Xenophysa naumanni Varga 1989

Varga, Zoltán, 2011, Revision of the genus Xenophysa Boursin, 1969 (Lepidoptera, Noctuidae), Zootaxa 3094, pp. 1-29 : 11-27

Xenophysa naumanni Varga 1989

Xenophysa naumanni Varga 1989 , Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 41 (1–2): 9.

Type material. Holotype: male “NO-Afghanistan, Prov.Badakhshan, Pamir-e-Kalon, Kotal-e-Shaur, 4300 m, 18.VIII.1972 (UV-Lichtfang), leg. Brade & Naumann” (coll. Naumann, Bonn, now LMK).

Paratype: 3, with the same data as the holotype (coll. Varga).

Other material: 1 3, N. Pakistan, Shandur pass, 3600 m, 36o 07’ N, 72o 38’ E, leg. M. Hreblay and G. Csorba (coll. Varga); 3, Pakistan, Hindukush Mts., Teru, cca 2500 m, 14. VII. 1994, leg. B. Herczig, Gy.M. László and G. Ronkay (colls. Herczig, G. Ronkay and Varga), 9 3, 6 Ƥ, Pakistan, Hindukush Mts., 5 km E of the Shandur pass, 3750 m, 72o 38’ E, 36o 07’ N, 26–27. 06. 2000, leg. G. Ronkay & Z. Varga (colls. Gyulai, Ronkay and Varga). Genital slides 3: Varga 4501 (Holotype), 4502 (Paratype), 6789, 7416, 7766, Ƥ: Varga 7420.

Redescription. X. naumanni was originally described based on 2 worn males, without females. Externally very similar to X. agnostica but generally with darker brownish and more concolorous colouration. Forewing 15–17 mm, expanse 32–35 mm, sexes similar. Head dark cocoa-brown, frons and tips of palpi with light greyish hairs. Antennae extremely long, reaching about the 5/6 length of forewing, finely bipectinate and ciliate (male) or filiform, finely ciliate (female). Collar and tegulae blackish brown with light grey margin, abdomen lighter brown grey. Forewing elongate triangular, acute apically, with nearly concolorous cocoa brown colouration and with short blackish basal stripe. Ground colour irrorated with light gray scales at the costa and hind margin. Orbicular and reniform spots oblique quadrangular, light grey with brownish filling, interconnected, with blackish brown intermaculation; claviform spot short. Transverse lines faint, subterminal line marked with light grey scales and bordered by faint and confluent arrowheads at the inner margin. Cilia fuscous with dark brown medial line. Hindwing smoky greyish brown, lighter basally, lunule faint (Plate 5, Figs 27–28). Male genitalia: Basic configuration like X. agnostica , but generally more compressed, stubby. Uncus strong, thick, slightly shorter and with obtuse hook terminally. with very strong, quadrangular cross-shaped extension of uncus at base. Lateral lobes of tegumen densely spinulose, larger, and more elongate than in X. agnostica . Juxta broad pentagonal. Valve shorter with shorter digitus than in X. agnostica , dorsal extension higher and more densely dentate. Aedeagus ventrally with strongly sclerotised knot, vesica without subbasal bladder and relatively shorter recurved diverticulum (Plate 10, Figs. 48a, b; 49a, b). Female genitalia: Papillae anales triangular, more acute and slightly more densely covered by long hairs than in X. agnostica . Antrum with deeper incision and with longer and stronger sclerotised bilateral arms; ductus bursae short, bursa elliptic, without signa; appendix recurved and larger than in X. agnostica (Plate 14, Figs 64).

PLATE 1. Xenophysa specimens. Fig. 1. Xenophysa junctimacula Christoph , female Lectotype, Persia. Fig. 2. Xenophysa junctimacula Christoph , male, Iran, Shahkuh, Lectotype. Fig. 3. Xenophysa junctimacula Christoph , male, Iran, Alborz Mts. Holotype of X. huberi persica . Fig. 4. Xenophysa junctimacula Christoph , female, Iran, Alborz Mts. Fig. 5. Xenophysa junctimacula huberi Varga , male, Turkey, Hakkari prov., Paratype. Fig. 6. Xenophysa junctimacula huberi Varga , male, Turkey, Hakkari prov., Paratype.

PLATE 2. Xenophysa specimens. Fig. 7. Xenophysa cacumena Brandt , female, Iran, Shahkuh, Paralectotype of X. junctimacula Chr. Fig. 8. Xenophysa cacumena Brandt , male, Iran, Khorassan, Barm-i-Firus. Fig. 9. Xenophysa cacumena Brandt , male, Turkmenistan, Kopet-Dagh. Fig. 10. Xenophysa cacumena Brandt , female, Turkmenistan, Kopet-Dagh. Fig. 11. Xenophysa cacumena afghanorea Varga , male, Paratype, Afghanistan, Dasht-i- Nawar.Fig. 12. Xenophysa cacumena afghanorea Varga , male, Holotype, Afghanistan, Dasht-i- Nawar.

