Cyana puer ( Elwes, 1890 )

Singh, Navneet, Volynkin, Anton V., Kirti, Jagbir Singh, Datta, Harvinder Singh & Ivanova, Maria S., 2020, A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini), Zootaxa 4738 (1), pp. 1-93: 13

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Cyana puer ( Elwes, 1890 )


Cyana puer ( Elwes, 1890)  

( Figs 22–24 View FIGURES 19–27 , 171 View FIGURES 170–172 , 244 View FIGURES 240–245 )

Bizone puer Elwes, 1890   ; Proceedings of the general meetings for scientific business of the Zoological Society of London 1890: 392, pl. 32, fig. 8 (Type locality: “near Darjeeling …, from the Khasia Hills, … Mao on the Manipur side of the Naga Hills, … from Assam ”).

Type material examined. Lectotype (designated herein) ( Fig. 22 View FIGURES 19–27 ): ♂, printed label “Darjeeling. 20 July 1886. H.J. Elwes. ” / handwritten label “ Bizone puer Elwes   Type” / round printed label with red circle “Type” / “ Collectio H.J. Elwes ” / “Rothschild Bequest B.M. 1939-I ” / printed label with a unique identifier “NHMUK010402104” (Coll. NHMUK).  

Other material examined. 217 ♂ and 39 ♀ from India, Nepal and North West Myanmar ( Colls MWM / ZSM, NHMUK and ZMB) listed by Singh et al. (2019a). MEGHALAYA   : 1 ♂, India, Meghalaya, Pynursla , 11.IX.2014, leg. H.S. Datta (Coll. NZCZSI)   ; 1 ♂, India, Meghalaya, Cherrapunji , 13.IX.2014, leg. H.S. Datta (Coll. NZCZSI)   ; 4 ♂, India, Meghalaya, Jowai , 17.IX.2015, leg. Kumar Kaustubh (Coll. NZCZSI)   ; SIKKIM: 1 ♂, India, Sikkim, Okhrey , 24.IX.2014, leg. H.S. Datta (Coll. NZCZSI)   ; ARUNACHAL PRADESH: 1 ♀, India, Arunachal Pradesh, Tippi , 16.V.2016, leg. H.S. Datta (Coll. NZCZSI)   .

Remark. The species was described based on five specimens of both sexes from several localities in India ( Elwes 1890). Singh et al. (2019a) erroneously cited the male syntype deposited in NHMUK as ‘Holotype’. As C. puer   has no external differences from C. neopuer   , to avoid a probable confusion in the future, here we designate this specimen as lectotype.

Diagnosis. Forewing length is 17–19 mm in males and 20–21 mm in females. Externally, the species is polymorphic: colour of the transverse bands of forewing varies from crimson to orange (the orange lined form is illustrated by Singh et al. (2019a)). The width and shape of transverse lines and the size of discal spots also vary significantly. In the male genitalia, the shape of uncus tip and the width of distal section of valva are variable, but the vesica structure is stable and diagnostic. Cyana puer   has no external differences from its closest relative C. neopuer   and can be distinguished by the genitalia structures only. The differences between the species are listed in the diagnosis of C. neopuer   .

Distribution. North East India (Arunachal Pradesh, Assam, Manipur, Meghalaya, Nagaland, Sikkim and West Bengal) ( Elwes 1890, Hampson 1894, Holloway 2001, Kirti & Singh 2015; Singh et al. 2019a), Nepal ( Kishida 1994, as Cyana dohertyi   ; Kishida 2000; Holloway 2001; Singh et al. 2019a) and North West Myanmar (Chin State) ( Singh et al. 2019a). The record for China (Xizang) ( Fang 2000) needs clarification.


Natural History Museum, London


Bavarian State Collection of Zoology


Museum für Naturkunde Berlin (Zoological Collections)














Cyana puer ( Elwes, 1890 )

Singh, Navneet, Volynkin, Anton V., Kirti, Jagbir Singh, Datta, Harvinder Singh & Ivanova, Maria S. 2020

Bizone puer

Elwes 1890