Hebynthus cleon, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 690-692

Hebynthus cleon sp.nov.

(Figs 1893-1899; Hab. E 280)

DIAGNOSIS. Female (overall length about 1.3mm): antenna (Fig, 1894) with scape dark brown; flagellum dark brown; coxae brown, legs mostly dark brown; fore wing (Fig. 1896) virtually hyaline, but with a very small infuscate area adjacent to marginal vein (Fig. 1899); head (Fig. 1898) about 3.2X as wide as frontovertex; malar sulcus absent; antenna (Fig. 1894) with scape about 4.4X as long as broad; pedicel nearly as long as F1-F3 combined; funicle with F2-F3 transverse, remaining segments quadrate or longer than broad; funicle with linear sensilla on F3- F6 or absent from F3; sensory part of clava extending about 0.4X along ventral surface; fore wing (Fig. 1896) about 2.3X as long as broad; ventral surface of costal cell with only one complete line of setae; mid tibial spur very slightly longer than basitarsus; ovipositor slightly exserted with exserted part about 0.4X as long as mid tibial spur; gonostylus slightly longer than mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 1.26mm; excluding ovipositor, 1.19mm ( CPD) .

Head mainly black; ocellar area with a dull, dark green sheen mixed coppery purple, remainder of frontovertex, scrobal area, temple and gena with a coppery purple sheen; frontovertex, interantennal prominence and gena with inconspicuous dark brown setae; antenna (Fig. 1894) with radicle dark brown; scape dark brown; pedicel dark brown, apex orange-brown; flagellum dark brown; maxillary palpus white; thorax generally dark brown to black; pronotum with a coppery sheen; mesoscutum with an overall metallic blue sheen mixed with some coppery and brassy reflections, posterior margin with a coppery purple sheen; tegula dark brown with a slight coppery purple sheen; axilla with a coppery purple sheen; scutellum dorsally virtually matt but very weakly coppery purple, laterally narrowly metallic green, extreme sides and apex shining blue-green mixed brassy and coppery; dorsum of thorax clothed with dark brown setae; metanotum dark coppery purple; mesopleuron with a weak coppery purple lustre; mesosternum dark brown; all coxae dark brown; fore femur dark brown, tibia pale orange with a broad, dark brown, subbasal band, tarsus brown; mid femur dark brown, tibia proximally dark brown, distal 0.6X pale orange, spur pale orange, tarsus dusky pale orange, apical tarsomere brown; hind femur dark brown, tibia dark brown, extreme apex pale orange, tarsus orange brown, apical tarsomere brown; fore wing (Fig. 1896) virtually completely hyaline, but with a very small, extremely weakly infuscate area adjacent to, and not wider than, marginal vein (Fig. 1899); hind wing hyaline, venation orange to pale brown; propodeum dark purple-brown with a weak brassy sheen, side shining metallic green mixed coppery and brassy, about 10 long, conspicuous pale setae outside spiracle extending nearly to posterior margin; gaster dark brown, with a fairly strong purple and coppery sheen dorsally weakly mixed brassy, Gt1 dark, metallic green mixed coppery, more coppery laterally, side of gaster with distinct metallic green, brassy, purple and coppery reflections, venter slightly duller with mixed green, purple and blue, outer plates of ovipositor relatively dull coppery, purple and blue; gonostylus dark brown, extreme apex pale orange.

Head (Fig. 1898) about 3.2X as wide as frontovertex, in profile nearly 2X as high as deep, anteriorly fairly evenly curved from occipital margin to top of scrobes and then nearly straight to mouth margin, interantennal prominence hardly protuberant; occipital margin sharp, carinate; ocelli forming an angle of about 95°; frontovertex with fairly regular, moderately deep, polygonally reticulate sculpture of mesh size generally subequal to eye facet; frontovertex with moderately deep, distinct, piliferous punctures, 4 or 5 setae in front of anterior ocellus, a line of sparse setae along inner eye margin continued along lower eye margin; temple and gena more shiny than frontovertex, with conspicuous, coarse, longitudinally elongate, strigose to striate-reticulate sculpture; eye slightly overreaching occipital margin, with very inconspicuous pale setae that are clearly shorter than diameter of facet, eye separated from scrobal margin by slightly less than diameter of ocellus, area between eye and scrobe with relatively, deep and coarse, transversely elongate, polygonally reticulate sculpture; scrobes very shallow, narrowly ∩-shaped, meeting, with shallow, polygonally reticulate sculpture, weakly margined dorsally and laterally; interantennal prominence weak, dorsally rounded, sculpture on interantennal prominence much shallower and finer than that on frontovertex; antenna as in Fig. 1894; scape slightly broadened and flattened about 4.4X as long as broad; funicle with F1-F3 subequal, transverse, F4 larger, quadrate, F5- F6 slightly longer than broad, linear sensilla present only on F3-F6 or absent from F3; clava hardly broadened, about 1.2X as wide as F6, as long as F2-F6 combined, sutures strongly oblique, sensory area enlarged, extending ventrally about 0.4X length of clava, apex strongly obliquely truncate; malar sulcus absent; clypeal margin weakly emarginate medially; mandible with three teeth, upper tooth truncate. Relative measurements: HW 69.5, HH 65, FV 22, POL 12, OOL 2, OCL 3, AOL 9, EL 42, EW 34, MS 28, SL 35, SW 8.

Thorax (Fig. 1897) in dorsal view with pronotum short, mostly hidden by head; pronotum dorsally with imbricate-reticulate to polygonally reticulate sculpture that is conspicuously shallower than sculpture on frontovertex; mesoscutum generally with similar sculpture to that on pronotum; axilla with fine, polygonally reticulate sculpture that is about as deep as that on mesoscutum; scutellum hardly convex, dorsal part with regular, slightly longitudinally elongate, polygonally reticulate sculpture that is conspicuously deeper and coarser than on axilla, vertical apex and side completely smooth and shiny; visible part of mesoscutum about 2X as broad as long; scutellum very slightly broader than long; fore wing with venation and distribution of setae as in Figs 1896, 1899; ventral surface of costal cell with only one complete line of setae; mid tibial spur very slightly longer than basitarsus; propodeum (Fig. 1897) medially about 0.1X as long as scutellum and virtually smooth medially. Relative measurements: FWL 105, FWW 46; HWL 80.5, HWW 23.5.

Gaster with hypopygium (Fig. 1895) reaching about 0.6X to apex; syntergum about 0.7X as long as mid tibia, with apex rounded; ovipositor (Fig. 1893) slightly exserted, the exserted part about 0.4X as long as mid tibial spur or about 0.14X mid tibia; gonostylus nearly 1.2X as long as mid tibia spur. Relative measurements: OL 85.5, GL 28.5 [MT 72].

Variation. Only holotype examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Puntarenas, Potrero Grande , Cerro Kasir, LS 349700 566850, 2959m, #59114, vii-viii.2000 (M. Alfaro) . Holotype in NHMUK .

COMMENTS. Hebynthus cleon is probably closest to solex and phyrra (see comments under solex , p. 694)