Bia rebeli aegina Penz & Simonsen

Penz, Carla M., Casagrande, Mirna M., Devries, Phil & Simonsen, Thomas J., 2017, Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae), Zootaxa 4258 (3), pp. 201-237 : 218

Bia rebeli aegina Penz & Simonsen , NEW SSP.

( Figs 5 c–f, 10g, 12)

Diagnostic description. Defined by the following combination of characters: (1) MF DFW white apical ocelli medium-small (most of the range) showing a cline to very small or absent (the DFW and VFW apical ocelli, and the VFW ocellus below M1 are very small or absent in Bolivian specimens). (2) M DFW orange band moderately wide, usually opaque; orange scales somewhat extended proximally along veins. (3) M DFW iridescent band from anal margin to approximately half of the CuA2 cell in most specimens, nearly reaching CuA 2 in others, and less frequently reaching CuA2 (southern Peru; variable within localities). F DFW iridescent band more diffuse and narrower than that of actorion , especially noticeable below CuA2 where the iridescence is less extended towards the tornus. (4) M DFW androconial organ on the CuA-CuA1-CuA2 intersection pale, contrasting scale color of surrounding area. (5) M DHW discal androconial pad varying from pale brown to silvery brown. (6) DHW discal hairpencil brown, darker in color than discal androconial pad. (7) F VFW ripple pattern less dense than that of M.

Etymology. In Greek mythology, Aegina, daughter of the river god Asopus, married Actor, son of king Deioneus of Phocis and his wife Diomede. Here we name one of the Bia rebeli subspecies after the nymph Aegina as means of acknowledging that this species is separate from Bia actorion .

Type material. Holotype M ( Fig. 5 c), deposited in the PJD collection, two labels separated by // and transcribed verbatim: Peru: Madre de Dios, 250m Los Amigos Research Center 12o34’10.0”S, 70o06’01.4’W 13 May 2004 FW 25.7 P.J. DeVries Trap: 4 und // HOLOTYPE Bia rebeli aegina Penz & Simonsen, 2017 . Paratypes are listed in Appendix, and Fig. 5 d shows a paratype F.

Distribution and examined specimens. Fig. 12 and Appendix.

Remarks. Specimens from Bolivia slightly differ from Peruvian by their minute (sometimes absent) dorsal forewing white apical ocelli, and a very small dark ocellus in ventral forewing cell M2 (compare Figs 5 e–f with 5c– d). Although examined specimens from Peru, Manu National Park generally conform to the rebeli aegina phenotype, one specimen has paler than expected hind wing discal androconial organs ( Fig. 5 g) although not as pale as those of rebeli rebeli or rebeli acreana .

Figures

FIGURE 5. Adults in dorsal and ventral views. Taxon names appear below each set. Bia rebeli rebeli: a, male, Peru (USNM); b, female, Peru, Huallaga, Tingo Maria (ZSM). Bia rebeli aegina Penz & Simonsen, new ssp.: c, male holotype, Peru, Madre de Dios, Los Amigos Research Center [Puerto Maldonado] (PJD); d, female paratype, same locality (PJD); e, male, Bolivia, Santa Cruz, Buenavista (CMNH); f, female, same locality (CMNH). Intermediate between B. rebeli rebeli and B. rebeli aegina: g, male, Peru, Madre de Dios, Manu, Pakitza (USNM).

FIGURE 12. Map showing collection localities of Bia species and subspecies (see Appendix). Some close localities are represented by one single dot. Dubious label data, localities that could not be accurately identified or were incomplete (e. g., Brazil, Pará) were not placed on map.