PLATE 3. Xenophysa specimens. Fig. 13. Xenophysa argyroramma Varga , male, Holotype, Afghanistan, Dasht-i-Nawar. Fig. 14. Xenophysa argyroramma Varga , male, Paratype, Afghanistan, Dasht-i-Nawar. Fig. 15. Xenophysa xenogramma Boursin , male, Holotype, Afghanistan, Band-i-Amir. Fig. 16. Xenophysa xenogramma Boursin , female, Paratype, Afghanistan, Band-i- Amir. Fig. 17. Xenophysa monastica Boursin , male, Holotype, Afghanistan, Dasht-i-Nawar. Fig. 18. Xenophysa monastica Boursin , male, Paratype, Afghanistan, Band-i-Amir.

PLATE 4. Xenophysa specimens. Fig. 19. Xenophysa sharhu Varga , male, Mongolia, Chovd aimak, Mongol Altai Mts. Fig. 20. Xenophysa sharhu Varga , female, Mongolia, Govi Altai aimak, Govi Altai Mts. Fig. 21. Xenophysa poecilogramma Varga , male, Pakistan, Hindukush Mts. Fig. 22. Xenophysa poecilogramma Varga , female, Paratype, Afghanistan, Salang pass. Fig. 23. Xenophysa pseudopoecila Varga , male, Holotype, Kirghisia, Karagai-tau. Fig. 24. Xenophysa pseudopoecila Varg a, fe mal e, Paratype, Pakistan, Hindukush Mts.

PLATE 5. Xenophysa specimens. Fig. 25. Xenophysa agnostica Varga , male, Paratype, Afghanistan, Darwaz Mts. Fig. 26. Xenophysa agnostica Varga , female, Paratype, Afghanistan, Darwaz Mts. Fig. 27. Xenophysa naumanni Varga , male, Pakistan, Hindukush Mts. Fig. 28. Xenophysa naumanni Varga , male, Pakistan, Hindukush Mts. Fig. 29. Xenophysa boursini Varga , male, Afghanistan, Salang-pass. Fig. 30. Xenophysa (Paraxenophysa) paradoxa Varga , male, Paratype, Darwaz Mts.

PLATE 6. Xenophysa male genitalia. Fig. 31a. Xenophysa (Paraxenophysa) paradoxa Varga , Paratype, Afghanistan, Darwaz Mts.genital capsula. Fig. 31b. Xenophysa (Paraxenophysa) paradoxa Varga , Paratype, Afghanistan, Darwaz Mts., aedeagus Fig. 32a. Xenophysa junctimacula Chr. „ Persia ”, genital capsula. Fig. 32b. Xenophysa junctimacula Chr. , „ Persia ”, aedeagus. Fig. 33a. Xenophysa junctimacula huberi Varga , Paratype, Turkey, Hakkari, genital capsula. Fig. 33b. Xenophysa junctimacula huberi Varga , Paratype, Turkey, Hakkari, aedeagus. Fig. 34a. Xenophysa junctimacula huberi Varga , Turkey, Paratype, Hakkari, genital capsula. Fig. 34b. Xenophysa junctimacula huberi Varga , Paratype, Hakkari, aedeagus.

PLATE 7. Xenophysa male genitalia. Fig. 35a. Xenophysa argyrogramma Varga , Paratype, Afghanistan, Dasht-i-Nawar, genital capsula. Fig. 35b. Xenophysa argyrogramma Varga , Paratype, Afghanistan, Dasht-i-Nawar, aedeagus. Fig. 36a. Xenophysa xenogramma Boursin , Paratype, Afghanistan, Band-i-Amir, genital capsula. Fig. 36b. Xenophysa xenogramma Boursin , Paratype, Afghanistan, Band-i-Amir, aedeagus. Fig. 37a. Xenophysa monastica Boursin , Paratype, Afghanistan, Band-i-Amir, genital capsula. Fig. 37b. Xenophysa monastica Boursin , Paratype, Afghanistan, Band-i-Amir, aedeagus. Fig. 38a. Xenophysa cacumena Brandt , Paratype, Iran, Khorassan, genital capsula. Fig. 38b. Xenophysa cacumena Brandt , Paratype, Iran, Khorassan, aedeagus.

PLATE 8. Xenophysa male genitalia. Fig. 39a. Xenophysa sharhu Varga , Mongolia, Mongol Altai, genital capsula. Fig. 39b. Xenophysa sharhu Varga , Mongolia, Mongol Altai, aedeagus. Fig. 40a. Xenophysa sharhu Varga , Mongolia, Govi Altai, genital capsula. Fig. 40b. Xenophysa sharhu Varga , Mongolia, Govi Altai, aedeagus. Fig. 41a. Xenophysa poecilogramma Va rg a, P ak - istan, Hindukush Mts., genital capsula. Fig. 41b. Xenophysa poecilogramma Varga, Hindukush Mts. , aedeagus. Fig. 42a. Xenophysa poecilogramma Varga , Pakistan, Hindukush Mts., genital capsula. Fig. 42b. Xenophysa poecilogramma Varga , Pakistan, Hindukush Mts., aedeagus.

