Scopaeus borneensis Cameron, 1941

Frisch, Johannes & Narakusumo, Raden Pramesa, 2022, Revision of Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae, Paederinae) of Indonesia, with description of 19 new species, Soil Organisms 95 (1), pp. 23-73 : 62-67

Scopaeus borneensis Cameron, 1941

( Figs 26 , 180, 181 )

Scopaeus borneensis Cameron, 1941: 229 .

Type specimen examined: Holotype ♀, Kalimantan Barat, Pontianak; labeled ‘Type’ (round, printed label with red margin), ‘Pontianak / Borneo’ (handwritten), ‘S. / borneensis / TYPE Cam.’ (handwritten), ‘M. Cameron./ Bequest./ B.M.1955-147.‘ (printed), ‘ Holotype / Scopaeus borneensis / Cameron 1941 / label by J. Frisch, 2022 ’ (red, printed) ( NHML).

Cameron (1941: 229) described Scopaeus borneensis based on a ‘unique’ specimen. Thus, it is a holotype by monotypy ( ICZN, Article 73.1.2) .

Redescription: Habitus and coloring as in Fig. 26 . Head subquadrate. Penultimate antennal segment slightly elongate. Mesotibia moderately thickened. Body surface subnitid, finely, densely punctate, without microreticulation. Pubescence of body surface short, decumbent, without conspicuous macrosetae. Forebody reddish dark brown; abdomen and antennae medium brown; maxillary palpi and legs light brown. Total body length 3.8 mm; forebody length 2.4 mm.

Male: Unknown.

Female: Chamber segment of sperm pump short, without apophysis, acute-angled attached to distal process with strongly widened apex ( Fig. 181 ); bursal duct strong, with S-shaped portion gradually widened, then strongly narrowed towards sperm pump ( Fig. 180 ). Figures 134–145. Abdominal sternite VII (138, 140, 142) and abdominal sternite VIII (134 – 138, 139, 141, 143– 145) of ♂ of Scopaeus cuspilobatus , holotype, Jawa Barat, Sukabumi: Sirnarasa (134); S. velifer , holotype, Banten, Lebak: Citorek Kidul (135); S. elegantulus , Jawa Barat, Sukabumi, Sirnarasa: Cisarua (136); S. uncinatus , holotype, Banten, Lebak: Citorek Kidul (137); S. bipenicillatus , paratype, Sulawesi Tengah, Sigi: Pagana (138); S. bipenicillatus , holotype, Sulawesi Tengah, Sigi: Pagana (139); S. bipectenatus , holotype, Sulawesi Tengah, Poso: Lake Poso (140, 141); S. cuspidatus , holotype, Bali, Jembrana: Cekik (142, 143); S. crassipunctatus , holotype, Jawa Barat, Sukabumi, Sirnarasa: Cisarua (144); S. sulawesianus , holotype, Sulawesi Selatan: Engrekang (145).

Phylogeny: Scopaeus borneensis must be classified in the subgenus Hyperscopaeus Coiffait, 1984 (see Frisch et al. 2002: 38).

Distribution: This species is known only from the type locality in Kalimantan Barat, Indonesian Borneo.

Figures

Figures 23–26. Habitus of Scopaeus cuspidatus, paratype, Jawa Barat: Mt Salak (23); S. sulawesianus, paratype, Sulawesi Tengah: Tambarana (24); S. niger, paralectotype, Singapore (25); S. borneensis, holotype, Kalimantan Barat: Pontianak (26).

Figures 179–181. Sperm pump (179, 181) and bursal duct (180) of ♀ of S. niger, lectotype, Singapore (179); S. borneensis, holotype, Kalimantan Barat: Pontianak (180, with lateral gonocoxal plates; 181). Long scale bar: 179, 181; short scale bar: 180.