Eumanota parahumeralis, Papp, 2004

Papp, L, 2004, Seven New Species Of Manotinae (Diptera: Mycetophilidae) From Asia And Papua New Guinea, Acta Zoologica Academiae Scientiarum Hungaricae 50 (3), pp. 227-244 : 229-232

Eumanota parahumeralis sp. n.

( Figs 5–8 )

Holotype: male. TAIWAN, Fu-Shan Botanical Garden , 700 m – along a forest path, Sep 25, 2000, leg. L. Papp, No. 3. ( HNHM, hind tarsomere 2–5 lost.)

Paratypes: 1 male ( HNHM, right 2 apical flagellomeres, left mid tarsus and hind tarsomeres 2–5 as well as right mid tarsomeres 2–5 lost): TAIWAN, Nantou Hsien: Suili – forest undergrowth, Sep 30, 2000, leg. L. Papp, No. 12 ; 1 male ( NMNS, wings greasy and wrinkled, formerly prepared from alcohol, consequently most of setae lost, left mid tarsomeres 2–5 and right mid tarsomeres 3–5 lost): TAIWAN, Nantou, Jenai Chunyang , VIII-IX/12–8 /1998, C.S. Lin & W. T. Yang, Malaise trap – NMNS ENT 3028–178 .

Measurements in mm: body length 4.15 (holotype), 4.10, 3.95 (paratypes), wing length 3.30 (holotype), 3.30, 3.20, wing breadth 1.34 (holotype), 1.24 and not measurable.

Scutum brown, lateral parts of mesonotum, notopleura and pleura (mainly) ochre with dorsal part of anepisternum and anepimeron diffuse brown, whole laterotergite and lateral parts of mediotergite brown.

Head 0.595 mm high, 0.495 mm long, yellow, medial part of occiput fuscous brown. Face not only broad (0.20 mm on holotype) but with numerous thick black setae. Lateral ocelli close to eye margin, distance only 0.03 mm. Complete row of long postocular setae present. Flagellum of holotype ca. 2.00 mm long (slightly curved, i.e. not precisely measurable. Apical flagellomere 0.198 mm long, 0.05 mm broad, penultimate one 0.138 mm long, 0.06 mm broad. Palpus whitish, not precisely measurable but longer than fore coxa, i.e. longer than 1.55 mm.

Length of thorax 1.32 mm (holotype). Anepisternum with small setae on anterior and dorsal parts and with 4 very long setae, laterotergite with long setae, particularly caudally and dorsally.

Wing light brown, veins darker brown. Wing membrane with perpendicular short dorsal macrotrichia in m1, m2 and m3 cells distally, as well as similar but more numerous macrotrichia on anal region. Length of M stalk 0.20 mm (holotype), length of Sc vein 0.22 mm, R 1 0.65 mm, R 2 0.072 mm, R-M 0.77 mm. Stalk of halteres light brown, knob fuscous dark brown.

moved). Scales: 0.4 mm for Figs 5–6 , 0.2 mm for Figs 7–8

Legs yellow, incl. coxae, apices of coxae brown. Length of femur, tibia, first tarsomere and tibial spurs (in mm, measured on holotype): fore: 1.25, 0.835, 0.88, 0.53; mid: 1.385, 1.63, 0.935, 0.65, 0.41; hind: 1.54, 2.30, 1.10, 0.68, 0.44.

Abdomen dark brown, sternites 1–5 lighter. Cerci and entire proctiger white. Tergite 9 much narrowed caudally ( Fig. 5 ), though caudal edges rounded, width caudally less than half of subbasal width. Tergite 9 with several long and thick setae. Cerci broad in dorsal view, covered by setulae of even thickness. Gonocoxites fused in their 1/2 ventrally (sagittally) ( Fig. 6 ), their mediodorsal process thin, with a blunt medioventral and very large, subtriangular lateral processes (lobes) ( Fig. 7 ). Gonostylus blunt apically, with a blunt medial apex. Gonostylus covered by dense short setae only. Aedeagus long, apex narrow; parameres short with lateroclinate apical setae ( Fig. 8 ).

Female unknown.

E. parahumeralis sp. n. is close to E. humeralis EDWARDS, 1933 known from Malaysia, Sabah, Mt Kinabalu. The medially directed gonostylar tip is blunt in E. parahumeralis (sharp in humeralis ) and its length / transversal breadth ratio is also different (longer in parahumeralis ). The shape of gonostylus also resembles that of E. racola [author and year is already given in the list of species in the introduction]. However, gonocoxite of E. parahumeralis has a pair of blunt medioventral and a pair of large lateral subtriangular processes caudally (latter ones are easily seen without preparation). If SØLI’ s (2002) fig. 8 depicts the lateral lobe of E. humeralis correctly, the difference in this respect is massive and most distinctive. If the tip of that process was broken (to which there is no reference in the paper), the extent of difference is unknown. It is certain that the difference in the shape of medial processes between the two species is distinct. The cerci of the new species seem broader ( Fig. 5 , cf. fig. 10 of SØLI) and the bare basal area of tergite 9 is larger.

Figures

Figs 5–8. Eumanota parahumeralis sp.n., paratype male, genitalia: 5 = tergite 9 and proctiger, dorsal view, 6 = gonocoxites and right gonostylus, ventral view, 7 = right gonocoxite and gonostylus, ventral view, 8 = right gonocoxite and gonostylus with genitalia, dorsal view (tergite 9 and proctiger re-