Hebynthus tamones, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 706-708

Hebynthus tamones sp.nov.

(Figs 1955-1961; Hab. E 291)

DIAGNOSIS. Female (length about 1.8-2.5mm): antenna (Fig. 1959) with scape pale orange ventrally; flagellum dark brown; fore and mid coxae brown, hind coxa pale orange; fore and mid femora with some brown, legs otherwise mostly yellow to pale orange; fore wing (Fig. 1960) virtually hyaline, but very weakly infused pale orange-brown in at least distal half; head about 3.5X as wide as frontovertex; malar sulcus present; antenna (Fig. 1959) with scape about 6.3X as long as broad; pedicel slightly longer than F1; funicle all segments longer than broad, F6 subquadrate; funicle with linear sensilla on all segments; sensory part of clava extending slightly more than 0.5X along ventral surface; fore wing (Fig. 1960) about 2.4X as long as broad; ventral surface of costal cell with two complete lines of setae; mid tibial spur very slightly shorter than basitarsus; ovipositor slightly exserted with exserted part about 0.2X as long as mid tibial spur; gonostylus slightly longer than mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 1.77mm; excluding ovipositor, 1.70mm (CPD).

Head mainly black; frontovertex generally with a dull, dark green sheen mixed purple, area between occipital margin and anterior ocellus with a stronger purple sheen; area between eye and scrobe, scrobal area, temple and gena with a dark coppery purple sheen mixed with some dark blue and dark green areas; frontovertex with inconspicuous dark brown setae, those on gena and interantennal prominence more conspicuous; antenna (Fig. 1959) with radicle pale orange-brown; scape pale orange; pedicel and flagellum dark brown; maxillary palpus dark brown; pronotum black with a mixed coppery and brassy sheen with a few scattered dark blue reflections; propleuron and prosternum dark brown; mesoscutum with an overall, dark blue sheen but mixed with some purple, posterior margin dark brown with a mixed brassy, coppery and purple sheen; tegula dark brown with a slight brassy and purple sheen; axilla dark purple-brown with a slight brassy sheen; scutellum dark metallic blue with some dark coppery purple reflections, vertical apex and side very shiny metallic green with some coppery reflections; dorsum of thorax clothed with dark brown setae; metanotum dark purple-brown with a weak brassy sheen; mesopleuron dark brown, anteriorly with a fairly strong purple and blue sheen with some violet, coppery and brassy reflections, posterior mainly with a coppery and brassy sheen; mesosternum dark brown; fore coxa dark brown; fore femur dark brown proximally, apical half pale orange, tibia and tarsus pale orange; mid coxa dark brown; mid femur brown proximally, apical half pale orange, tibia pale orange with a diffuse, brown, subbasal band, spur and tarsus pale orange; hind leg, including coxa, pale orange; fore wing (Fig. 1960) virtually hyaline, but very weakly infuscate in distal half, hind wing hyaline, venation orange; propodeum dark purple-brown with a weak brassy sheen, laterally dark metallic blue-green, green, coppery and brassy, side metallic green, about 20 long, conspicuous pale setae outside spiracle extending nearly to posterior margin; gaster dark brown, with a fairly strong purple and coppery sheen dorsally weakly mixed brassy, Gt1 dark, metallic blue, green and coppery laterally, side of gaster with distinct metallic green, brassy, purple and coppery reflections, venter slightly duller with mixed green, purple and blue, outer plates of ovipositor relatively dull coppery, purple and blue; gonostylus dark brown, extreme apex pale orange.

Head (Fig. 1957) sublenticular, about 3.5X as wide as frontovertex, in profile, about 2.2X as high as deep, anteriorly fairly evenly curved from occipital margin to mouth margin, interantennal prominence hardly protuberant occipital margin sharp, carinate; ocelli forming an angle of about 80°; frontovertex shiny, with fairly regular, moderately deep, polygonally reticulate sculpture of mesh size generally slightly smaller than eye facet; frontovertex with piliferous punctures moderately deep and distinct, naked in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; temple and gena more shiny than frontovertex, with conspicuous, coarse, longitudinally elongate, strigose to striate-reticulate sculpture; eye slightly overreaching occipital margin, with fairly conspicuous pale setae that are a little longer than diameter of facet, eye separated from scrobal margin by about 1.2X diameter of ocellus, area between eye and scrobe with relatively, deep and coarse, transversely elongate, polygonally reticulate sculpture; scrobes very shallow, narrowly ∩-shaped, meeting, with distinct, polygonally reticulate sculpture, weakly margined dorsally and laterally; interantennal prominence weak, dorsally rounded, sculpture on interantennal prominence much shallower and finer than that on frontovertex; antenna as in Fig. 1959; scape subcylindrical, about 6.25X as long as broad; all funicle segments clearly longer than broad, segments broader distad; clava hardly broadened, about 1.2X as wide as F6, slightly longer than F4-F6 combined, sutures strongly oblique, sensory area enlarged, extending ventrally nearly 0.6X length of clava, apex strongly obliquely truncate; malar sulcus present, but very inconspicuous; clypeal margin weakly emarginate medially;; mandible with two acute teeth and a truncate upper tooth. Relative measurements: HW 99, HH 98.5, FV 28, POL 15, OOL 0.5, OCL 6, AOL 11.5, EL 69, EW 48, MS 30, SL 50, SW 8.

Thorax (Fig. 1961) in dorsal view with pronotum short, mostly hidden by head; pronotum dorsally with imbricate-reticulate to polygonally reticulate sculpture that is slightly shallower than sculpture on frontovertex; mesoscutum generally with similar sculpture to that on pronotum; axilla with polygonally reticulate sculpture that is about as deep as that on frontovertex; scutellum clearly convex, dorsal part with regular, slightly longitudinally elongate, polygonally reticulate sculpture that is conspicuously deeper and coarser than on axilla, vertical apex and side completely smooth and shiny; visible part of mesoscutum about 1.9X as broad as long; scutellum about as long as broad; fore wing with venation and distribution of setae as in Figs 1958, 1960, ventral surface of costal cell with two complete line of setae; mid tibial spur very slightly shorter than basitarsus; propodeum (Fig. 1961) medially about 0.21X as long as scutellum and with several longitudinal carinae and irregular reticulate sculpture medially. Relative measurements: FWL 148, FWW 61; HWL 108, HWW 30.

Gaster with hypopygium reaching about 0.6X to apex; syntergum about 0.7X as long as mid tibia, with apex acute; ovipositor hardly exserted, the exserted part about 0.2X as long as mid tibial spur or less than 0.1X mid tibia; gonostylus slightly longer than mid tibial spur.

Paratype. Funicle with linear sensilla present on all segments; hypopygium Fig, 1956; ovipositor Fig. 1955. Relative measurements: OL 117, GL 43 [MT 120].

Variation. Females vary in overall length from 1.77-2.48mm, larger specimens from Manuel Antonio tend to have the fore wing generally infused very pale orange-brown, otherwise very little variation in material examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Puntarenas, Est. Biol. Monteverde, 10°20’N 84°49’W, 1540m, 18-24.ii.2004 (C. Hansson) . Paratypes: COSTA RICA,1E, Puntarenas, Monteverde , 15-16.vii.1986 (L. Masner); 1E, same data as holotype; 2E, Puntarenas, Manuel Antonio, 23-28.viii.1986 (L. Masner) . Holotype in MZUCR, paratypes in NHMUK and CNC .

COMMENTS. See comments under Hebynthus maros (p. 688).