Notsodipus, PLATNICK, 2000

PLATNICK, NORMAN I., 2000, A Relimitation And Revision Of The Australasian Ground Spider Family Lamponidae (Araneae: Gnaphosoidea), Bulletin of the American Museum of Natural History 2000 (245), pp. 1-328 : 269-271

Notsodipus , new genus

TYPE SPECIES: Notsodipus dalby , new species.

ETYMOLOGY: The generic name is an arbitrary combination of letters, considered masculine in gender.

DIAGNOSIS: Members of this genus resemble species of Asadipus in having a relatively long and sinuous embolus, and equally long and sinuous epigynal ducts, but can be distinguished by the unexcavated retrolateral tibial apophysis of males (figs. 651, 663) and the deep anterior epigynal atrium and highly coiled posterior ducts of females (figs. 652, 653).

DESCRIPTION: Small to medium­sized spiders, total length of males 1.8–5.0, of females 1.9–5.7. Carapace dark orange, coated with large tubercles bearing long, erect setae, margins with long, dark setae; thoracic groove short, almost circular, deep. Eight eyes in two rows, posterior medians largest, other subequal, anterior medians circular, dark, laterals almost circular, light, posterior medians irregularly oval, flattened, light; from above, anterior eye row slightly recurved, posterior row slightly procurved, from front, both rows procurved; anterior medians separated by more than their radius, closer to anterior laterals; posterior medians separated by less than their radius, much farther from posterior laterals; anterior and posterior laterals separated by about their radius; median ocular quadrangle slightly wider in back than in front, about as long as wide in back. Chelicerae, sternum, and mouthparts orangish red; chilum wide, triangular, short, accompanied by second, I­shaped, posterior chilum (extremely narrow sclerite separating bases of chelicerae posteriorly); chelicerae with distinct lateral boss and scattered tubercles bearing short, stiff setae, anterior surface with depressed, relatively unsclerotized oval area near promargin; promargin with row of long setae originating in line along base of fang, seta closest to fang bent near base at almost 908 angle, extending behind other promarginal setae toward midline; promargin with three teeth, median one largest, proximal one smallest; retromargin with two well separated teeth. Labium elongate, base narrowed at about one­fifth of labial length, posterior margin procurved, anterior margin with slight medial invagination, surface not depressed medially. Endites obliquely depressed, with sharply demarcated, deep groove along margin near labium, groove wider anteriorly than posteriorly; serrula long, with single row of teeth; anterior surface with distinct sieve plate of maxillary gland openings set in oval of unsclerotized cuticle. Sternum slightly elevated, with inclined lateral margins, not expanded anteriorly, with slight triangular extensions to and longer extensions between coxae; surface not tuberculate, with only slight elevations opposite, and depressions between, coxae. One epimeric sclerite on each side, above each coxa, extending toward but not fused with sternal triangles, not fused to carapace or to sclerite surrounding pedicel. Pedicel consisting of large, diamond­shaped sclerite with narrow anterior margin meeting posterior tip of sternum at slightly procurved line.

Anterior edge of abdomen of male with complete sclerotic ring formed by strong epigastric scutum separated from strong dorsal abdominal scutum covering almost entire dorsal surface of abdomen, females with dorsal scutum represented only by small, almost circular plate above pedicel, extending slightly on to abdominal dorsum (except in N. keilira , where extending over length of abdominal dorsum); cuticle with small tubercles bearing long, erect setae; epigastric scutum accompanied posterolaterally by pair of oval, deeply invaginated sclerites bearing clearly elevated anterior rim; sclerites separated by membranous lobe, anterior rim of sclerites fitting under epigastric scutum; colulus represented only by pair of setae; tiny transverse sclerite, well removed from spinnerets, marking position of small posterior spiracle. Anterior lateral spinnerets tubular, separated by less than their diameter, cuticle representing distal, second spinneret segment restricted to semicircle surrounding major ampullate gland spigots (piriform gland spigots surrounded only by soft cuticle); poste­ rior median spinnerets of males large, tubular, of females with anteriorly expanded tips, bases occupied by three enormously widened cylindrical gland spigots; posterior lateral spinnerets two­segmented, those of females with two greatly widened cylindrical gland spigots.

Leg spination greatly reduced; typical leg spination pattern (only surfaces with spines listed): tibiae: III v0­0­1p; IV v0­0­2. Most leg surfaces coated with long setae; males with all coxae and trochanters dorsally tuberculate; anterior coxae with slightly protuberant posterolateral corners; trochanters unnotched; anterior metatarsi and tarsi without scopulae; posterior metatarsi with thick, distal preening brushes; tarsi III, IV without scopulae; tarsi with two dentate claws, claw tufts composed of lateral pads of closely appressed setae; trichobothria present on tibiae, metatarsi, and tarsi. Female palpal tibia and tarsus with long, thin, prolateral spines; female palpal tarsus with long, basally dentate claw.

Male palp with long, unexcavated retrolateral tibial apophysis; cymbial surface flattened opposite tibial apophysis; tegulum occupying retrolateral surface of bulb, produced into lobe medially, bearing large, elaborate terminal apophysis; embolus originating prolaterally, long, looping around prolateral, distal, and proximal sides of bulb; median apophysis absent. Epigynum flattened, with conspicuously excavated anterior atrium; spermathecal ducts long, sinuous.

