Orixia demos, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 146-147

Orixia demos sp.nov.

( Figs 350-355; Hab. E 56)

DIAGNOSIS. Female (length about 1.1-1.2mm): head orange; F6 and clava dark brown; hind tibia orange-brown; fore wing ( Fig. 365) weakly, but distinctly infuscate; hind wing hyaline; ocelli forming an angle of about 55°; antenna with linear sensilla on F2-F6 sometimes absent from F2; fore wing about 2.5X as long as broad, postmarginal vein ( Fig. 351) about 0.6X as long as stigmal vein; sculpture on mesoscutum and scutellum ( Fig. 355) composed of longitudinally elongate imbricate-reticulate to polygonally reticulate cells that are much more strongly elongate laterally; mid tibial spur about 0.8X as long as basitarsus. Male: unknown.

Female (holotype): length, including ovipositor, 1.23mm; excluding ovipositor, 1.21mm (CPD).

Head generally orange with a weak brassy sheen, occiput with a brown mark above foramen; antenna with radicle, scape and pedicel orange; F1-F3 brown, F4-F5 orange (slightly dusky on right antenna), F6 brown, clava dark brown; mandible orange; palpi pale orange; frontovertex and gena with relatively scattered, conspicuous brown setae, interantennal prominence with relatively dense brown setae; thorax generally orange with a weak brassy sheen, scutellum subapically orange-brown, mesopleuron pale orange, setae on dorsum of thorax relatively dense, dark brown and conspicuous; legs pale orange, hind tibia mostly orange-brown; fore wing ( Fig. 350) mostly weakly infuscate, virtually hyaline proximad of linea calva and with a pair of opposite, poorly defined, paler, subtriangular areas distad of apex of wing venation, submarginal and stigmal veins pale orange, rest of venation brown; hind wing hyaline; propodeum orange, side slightly dusky; gaster pale orange, Gt3-Gt6 slightly dusky, apices of syntergum and outer plates of ovipositor dusky; gonostylus pale orange.

Head ( Fig. 352) about 3.9X as wide as frontovertex, in profile about 1.4X as high as deep; occipital margin not sharp; ocelli forming an angle of about 55°; about 10 punctures in two rows on frontovertex below anterior ocellus and with relatively shallow, polygonally reticulate sculpture of slightly smaller mesh than diameter of eye facet; antenna as in Fig. 353; scape about 2.4X as long as broad; F1-F2 subquadrate, F3-F6 clearly transverse, segments slightly broader distad; clava slightly broadened, slightly longer than F4-F6 combined, 2-segmented, sutures not oblique, sensory area slightly enlarged, at apex only giving the clava a distinctly transversely truncate appearance; inner eye margins subparallel; malar sulcus present, distinct. Relative measurements: HW 60, HH 48, FV 15.5, POL 4.5, OOL 1, OCL 5.5, AOL 5.5, EL 35, EW 33, MS 20, SL 26, SW 11.

Thorax ( Fig. 355) with polygonally reticulate sculpture on pronotum and axilla that is of similar depth to that on frontovertex but mostly composed of distinctly longitudinally elongate, polygonal cells that are more strongly elongate laterally; mesoscutum and scutellum with rounded to longitudinally elongate, imbricate-reticulate to polygonally reticulate sculpture that is clearly deeper than that on frontovertex; scutellum with three pairs of very small subapical pits or pores that are each of much smaller diameter than cell of reticulate sculpture; visible part of mesoscutum about 1.6X as broad as long; scutellum about as long as broad; setae on dorsum moderately dense and of more or less uniform length, separated by less than their own lengths, apical setae on scutellum slightly longer; fore wing with venation and setation as in Figs 350, 351. Relative measurements: FWL 67, FWW 26.5; HWL 44, HWW 9.5.

Gaster with hypopygium reaching about 0.8X towards apex; syntergum about 0.7X as long as mid tibia, with apex truncate; ovipositor very slightly exserted, the exserted part less than 0.2X as long as mid tibial spur.

Paratype. Funicle with linear sensilla present only on F2-F6 or absent from F1; ovipositor Fig. 354;. Relative measurements: OL 39.5, GL 11.5 [MT 53].

Variation. Females vary in overall length from 1.12-1.22mm, F4 may be brown, otherwise little variation in material available.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Guanacaste, Pitilla ( ACG), 700m, MT/YPT, 24.i-27.ii.1996 (J.S. Noyes) . Paratypes: COSTA RICA, 1E, Guanacaste, same data as holotype; 1E, Guanacaste, Est. Pitilla ( ACG), 10°59’N 85°25’W, 700m, FIT, iii.2001 (C. Moraga, P.N. Thomas) . Holotype in MZUCR, paratypes in NHMUK .