PLATE 9. Xenophysa male genitalia. Fig. 43. Xenophysa pseudopoecila Varga , Holotype, Kirghisia, Karagai-tau, genital capsula. Fig. 44. Xenophysa pseudopoecila Varga , Paratype, Pakistan, Hindukush Mts., genital capsula. Fig. 45a. Xenophysa pseudopoecila Varga , Paratype, Pakistan, Hindukush Mts., genital capsula. Fig. 45b. Xenophysa pseudopoecila Varga, P arat y pe, Pakistan, Hindukush Mts. aedeagus. Fig. 46a. Xenophysa pseudopoecila Varga , Pakistan, Hindukush Mts., genital capsula. Fig. 46b. Xenophysa pseudopoecila Varga , Pakistan, Hindukush Mts., aedeagus.

PLATE 10. Xenophysa male genitalia. Fig. 47a. Xenophysa agnostica Varga , Paratype, Pakistan, Darwaz Mts., genital capsula. Fig. 47b. Xenophysa agnostica Varga , Paratype, Pakistan, Darwaz Mts., aedeagus. Fig. 48a. Xenophysa agnostica Varg a, P ar atype, Pakistan, Darwaz Mts., genital capsula. Fig. 48b. Xenophysa agnostica Varga , Paratype, Pakistan, Darwaz Mts., aedeagus. Fig. 49a. Xenophysa naumanni Varga , Paratype, Afghanistan, Pamirs, Wakhan, genital capsula. Fig. 49b. Xenophysa naumanni Varga , Pakistan, Hindukush Mts., aedeagus. Fig. 50a. Xenophysa naumanni Varga , Pakistan, Hindukush Mts., genital capsula. Fig. 50. Xenophysa naumanni Varga , Pakistan, Hindukush Mts., aedeagus.

PLATE 11. Xenophysa female genitalia. Fig. 51. Xenophysa junctimacula Christoph, LectoHolotype , Iran, Shahkuh. Fig. 52. Xenophysa junctimacula Christoph , Khorassan, Mts. Kopet-Dagh. Fig. 53. Xenophysa argyrogramma Varga , Afghanistan, Dasht-i-Nawar. Fig. 54. Xenophysa xenogramma Boursin , Afghanistan, Band-i-Amir.

PLATE 12. Xenophysa female genitalia. Fig. 55. Xenophysa cacumena Brandt , Iran, Elburz Mts. Fig. 56. Xenophysa cacumena Brandt , Turkmenistan, Kopet-dagh. Fig. 57. Xenophysa monastica Boursin , Afghanistan, Band-i-Amir. Fig. 58. Xenophysa sharhu Varga , Mongolia, Mongol Altai.

PLATE 13. Xenophysa female genitalia. Fig. 59. Xenophysa poecilogramma Varga , Pakistan, Hindukush Mts. Fig. 60. Xenophysa poecilogramma Varga , Holotype, Afghanistan, Salang pass. Fig. 61. Xenophysa pseudopoecila Varga , Paratype, Pakistan, Hindukush Mts. Fig. 62. Xenophysa pseudopoecila Varga , Paratype, Pakistan, Hindukush Mts.

PLATE 14. Xenophysa female genitalia. Fig. 63. Xenophysa agnostica Varga , Paratype, Afghanistan, Badakhshan. Fig. 64. Xenophysa agnostica Varga, Kirghisia , Karagai-tau. Fig. 65. Xenophysa naumanni Varga , Pakistan, Hindukush Mts. Fig. 66. Xenophysa agnostica Varga , „Samarkand”, Holotype.

FIGURE 67. Geographical distribution of the western species group of Xenophysa : ٭ Xenophysa (Paraxenophysa) paradoxa , Ο X. junctimacula , Ο h X. junctimacula huberi , ● X. cacumena , ª X. cacumena afghanorea , □ X. argyrogramma , ̻ X. xenogramma , X. monastica , ˔ X. boursini .

Biological notes. This species was previously known only by two males, collected in high altitudes (4300 m) by the second Pamir expedition of C. M. Naumann, and it was thought to be extremely localised and rare, since several large museum (HNHM, ZMRA, ZSM, colls. of Klapperich, Naumann, Omoto, Tsvetaev, etc., more than 200 specimens of Xenophysa !) from the Hissaro-Darwaz, Badakhshan and Pamirs regions did not contain specimens of this species. The more recent expeditions of Hungarian collectors were able, however, to bring some new materials from the high parts of the Hindukush Mts. (N Pakistan) consisting of both sexes in good condition. The species has a wider vertical distribution (from about 2500 m to the subnival altitudes) but is not common and with a relatively short flying period (from mid-June to very early July). The rather worn type specimens were collected atypically late and at extremely high altitudes. Thus, the range of distribution of this species is as yet incompletely known.