KEY TO SPECIES OF NOTSODIPUS

1. Males (those of N. blackall unknown)... 2

– Females (those of N. broadwater unknown) ............................... 17

2. Ventral portion of terminal apophysis terminating in narrow, sharp point (as in figs. 650, 654)........................ 7

– Ventral portion of terminal apophysis not terminating in narrow, sharp point (figs. 666, 670, 678, 706, 710)................ 3

3. Ventral portion of terminal apophysis relatively broad at tip (figs. 666, 670).... 4

– Ventral portion of terminal apophysis relatively narrow at tip (figs. 678, 706, 710) ................................ 5

4. Tip of terminal apophysis with three points (fig. 670).................. magdala

– Tip of terminal apophysis with single point (fig. 666).................... marun

5. Embolus relatively long, looping to proximal half of tegulum (fig. 678)..... renmark

– Embolus relatively short, restricted to distal half of tegulum (figs. 706, 710)...... 6

6. Embolus relatively wide at base (fig. 706).............................. keilira

– Embolus relatively narrow at base (fig. 710)........................... domain

7. Tegulum with large proximal apophysis (figs. 702, 703)..................... visio

– Tegulum without proximal apophysis.... 8

8. Tip of terminal apophysis bifid (fig. 674).......................... broadwater

– Tip of terminal apophysis not bifid..... 9

9. Embolus very long, making extra loop proximally (fig. 662)............. muckera

– Embolus shorter, not making proximal loop ............................... 10

10. Dorsal prong of terminal apophysis forming tube­shaped structure behind ventral prong (figs. 650, 654, 658).............. 11

– Dorsal prong of terminal apophysis otherwise ............................... 13

11. Retrolateral tibial apophysis abruptly narrowed at tip (figs. 655, 659)........ 12

– Retrolateral tibial apophysis gradually narrowed toward tip (fig. 651)...... dalby

12. Tegular bulge situated near anterior edge of tegulum (fig. 654).............. innot

– Tegular bulge situated farther posteriorly (fig. 658)....................... upstart

13. Dorsal prong of terminal apophysis with prolaterally directed spur near base (fig. 686)............................ barlee

– Dorsal prong of terminal apophysis without prolaterally directed spur........... 14

14. Retrolateral tibial apophysis relatively wide at base (figs. 683, 691).............. 15

– Retrolateral tibial apophysis relatively narrow at base (figs. 695, 699)............ 16

15. Tegulum relatively narrow, embolus relatively short (fig. 682)........... capensis

– Tegulum relatively wide, embolus relatively long (fig. 690)................ meedo

16. Retrolateral tibial apophysis relatively short (fig. 695)................... quobba

– Retrolateral tibial apophysis relatively long (fig. 699).................. bidgemia

17. Epigynum with pair of lateral pockets and anteromedian septum (fig. 712)... domain

– Epigynum without lateral pockets..... 18

18. Epigynum relatively long, narrow (figs. 592, 704, 708)....................... 19

– Epigynum not narrow and elongated (as in figs. 652, 692)................... 21 19. Epigynal atrium situated medially (fig. 708)............................ keilira

– Epigynal atrium situated anteriorly (figs. 591, 704)........................... 20

20. Epigynal ducts extending to anterior half of epigynum (figs. 592, 593)...... blackall

– Epigynal ducts restricted to posterior half of epigynum (figs. 704, 705)........ visio

21. Epigynal atrium relatively small (figs. 664, 692)........................... 22

– Epigynal atrium relatively large (as in figs. 652, 660)....................... 23

22. Epigynal atrium very short (fig. 692).................................. meedo

– Epigynal atrium longer (fig. 664)... muckera

23. Epigynal atrium relatively short (figs. 652, 660)........................... 24

– Epigynal atrium longer (as in figs. 680, 700) ............................... 25

24. Epigynal atrium wider than long (fig. 660)........................... upstart

– Epigynal atrium longer than wide (fig. 652)............................ dalby

25. Epigynal atrium widest anteriorly or medially ............................... 27

– Epigynal atrium widest posteriorly (figs. 668, 688)........................... 26

26. Lateral epigynal ducts relatively long (figs. 668, 669).................... marun

– Lateral epigynal ducts relatively short (figs. 688, 689).................... barlee

27. Epigynal atrium widest anteriorly (figs. 680, 696, 700)....................... 28

– Epigynal atrium widest at middle (figs. 656, 672, 684)....................... 30

28. Epigynal atrium relatively long (fig. 696)............................. quobba

– Epigynal atrium relatively short (figs. 680, 700)........................... 29

29. Epigynal atrium relatively small (fig. 680)............................ renmark

– Epigynal atrium relatively large (fig. 700)............................ bidgemia

30. Epigynal atrium relatively short (fig. 656)............................... innot

– Epigynal atrium relatively long (figs. 672, 684)........................... 31

31. Lateral epigynal ducts relatively long (figs. 672, 673).................. magdala

– Lateral epigynal ducts relatively short (figs. 684, 685................... capensis