Nalassus Mulsant, 1854

Nabozhenko, Maxim & Ando, Kiyoshi, 2018, Subtribal, Generic And Subgeneric Composition Of Darkling Beetles Of The Tribe Helopini (Coleoptera: Tenebrionidae) In The Eastern Palaearctic Region, Acta Zoologica Academiae Scientiarum Hungaricae 64 (4), pp. 277-327 : 296-298

Subgenus Nalassus Mulsant, 1854

Nalassus (Nalassus) brunneus brunneus ( Marseul, 1876), comb. n. ( Fig. 8 )

Marseul, 1876: 140 ( Helops ); Chûjô, 1975: 22 ( Tarpela ); Masumoto, 1993: 129, figs 1, 7 ( Tarpela ); Nabozhenko & Löbl, 2008: 256 ( Tarpela ); Jung, 2012: 44, fig. 22 in plates II, V, X, XIII ( Tarpela ).

= Lamperos japonica Allard, 1876: 46 (Type locality: “ Japon ”) .

Material examined. 1 m ( CKA): Japan, Kagoshima Pref., Yakushima, Malaise trap, 250 m, 1–30.xii.2007 (leg. Takeo Yamauchi et al.); 1 m, 2 f ( CKA): Japan, Ohkawa Rindô, Yakushima, Satsuma, 3.v.2008 (leg. Kiyoshi Ando); 3 exs. ( CKA), Dazaifu, Fukuoka, 3.vi.1974 (leg. Y. Kitsuki); 1 m, 1 f ( CKA), Suwa Temple, Nagasaki, 14.vi.1986 (leg. K. Ando); 1 ex. ( CKA), Oshima, Kii, 29.iv.1959 (leg. Y. Kimura); 1 m ( CKA), Ashyû, Kyoto, 27.vi.1976 (leg. K. Ando); 3 exs. ( CKA), Vill. of Four Seasons, Kyokushi, Kikuchi City, Kumamoto (Higo), 21.iii.2011 (leg. K. Ando); 2 exs. ( CKA), ditto, 18.iii.2012 (leg. K. Ando); 2 exs. ( CKA), Yukawa, Nachikatsuura, Kii, 21.iii.1994 (leg. M. Yoshida); 1 f ( CKA), Mt. Tara-dake, Nagasaki, 3.v.2006 (leg. K. Ando); 3 exs. ( CKA), Yosakoe-Pass, Kikawamura Vill., Tosa, 25.ix.1999 (leg. S. Nagashima); 2 f ( CKA), Anbô, Yakushima, Satsuma, 6.v.2008 (leg. K. Ando); 2 exs. ( CKA), Ohkawa Rindô, Yakushima, Satsuma, 3.v.2008 (leg. K. Ando); 1 ex. ( CKA), Mt. Ukidake, Nanatsu Vill., Karatsu, Saga, 26.v.2007 (leg. K. Ando); 7 exs. ( CKA), Japan, Kagoshima Pref., Hanyama, Yakushima Is., Malaise trap (H1 & H2), 250 m alt., 1–30.xii.2007, 25–28.xii.2006, and 2–29.iii.2008 (leg. T. Yamauchi et al.); 4 exs. ( CKA), Japan, Kagoshima Pref., Yakushima Is., Mt. Aiko-dake, Malaise trap ( AN 3), 170 m alt., 2–30.iii.2008 (leg. T. Yamauchi et al.); 1 ex. ( CKA), Japan, Kagoshima Pref., Yakushima Is., Mt. Aiko-dake, Malaise trap ( AC 2), 150 m alt., 26–29.i.2007 (leg. T. Yamauchi et al.); 1 ex. ( CKA), Japan, Kagoshima Pref., Yakushima Is., Kankakei, Malaise trap (K1), 220 m alt., 27. II.–2.iii.2007 (leg. T. Yamauchi et al.).

Distribution. Japan: Honshu (Southwestern districts), Shikoku, Kyushu, Izu-Ôshima Is., Nii-jima Is., Hachijô-jima Is., Hirado-jima Is., Tsushima Is., Yakushima Is.; Korea? The species was listed for Korea only by HEYDEN (1887) and probably by misidentification.

protrochanters

Nalassus (Nalassus) brunneus konoi ( Nakane, 1963), comb. n.

( Fig. 9 )

Nakane, 1963: 28 ( Tarpela ); Chûjô, 1977: 4; Chûjô, 1978a: 75 ( Tarpela ); Chûjô, 1978b: 15 ( Tarpela ); Masumoto, 1993: 129 ( Tarpela ); Nabozhenko & Löbl, 2008: 256 ( Tarpela ).

Type material examined. Holotype ( MHU), m with labels: “ Holotype ”, “Naka-no- Shima, Iss. Tokara, S. Japan, IV.3–13.1953. Coll. Hiroshi Kono ”, “ Tarpela brunnea konoi Nak. Det. T. Nakane. 1963”, “115–21b”, “Nakane Coll. SEHU Japan 1999”.

Material examined. 1 m ( CKA): Japan, Sokobaru, Ishigaki Is., Yaeyama, 20.i.2012 (leg. T. Nakata) .

Comments. This subspecies was interpreted as the junior synonym of the nominotypical subspecies by Jung (2012) without any arguments. We consider N. brunneus konoi as a good subspecies (see differences in the key to the species).

Distribution. Japan: Tokara Nakanoshima Is., Amami-Ôshima Is., Okinawa-jima Is., Ishigaki-jima Is., and Iriomote-jima Is.

Figures

Fig. 8. Nalassus (s. str.) brunneus brunneus, details of structure: A = head and prothorax, ventral view, B = head, ventral view (arrow shows weakly developed posterior grooves beginning at lower aspect of eyes), C = prosternum, D = prosternal process, procoxae and

Fig. 9. Nalassus brunneus konoi, holotype, female: A = habitus, B = labels, C = protibia, D = mesotibia