CETACEA Brisson, 1762

Bouetel, Virginie & Muizon, Christian de, 2006, The anatomy and relationships of Piscobalaena nana (Cetacea, Mysticeti), a Cetotheriidae s. s. from the early Pliocene of Peru, Geodiversitas 28 (2), pp. 319-395 : 383-385

Order CETACEA Brisson, 1762 Infraorder MYSTICETI Cope, 1891

Infraorder ARCHAEOCETI Flower, 1883 Superfamily AETIOCETOIDEA

Family BASILOSAURIDAE Cope, 1868 Sanders & Barnes, 2002

Subfamily BASILOSAURINAE Cope, 1868 Family AETIOCETIDAE Emlong, 1966 Subfamily AETIOCETINAE Emlong, 1966

Genus Zygorhiza True, 1908

Genus Aetiocetus Emlong, 1966

TYPE SPECIES. — Basilosaurus kochii Reichenbach,

1847. TYPE SPECIES. — Aetiocetus cotylalveus Emlong, 1966 .

Zygorhiza kochii (Reichenbach, 1847)

Aetiocetus cotylalveus Emlong, 1966

HOLOTYPE. — Geologisch-Paläontologisches Institut HOLOTYPE. — USNM 25210, sub-complete skull, both and Museum der Universität Berlin no. 15324, partial auditory areas and post-cranial elements .

skull. LOCALITY AND HORIZON. — Yaquina Formation, Lincoln

LOCALITY AND HORIZON. — Clark County, Alabama, County, Oregon, USA, latest Oligocene.

USA, late Eocene. REFERENCES. — Barnes et al. 1994; Emlong 1966;

REFERRED SPECIMEN. — USNM 11962. Sanders & Barnes 2002.

REFERENCE. — Kellogg 1936.

Subfamily DORUDONTINAE Miller, 1923

Genus Dorudon Gibbes, 1845

TYPE SPECIES. — Dorudon serratus Gibbes, 1845 . Subfamily CHONECETINAE Barnes, Kimura, Furusawa & Sawamura, 1994

Genus Chonecetus Russel, 1968

TYPE SPECIES. — Chonecetus sookensis Russel, 1968 .

Dorudon atrox (Andrews, 1906) Chonecetus goedertorum

Barnes & Furasawa, 1994

HOLOTYPE. — CGM 9319, cranium with right ramus of the lower jaw. HOLOTYPE. — LACM 131146 , partial skull, dentary and post-cranial elements .

LOCALITY AND HORIZON. — Birket Qarun Formation, Fayum, Egypt, Bartonian to early Priabonian. LOCALITY AND HORIZON. — Pysht Formation, Clallam County, Washington, USA, latest Oligocene.

REFERRED SPECIMENS. — Those figured by Uhen 2004. REFERENCE. — Barnes et al. 1994. REFERENCE. — Uhen 2004. REMARKS As Ichishima (1997) mentioned it, the systematics of fossil mysticetes is far from clear in some cases

(e.g., Cetotherium ...). This is due to the fact that some taxa “are represented by non-diagnostic skel-

etal elements”. According to this, and in order to minimize the number of missing data and to avoid

Bouetel V. & Muizon C. de problems of uncertain taxa, when possible, only LOCALITY AND HORIZON. — Astoria Formation, Newtype species were used in the present study. port, Oregon, USA, Miocene. REFERRED SPECIMENS. — (1) USNM 13646, (2) CMUO

305, skull studied from Packard & Kellogg 1934.

Infraorder MYSTICETI Cope, 1891 REFERENCE. — Packard & Kellogg 1934.

Genus Aglaocetus Kellogg, 1934

Genus Diorocetus Kellogg, 1968

TYPE SPECIES. — Cetotherium moreni Lydekker, 1894 . TYPE SPECIES. — Diorocetus hiatus Kellogg, 1968 .

Aglaocetus patulus (Kellogg, 1968) Diorocetus hiatus Kellogg, 1968

HOLOTYPE. — USNM 23690, sub-complete skull (lacking both lachrymals and jugals, and pterygoids), both HOLOTYPE. — USNM 16783, sub-complete skull lacking tympanics, left petrosal, and post-cranial elements. its left half and the right petrosal, both mandibles lacking condyles and adjacent portion of the vertical ramus

LOCALITY AND HORIZON. — Calvert Formation, Virginia, behind coronoid process, and post-cranial elements. USA, middle Miocene. LOCALITY AND HORIZON. — Calvert County, Calvert

REFERRED SPECIMENS. — (1) USNM 13472, (2) USNM Formation, Maryland, USA, middle Miocene. 23049. REFERRED SPECIMENS. — (1) USNM 16871, (2) USNM

REFERENCE. — Kellogg 1968. 23494, (3) USNM 16567.

Genus Cetotherium Brandt, 1843

TYPE SPECIES. — Cetotherium rathkei Brandt, 1843 .

REFERENCE. — Kellogg 1968.

