Togoperla Klapálek, 1907

Stark, Bill P. & Sivec, Ignac, 2008, The Genus Togoperla Klapálek (Plecoptera: Perlidae), Illiesia 4 (20), pp. 208-225 : 209

Genus Togoperla Klapálek, 1907

Type species.‐ Perla limbata Pictet, 1841 .

Distribution.‐ Eastern Palearctic and Oriental regions. Known from China, Japan, Thailand and Vietnam.

Diagnosis. Adult Togoperla are large, typically dark, and fully winged. Males of all known species have a well developed, truncate, emarginate or slightly notched lobe on the 5 th abdominal tergum and large mesal membranous areas on terga 6‐9 which often bear setal clusters but seldom have sensilla basiconica patches. Male hemiterga have a basal callus and a relatively short (in comparison to Agnetina ) anterior process. The aedeagal tube of all species has a basodorsal lobe and most also have a pair of lateral lobes on the tube; the membranous apical sac is coarsely armed with small triangular spines. Females have a large parabolic subgenital plate, frequently with a median sclerite in the dorsal membrane of the plate. The subgenital plate in several species is emarginate or shallowly notched but no examples of narrow deep notches are known for the group. Egg collars are absent for most species, but two species have a sessile collar without a flanged rim and another has a small thickened polar disc. Larvae are associated for only one species, T. limbata (Pictet) ( Sivec et al. 1988) , which lacks anal gills, has few (usually 1‐2) slender intercalary setae on abdominal terga, and lacks a median dorsal fringe of long silky setae. Provisional keys for adult Togoperla are presented below the species accounts.

Current species. 1) canilimbata ( Enderlein, 1909) ; 2) clavata sp. n.; 3) fortunati Navas, 1926 ; 4) limbata ( Pictet, 1841) ; 5) noncoloris Du & Chou, 1999 ; 6) perpicta Klapálek, 1921 ; 7) poilanina ( Navas, 1934) ; 8) shan Stark & Sivec, 1991 ; 9) totanigra Du & Chou, 1999 ; 10) triangulata Du & Chou, 1999 ; 11) tricolor Klapálek, 1921 .