Coccidencyrtus fescus, Noyes, 2023

Noyes, John Stuart, 2023, ENCYRTIDAE OF COSTA RICA (HYMENOPTERA: CHALCIDOIDEA), 4 Subfamily Encyrtinae: tribes Arrhenophagini, Habrolepidini, Cerapterocerini, Cheiloneurini, Trechnitini, Cercobelini, Polaszekiini, Protyndarichoidini, Gahaniellini and Syrphophagini (part), mainly primary parasitoids and hyperparasitoids of Coccoidea and Psylloidea (Hemiptera), Taxonomic Monographs on Neotropical Hymenoptera (Oxford, England) 2 (11), pp. 1-921 : 113-115

Coccidencyrtus fescus sp.nov.

( Figs 254-259)

DIAGNOSIS. Female (overall length about 1.1mm): scape mostly dark brown; pedicel brown; funicle yellow; clava pale brown; all coxae white to pale yellow; fore femur white, distally pale brown; fore tibia pale orange with a diffuse, pale brown subbasal band; mid femur white with a subapical brown band; mid tibia white to pale orange with a narrow subbasal brown band; hind femur white with a broad, brown subapical band; hind tibia mostly dark brown; gaster dorsally dark brown, medially and ventrally dusky pale yellow, hypopygium brown; frontovertex about 0.4X head width; antenna ( Fig. 254) with scape about 3.5X as long as broad, funicle with F1 narrower than pedicel, F1-F4 relatively small, subequal, F5-F6 largest and longer than broad; linear sensilla present only on F4-F6; clava very nearly as long as funicle, about 3.4X as long as broad; palp formula 3-2; scutellum ( Fig. 259) with 3 pairs of setae and longitudinally elongate polygonally reticulate to striate-reticulate sculpture, deeper than that on mesoscutum; mid basitarsus with 6 or 7 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with stigmal vein slightly longer than postmarginal vein but slightly shorter than marginal vein ( Fig. 255); hypopygium reaching about 0.8X towards apex of gaster; ovipositor about 3.5X as long as gonostylus and slightly exserted with exserted part slightly shorter than mid tibial spur. Male: unknown.

Female (holotype): length, including ovipositor, 1.14mm; excluding ovipositor, 1.09mm ( CPD) .

Head dark brown to black with a weak to moderate metallic lustre; occiput with a weak coppery sheen; frontovertex generally metallic dark green mixed coppery, area between posterior ocelli and occipital margin coppery; area above scrobes coppery; scrobal area, temple and gena with a weak coppery to coppery purple sheen; frontovertex with a few, inconspicuous, scattered, brown setae; gena and interantennal prominence with a few scattered, very inconspicuous, brown setae; maxillary palpus white; radicle dark brown; scape mostly dark brown, apical 0.25X pale yellow; pedicel brown; funicle yellow, clava pale brown with proximal segment more or less yellow-brown; thorax dark brown to black with a metallic lustre; pronotum with a weak coppery sheen; mesoscutum slightly brighter metallic dark blue-green mixed coppery especially near margins; tegula dark purple-brown with a weak sheen; axilla with a weak coppery purple sheen; scutellum dorsally virtually matt, but with a very weak coppery purple sheen, extreme side and apex weakly shining and slightly brassy; mesopleuron with a moderate purple sheen; fore coxa white; fore femur white in proximal 0.5X, distally orange-brown to pale brown, tibia basically pale orange with a broad, diffuse, subbasal pale brown ring in proximal 0.6X, tarsus pale orange, apical tarsomere pale orange-brown; mid coxa very pale yellow; mid femur white in proximal 0.5X, a broad, dark brown, subapical band in distal 0.5X, extreme apex white, tibia white to pale orange with a narrow, subbasal dark, brown band, spur and tarsus very pale orange; hind coxa very pale yellow, slightly dusky proximally; hind femur white in proximal 0.5X, a broad, dark brown, subapical band in distal 0.5X, extreme apex white, tibia with extreme base white, dark brown to about 0.8X, apex pale orange, tarsus slightly dusky pale orange, apical tarsomere pale brown; wings mainly hyaline, but fore wing weakly infuscate immediately below marginal vein ( Fig. 255); propodeum dark brown with a weak purple sheen; gaster dorsally purple-brown with coppery purple and green reflections, Gt1-Gt2 medially dusky, pale yellow, side and venter mostly dark brown, but basal sternites dusky, pale yellow; gonostylus dark brown.

