Anomala kanshireiensis Zorn & Zhao, 2022

Zhao, Ming-Zhi & Zorn, Carsten, 2022, Contribution to the genus Anomala Samouelle, 1819 (Coleoptera: Scarabaeidae Rutelinae) of China and adjacent regions. Part II: six new species from Taiwan and Hainan, Zootaxa 5168 (2), pp. 175-195 : 184-187

Anomala kanshireiensis Zorn & Zhao , new species [关T岭异fflŵoi/ṞTĂ條ŵff]

( Figs. 6 , 8E–F , 9H–J, 9N–O )

Type material. Holotype: ♂ ( MFNB), Formosa, Sauter, Kanshirei [= Guanziling in Tainan City, ṞTĂ], 908. V. 19-27 . Paratypes: 1♂, 3♀♀ ( MFNB), same data as holotype ; 3♂♂ ( MFNB), Formosa , Sauter, Kanshirei, 908. V. 17-24 .; 1♂ ( MFNB), Formosa , Sauter, Kanshirei, 908. V. 1-9 ; 1♂ ( MFNB), Formosa , Sauter, Kanshirei, 908. V. 19-26 .; 3♂♂, 1♀ ( SMNS), TAIWAN, Kanshirei , 2. - 14. 6. 1908, H. Sauter leg. ; 1♂ ( HNHM), Formosa , Sauter, Kanshirei, 908. V-1-2-14 .

Description. Holotype male ( Figs. 6A–C , 8E , 9K–M ). Body length: 11.8 mm, greatest width: 6.7 mm.

General appearance. Body shape elongated ovoid, weakly convex. Entirely dark reddish brown, with weak green metallic sheen.

Head ( Fig. 8E ). Clypeus subtrapezoidal, anterior corner broadly rounded; anterior margin somewhat rounded and strongly reflexed; surface densely rugopunctate. Frontoclypeal suture distinct. An inverted triangular area at anterior half of frons densely rugopunctate, other portions of frons and vertex with irregularly distributed dense small punctures. Antennal club longer than antennomeres 2–6 combined. Inner margin of eye with several moderately long setae (mostly worn in the holotype).

Pronotum. Sides gently arched and convergent anteriad. Anterior angle acute and protruding, posterior angle obtuse, rounded off. Basal marginal line interrupted before scutellum, other marginal lines complete. With dense small punctures, punctures being partly coalescent at marginal portions; with a very weakly indicated longitudinal medial furrow not reaching the base. Lateral margin with several long setae, two moderately long to long semierect setae on disc of pronotum.

Scutellum. Subtriangular, lateral margin arched. Margins with polished appearance, disc with irregularly distributed moderately large punctures. Disc with two semierect long setae.

Elytra. Intervals strongly convex; strial punctures dense to very dense, annulated and large, punctures of secondary striae unequally spaced; the whole surface with scattered minute punctures. Interstices I to III each with a secondary stria; interstice I (the subsutural interstice) broadest, with secondary stria shortly doubled in basal fourth; the other secondary striae reaching the level of apical protuberance and usually interrupted, secondary stria of interstice II obsolete between level of scutellar apex to midway of elytron. Primary costae without additional striae. Humeral umbone and apical protuberance moderately bulging. Lateral carina indistinct. Epipleura with a row of moderately dense long setae. Marginal membrane complete.

Propygidium. With dense transverse punctures, usually coalescent into transverse striolation; glabrous.

Pygidium. Distinctly bulging, posterior margin protruding. Disc with dense ovoid large punctures, becoming striate near all margins; punctures encircling greatest tumidity. With several short setae distributed along posterior margin, posterior margin with a row of long setae near apex.

Ventral thoracic surface. Hypomeron with dense coalescent striolation and moderately dense long setae. Collar of mesosternite transversely striolate with dense recumbent short setae. Other portions of mesosternite with small punctures and somewhat rugopunctate, with dense short setae. Ventral metathoracic surface with dense annulated moderately large punctures and dense, rather long setae; with minute punctures and almost glabrous medially.

Abdominal ventrites. Ventrites 1–2 and anterior half of 3 weakly carinate laterally. Ventrites 2–5 with irregularly distributed small punctures, punctures coalescent into striolation in ventrite 6. Ventrites 2–5 each with a transverse row of sparse and moderately long setae; ventrite 6 with a complete row of long setae along posterior margin.

