Amynthas tokioensis (Beddard, 1892)

Blakemore, Robert J., 2013, Jeju-do earthworms (Oligochaeta: Megadrilacea) - Quelpart Island revisited, Journal of Species Research 2 (1), pp. 15-54 : 34-35

Amynthas tokioensis (Beddard, 1892)

Material examined. [IV0000250398 a sub-adult specimen from Hyonyungsa Temple (12 th June, 2012, RJB) that provided DNA sample ( WO 49 detailed in Appendix) complying with either (or both?) A. tokioensis and Metaphire hilgendorfi using megaBLAST searches].

Remarks. Amynthas tokioensis (Beddard, 1892) [syns.?s chizopora,? irregularis, levis,? parvicystis;? verticosa; ? yongshilensis Hong & James, 2001,? eastoni Hong & James, 2001;? boletiformis Hong & James, 2001 (these last two synonyms as per Blakemore, 2003: 43, addenda); ? gucheonensis Song & Paik, 1970 ; jiriensis Song & Paik, 1971; ? paiki Hong in Hong, Lee & Kim, 2001: 266 (these last three synonyms as per Blakemore, 2006; 2008)]. Plus? Pheretima gucheonensis Song & Paik, 1971 ,? Pheretima surcata Ishizuka, 1999 and A. sonjaesiki Hong & James, 2009 were placed in synonymy of A. tokioensis by Blakemore (2010a: 13, fig. 2) based on reinspection of its London types. Blakemore & Ueshima (2011) & Blakemore (2012d, f) described syntypes of Amynthas levis (Goto & Hatai, 1899) as being similar to a parthenogenetic form of A. tokioensis and thus a possible synonym, as was already noted by Blakemore (2012b).

Amynthas conferticurtus Hong & James, 2009: 1241 appears to be misdescribed in several key characters: e.g., the spermathecal pores are said to be on 7 and 8 (thus allegedly qualifying for an A. pomellus species group; however, they are shown in their fig. 7 to be in 6/7/8! Moreover the supposed genital markings on 7 & 8 and 18 are not shown in their figure 7! Their description with this new name appears to be indistinguishable from the subsequent A. sonjaesiki Hong & James, 2009: 1243 that has markings resembling those claimed for A. conferticurtus and is moreover similar to A. paiki Hong, 2001 and/ or to A. fasciiformis Hong & James, 2001, thus both latter names are probably in an A. tokioensis species-group, if not also synonyms. It seems that the A. conferticurtus name also belongs.

A. tokioensis is included due to record from Jeju of the likely synonym A. yongshilensis Hong & James, 2001; cf. Metaphire hilgendorfi noted below.

Note. Regarding types, Amynthas conferticurtus Hong & James, 2009 with types IV0000120468 (H) & 479 (P) not traceable in NIBR , its synonym, A. sonjaesiki Hong & James, 2009 with types IV0000120469 (H) & 480 (P) also not traceable in NIBR .