Head ( Fig. 257) about 2.4X as wide as frontovertex, in profile about 1.4X as high as deep, tangent to frontovertex forming an angle of about 105° with that of face; occipital margin more or less sharp, not carinate; no distinct oval depression adjacent to eye near occipital margin; ocelli forming an angle of about 90°; frontovertex not widening in front of anterior ocellus with distinct polygonally reticulate sculpture of mesh size generally a little smaller than eye facet, not shallower above top of scrobes; temple and gena with slightly shallower, longitudinally elongate, fine polygonally reticulate sculpture; top of scrobes with distinct, regular, polygonally reticulate sculpture, scrobes smooth; interantennal prominence and mouth margin with slightly shallower, polygonally reticulate sculpture; scrobes shallow, indistinctly margined, broadly ∩-shaped; interantennal prominence dorsally rounded; antenna as in Fig. 257; scape very slightly broadened and flattened, about 3.4X as long as broad; funicle with F1-F4, subequal, relatively small, subquadrate, F5-F6 largest and clearly longer than broad, linear sensilla present only on F4-F6; clava about 3.4X as long as broad, about 1.6X as wide as F6, not curved downwards, very nearly as long as funicle, sutures parallel, sensory area relatively small, at apex only giving it a weak, obliquely truncate appearance; eye slightly overreaching occipital margin, clothed with dense, conspicuous setae, each slightly longer than diameter of facet; malar sulcus present, but shallow and indistinct; clypeal margin concave, but more or less straight medially; mandible with two acute teeth and obtuse upper tooth; palp formula 3-2. Relative measurements: HW 60.5, HH 44, FV 25, POL 12, OOL 3, OCL 2.5, AOL 8, EL 33, EW 28, MS 16.5, SL 24, SW 7.

Thorax ( Fig. 259) with pronotum hardly exposed behind head, its posterior margin broadly concave, not angular medially; pronotum with slightly coarser and deeper polygonally reticulate sculpture than that in ocellar area; sculpture on mesoscutum similar but less coarse and clearly shallower; axilla with similar sculpture to mesoscutum; scutellum dorsally with very fine, polygonally reticulate sculpture that is a much deeper than sculpture on mesoscutum, slightly longitudinally elongate laterally, extreme side and apex completely smooth and shiny; visible part of mesoscutum about 1.6X as broad as long; scutellum about 1.1X as broad as long, apex with a pair of slender, elongate setae; hind femur about 4.2X as long as broad; mid basitarsus with 6 or 7 ventral pegs; mid tibial spur about as long as basitarsus; fore wing with venation and setation as in Figs 255, 256; propodeum ( Fig. 259) medially about 0.11X as long as scutellum. Relative measurements: FWL 81, FWW 37; HWL 52.5, HWW 13.

Gaster with hypopygium ( Fig. 258) reaching about 0.8X to apex; last tergite about 0.7X as long as mid tibia, with apex broadly rounded; ovipositor exserted, the exserted part about 0.6X as long as mid tibial spur or about 0.2X mid tibia. Relative measurements: OL 57, GL 16 [MT 53].

Variation. Only holotype examined.

Male. Unknown.

DISTRIBUTION. Costa Rica.

HOSTS. Unknown.

MATERIAL EXAMINED.

Type material. Holotype E: COSTA RICA, Cartago, PN Tapanti, 1200-1500m, 9°45’N 83°47’W, 20.iii- 10.iv.2000 (C. Hansson, D. Rubí ) . Holotype in NHMUK .

COMMENTS. See comments under jacintha (p. 109).