Genus Eomysticetus Sanders & Barnes, 2002

Cetotherium rathkei Brandt, 1843 TYPE SPECIES. — Eomysticetus whitmorei Sanders &

Barnes, 2002. HOLOTYPE. — Paleontological Institute of the Academy of Science , Moscow (uncatalogued), sub-complete skull Eomysticetus whitmorei Sanders & Barnes, 2002 lacking rostrum, left tympanic. LOCALITY AND HORIZON. — South of Russia, late Mi- HOLOTYPE. — ChM PV 4253, sub-complete skull, ocene. both petrosals, both tympanics, both dentaries, postcranial elements. REFERENCES. — Brandt 1843, 1873 ; Pilleri 1986.

LOCALITY AND HORIZON. — Chandler Bridge Formation, South Carolina, USA, late Oligocene. Genus Cophocetus Packard & Kellogg, 1934 REFERENCE. — Sanders & Barnes 2002.

TYPE SPECIES. — Cophocetus oregonensis Packard & Kellogg, 1934 . Genus Herpetocetus

Van Beneden, 1872

Cophocetus oregonensis

Packard & Kellogg, 1934 TYPE SPECIES. — Herpetocetus scaldiensis Van Beneden, 1872 .

TYPE SPECIMEN. — CMUO, No. 305, sub-complete skull with tympanic and petrosal, both mandibles lacking the Herpetocetus sendaicus anterior end, and post-cranial elements. (Hatai, Hayasaka & Masuda, 1963)

Piscobalaena nana ( Cetacea , Cetotheriidae ) from the early Pliocene of Peru

HOLOTYPE. — NSMT-PV 19540, sub-complete skull skull, imperfectly preserved tympanics and petrosals, lacking anterior third of rostrum, with left tympanic both mandibles, and post-cranial elements. and both petrosals still in situ in the skull, and prepared sub-complete right tympanic, and sub-complete post- LOCALITY AND HORIZON. — Modelo Formation, Los cranial skeleton. Angeles County, California, USA, late Miocene. LOCALITY AND HORIZON. — Yushima Formation, Ja- REFERENCE. — Kellogg 1934. pan, Pliocene. REFERRED SPECIMENS. — (1) NSMT-PV 19540, cast of Genus Nannocetus Kellogg, 1929 the type specimen, (2) USNM 299652, right petrosal, Yorktown Formation, North Carolina, USA, Pliocene, TYPE SPECIES. — Nannocetus eremus Kellogg, 1929 . from Geisler & Luo 1996, (3) ChM PV 5028, right petrosal, Yorktown Formation, North Carolina, USA, Pliocene, from Geisler & Luo 1996. Nannocetus eremus Kellogg, 1929

REFERENCES. — Geisler & Luo 1996; Oishi et al. HOLOTYPE. — UCMP 26502, incomplete cranium, right 1985. tympanic and petrosal in situ in the skull, left tympanic and petrosal prepared.

REMARKS LOCALITY AND HORIZON. — Towsley Formation, Cali- Since Herpetocetus sendaicus (NSMT-PV 19540) fornia, USA, late Miocene.

is not the type species of the genus Herpetocetus , REFERENCE. — Kellogg 1929.

the specimen NSMT-PV 19540 was used, and referred petrosal were studied. The choice of using these petrosals was supported by the fact that they Genus Parietobalaena Kellogg, 1924 noticeably resemble those that are still in situ in the skull of NSMT-PV 19540. TYPE SPECIES. — Parietobalaena palmeri Kellogg, 1924 .

Genus Metopocetus Cope, 1896

Parietobalaena palmeri Kellogg, 1924

TYPE SPECIES. — Metopocetus durinasus Cope, 1896 . HOLOTYPE. — USNM 10668, sub-complete cranium lacking rostrum, with both petrosals and tympanic.

Metopocetus durinasus Cope, 1896 LOCALITY AND HORIZON. — Calvert Formation, Maryland, USA, middle Miocene.

HOLOTYPE. — USNM 8518, an incomplete cranium with right petrosal in situ in the skull and left petrosal REFERRED SPECIMENS. — (1) USNM 7424, (2) USNM prepared. 10677, (3) USNM 10909, (4) USNM 11535, (5) USNM 12697, (6) USNM 13874, (7) USNM 13903, (8) USNM

LOCALITY AND HORIZON. — Calvert Formation, Mary- 15576, (9) USNM 16119, (10) USNM 16568, (11)

land, USA, middle Miocene. USNM 16570, (12) USNM 16667, (13) USNM 16838, (14) USNM 20376, (15) USNM 23015, (16) USNM

REFERENCES. — Cope 1896; Kellogg 1968. 23022, (17) USNM 23055, (18) USNM 23203, (19) USNM 23448, (20) USNM 23725.

Genus Mixocetus Kellogg, 1934

TYPE SPECIES. — Mixocetus elysius Kellogg, 1934 .

Mixocetus elysius Kellogg, 1934

REFERENCES. — Kellogg 1924, 1968.