Legs. Protibia tridentate, apical and second tooth acute at apex, proximal tooth small; apical tooth extending to level of halfway of protarsomere 2. Inner spur inserted at the level of the indention between proximal and second tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branches longer and wider, the lower branch of inner protarsal claw concave internobasally. Each tarsomere 5 with an internomedial protuberance. Outer metatarsal claw slightly longer than inner one. Mesofemoral surface with transverse rows of long setae, several irregular rows between two regular rows, one at anterior margin and another situated behind middle, the row at posterior margin usually with shorter setae. Metafemoral surface with three transverse rows of sparse long setae, the first row at anterior margin, a second row situated behind middle with robuster setae, and a third row at posterior margin with short setae.

Male genitalia. See Fig. 9H–J . Parameres asymmetric, right paramere longer. Ventral plate flat, apex acute.

Paratypes. Males. Body length: 11.5–13.0 mm, greatest width: 6.5–7.5 mm. External features and shape of parameres consistent. Semierect longer setae on scutellum and disc of pronotum are only present in few males but might be worn off in other specimens. Females ( Figs. 6D–F , 8F ). Body length: 12.3–13.0 mm, greatest width: 7.0– 7.9 mm. Generally similar to male. Antennal club almost equally as long as antennomeres 2–6 combined. Sides of pronotum more strongly curved. Pygidium more bulging, shorter than in male. Protibia and protarsomeres thinner than in male; apical protibial tooth tongue-shaped, almost extends to level of apex of protarsomere 3; the other two teeth blunt at apex, proximal tooth indistinct; inner spur situated at the level of the proximal tooth; internomedial protuberance of protarsomere 5 small; upper and lower branches of inner protarsal claw and outer mesotarsal claw almost equal in length. Ventral metathoracic surface less setose than in male. Semierect longer setae on scutellum and disc of pronotum are only present in the minority of females but might be worn off in some specimens.

Differential diagnosis. Anomala kanshireiensis Zorn & Zhao , new species is most similar to A. inclinata Zhao & Zorn , new species. Above all, the general body color is reddish brown in A. kanshireiensis but blackish brown in A. inclinata . The interrupted secondary stria of interstice II is more complete than in A. inclinata . The aedeagi of the two species are similar in shape. The left paramere is shorter and has a less curved apex compared to A. inclinata . The left paramere is placed slightly above the right paramere in lateral view (parameres are overlapping in A. inclinata ). The apex of the ventral plate is acute in A. kanshireiensis but arched in A. inclinata . The endophallites of the two species show considerable differences: the two dorsal endophallites are spaced with one being relatively long in A. kanshireiensis ( Fig. 9N–O ), they are adjacent and both short in A. inclinata ( Fig. 9P–Q ); the ventral endophallite is a curved spine in A. kanshireiensis , while it is a serrated sclerite in A. inclinata .

Etymology. The specific epithet is alluding to the historical name of the type locality, Kanshirei.

Distribution. Taiwan Island (Tainan).

Figures

FIGURE 6. Habitus of Anomala kanshireiensis Zorn & Zhao, new species. A–C, holotype; D–F, female paratype. A, D, dorsal view; B, E, lateral view; C, F, ventral view. Scale bar = 5 mm.

FIGURE 8. Head of Anomala spp. A, A. linwenhsini Zhao & Zorn, new species, holotype; B, ditto, female paratype; C, A. wutaiensis Zhao & Zorn, new species, holotype; D, ditto, female paratype; E, A. kanshireiensis Zorn & Zhao, new species, holotype; F, ditto, female paratype; G, A. inclinata Zhao & Zorn, new species, holotype; H, ditto, female paratype. Scale bar = 5 mm.

FIGURE 9. Male genitalia (A–M) and everted endophallus (N–Q) of Anomala spp. A–C, A. linwenhsini Zhao & Zorn, new species, holotype; D–G, A. wutaiensis Zhao & Zorn, new species, holotype; H–J, N–O, A. kanshireiensis Zorn & Zhao, new species, holotype; K–M, P–Q, A. inclinata Zhao & Zorn, new species, holotype. A, D, H, K, dorsal view; B, F, I, L, N, P, right lateral view; E, O, Q, left lateral view; C, G, J, M, ventral view. Sc 1, scale bar for A–M = 2 mm; Sc 2, scale bar for N–Q = 5 mm.