designation: Hoplomolgus raeticus Schweizer & Bader.

Type deposition. NMB (NP 1845).

274. Neomolgus raptor Kuznetsov & Barilo, 1984: 934 ; near Samarkand (Tchapan-Atha), Uzbekistan, under stones.

Type deposition. NBG.

275. Neomolgus reticulatus ( Schweizer & Bader, 1963) : 241 comb. nov.; S-chanf, Switzerland. Original designation: Hoplomolgus reticulatus Schweizer & Bader.

Type deposition. NMB (NP 283).

276. Neomolgus sabulosus Shiba, 1971: 96 ; Shigenobu River, Matsuyama, Ehime pref., Japan.

Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan.

277. Neomolgus seriatus Shiba, 1971: 101 ; Otanomôsu-no-taira, Shigakôgen, Nagano pref., Japan.

Type deposition. Biological Laboratory of Matsuyama Shinonome Junior College, Matsuyama, Japan.

278. Neomolgus thorianus ( Berlese, 1923) : 239 comb. nov.; Corfu, Greece. Original designation: Molgus (Hoplomolgus) thorianus Berlese. Other name: Molgus (Molgus) thorianus, Thor, 1931a: 62 . Distribution. Greece (Corfu Island) ( Thor 1931a), Georgia ( Gomelauri 1963b). Type deposition. CRA.