Cyana khasiana Hampson, 1897

Singh, Navneet, Volynkin, Anton V., Kirti, Jagbir Singh, Datta, Harvinder Singh & Ivanova, Maria S., 2020, A review of the genus Cyana Walker, 1854 from India, with descriptions of five new species and three new subspecies (Lepidoptera: Erebidae: Arctiinae: Lithosiini), Zootaxa 4738 (1), pp. 1-93: 43

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Cyana khasiana Hampson, 1897


Cyana khasiana Hampson, 1897  

( Figs 112–117 View FIGURES 112–119 , 210–213 View FIGURES 210–213 , 273, 274 View FIGURES 273–276 )

Cyana khasiana Hampson, 1897   , Journal of the Bombay Natural History Society 11: 296 (Type locality: [NE India, Meghalaya, Khasi Hills] “Khásis”).

= Chionaema hampsoni Kaleka, 2003   , Entomon, 28 (4): 380, figs 1–7 (Type locality: “ Arunachal Pradesh: West Kameng District, Bombdila, 2430 m ”), syn. nov.

Type material examined. Lectotype (designated herein): 1 ♂ ( Fig. 112 View FIGURES 112–119 ), printed label “Khasis. Nat. Coll.” / hand- written label “96-43 (upper side) Cyana khasiana   type ♂. Hmpsn. (under side)” / printed round label with a red circle “Type” / printed label with a unique identifier “NHMUK010597911” (Coll. NHMUK); paralectotypes: 2 ♀, printed label “Khasis. Nat. Coll.” (Coll. NHMUK).  

Other material examined. SIKKIM: 1 ♂, India, Sikkim, Pemayangtse, 2000m, 23–28.VII.1990, leg. Dr. W. Thomas, slide MWM 34441 View Materials (♂) Volynkin (Coll. MWM / ZSM)   ; 4 ♂, 2 ♀, India-Sikkim, Pemayangtse, 1200 m, be- low Rimbi, 26.VIII.1988, Lf., leg. Dr. W. Thomas, slide MWM 34442 View Materials (♀) Volynkin (Coll. MWM / ZSM)   ; 1 ♂, Sik- kim, Golitar, 26.IV.2009, leg. Rahul Joshi (Coll. NZCZSI)   ; 3 ♂, Sikkim, Golitar, 30.IV.2014, leg. H.S. Datta (Coll. NZCZSI)   ; 1 ♂, Sikkim, Golitar, 01.V.2014, leg. H.S. Datta (Coll. NZCZSI)   ; WEST BENGAL: 2 ♀, India WB, Darjeeling, 2100 m, 28.VIII.–1.IX.1988, leg. W. Thomas (Coll. MWM / ZSM)   ; 1 ♀, India W.B., Darjeeling, above Rambi, ca. 800 m, 7–8.VIII.1989, leg. Dr. W. Thomas (Coll. MWM / ZSM)   ; ASSAM: 1 ♀, Assam (Coll. NHMUK)   ; MEGHALAYA: 1 ♂, Khasis. Nat. Coll. / Rothschild Bequest B.M. 1939–1 / 448, slide NHMUK010315750 (♂) Volynkin (Coll. NHMUK)   ; 1 ♂, 1 ♀, Khasis. Nat. Coll. / Collectio H. J. Elwes / Rothschild Bequest B.M. 1939– 1, slides NHMUK010315751 (♂), NHMUK010313269 (♀) Volynkin (Coll. NHMUK)   ; 1 ♀, Khasis. Nat. Coll. / 97.164, slide NHMUK010315752 (♀) Volynkin (Coll. NHMUK)   ; 1 ♀, Assam: Shillong, H.M. Parish, B.M. 1923- 247 (Coll. NHMUK)   ; 1 ♂, Khasis. Nat. Coll. / Collectio H. J. Elwes / round label with a yellow circle “ paratype ” / Chionaema khasiana Hampson   det. A. Watson 1962, slide AV1877 ♂ Volynkin (Coll. ZFMK)   ; 1 ♂, Meghalaya, Umkiang, 8.IX.2014, leg. H.S. Datta (Coll. NZCZSI)   ; ARUNACHAL PRADESH: 1 ♂, Arunachal Pradesh, Go- dak, 05.IV.2009 leg. Rahul Joshi (Coll. NZCZSI)   .

Remarks. 1. Hampson (1897) described the species based on an unknown number of syntypes of both sexes. Three of them are found in the NHMUK collection. In order to stabilize the nomenclature, here we designate the male specimen labeled as “Type” as lectotype. 2. All syntypes of various Lithosiini species received by ZFMK as a result of exchange with NHMUK are labeled as paratypes. However, the male specimen of C. khasiana   deposited in the ZFMK collection and labeled as “ Paratype ” is not a syntype in fact.

Diagnosis. Forewing length is 16–20 mm in males and 20.5–24 mm in females. The species is represented by two color forms: the widespread red lined one (described as hampsoni Kaleka, 2003   ) ( Figs 114–117 View FIGURES 112–119 , 212, 213 View FIGURES 210–213 , 274 View FIGURES 273–276 ) and the typical form with amber yellow transverse lines found in Khasi Hills only ( Figs 112, 113 View FIGURES 112–119 , 210, 211 View FIGURES 210–213 , 273 View FIGURES 273–276 ). In addition, in males distal black spots may be connected ( Figs 112, 114, 116 View FIGURES 112–119 ) or separated ( Fig. 117 View FIGURES 112–119 ) as well. This variation is found in both color forms. The yellow lined form of C. khasiana   resembles externally C. arama   , but differs by its narrower antemedial line being almost straight and only slightly curved at wing costa (that is strongly S-like curved in C. arama   ), smaller discal black spots, narrower postmedial line, the absence of a subterminal line, and the pure white hindwing (that is pale ochreous orange in C. arama   ). The red lined form is similar to C. signa   and C. adita   , but can be distinguished by their slightly broader forewings with broader transverse lines and larger black spots; in addition, females of C. khasiana   have the postmedial line curved inwards at costa, whereas in C. signa   and C. adita   that is curved outwards. In the male genitalia, C. khasiana   differs clearly from C. arama   by its narrower and U-shaped uncus, less developed medial crest of valva, broader distal section of valva, slightly shorter and broader distal saccular process, narrower aedeagus and the vesica shape. Female genitalia of C. khasiana   differ from those of C. arama   by the smaller anterior section of the corpus bursae, the smaller signum, the sclerotized basal section of the appendix bursae, and the much larger distal membranous section of the appendix bursae.

Distribution. North East India (Sikkim, Assam and Meghalaya) ( Singh et al. 2014), Arunachal Pradesh, north of West Bengal (present study), China (Sichuan and Yunnan Provinces) (present study), North Thailand ( Černý & Pinratana 2009, as a part of C. signa   ) and North Vietnam (Fan-si-pan Mts.) (present study).


Natural History Museum, London


Bavarian State Collection of Zoology


Zoologisches Forschungsmuseum Alexander Koenig














Cyana khasiana Hampson, 1897

Singh, Navneet, Volynkin, Anton V., Kirti, Jagbir Singh, Datta, Harvinder Singh & Ivanova, Maria S. 2020

Chionaema hampsoni

Kaleka 2003

Cyana khasiana

Hampson 1897