Broscini, Hope, 1838

Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, Zootaxa 4598 (1), pp. 1-148 : 13

Key to New Zealand Broscini subtribes and genera

(adapted from Liebherr et al. 2011).

1. Mentum median process bifid ( Fig. 8 ), glossal sclerite with two setae; basal orifice of median lobe completely closed dorsally; internal sac of aedeagus with sclerites X and Y; female reproductive tract with helminthoid sclerite ( Fig. 5 ) (subtribe Nothobroscina, Roig-Juñent 2000 )............................................................................. 2

- Mentum median process simple, glossal sclerite with four apical setae; basal orifice of median lobe completely or partially open dorsally; internal sac of aedeagus without sclerites X and Y; female reproductive tract without helminthoid sclerite.......................................................... (subtribe Creobiina, Jeannel 1941) Bountya Townsend 1971

2. Each side of head with one or more supraorbital punctures ( Fig. 9 ), each bearing a single seta; external apex of protibia not distally expanded; elytra without a row of setiferous punctures along 7 th stria/interval................................. 3

- Each side of head with a single, plurisetose supraorbital puncture ( Fig. 9 ); external apex of protibia greatly distally expanded ( Fig. 10A ) or distally expanded ( Fig. 10B ); elytra with a row of setiferous punctures along 7 th stria/interval ( Fig. 1 )........................................................................................ Mecodema Blanchard 1853

3. Head with one supraorbital seta; mentum with two deep paramedial foveae; base of maxillary stipe bisetose.............. 4

- Head with two supraorbital setae; mentum broadly and shallowly depressed paramedially, lacking deep paramedial foveae; base of maxillary stipes trisetose; protibia slender............................................ Oregus Putzeys 1868

4. Pronotal base smooth (except Stewart Island species); only two basal male protarsomeres with adhesive articulosetae on ventral surface.......................................................................... Diglymma Sharp 1886

- Pronotal base distinctly punctate, as well as in pronotal foveae; all male protarsomeres glabrous medioventrally..................................................................................... Orthoglymma Liebherr et al. 2011

Figures

FIGURE 1. Dorsal view of a Mecodema specimen showing taxonomic structures used in the species descriptions. H =head; P = pronotum; E = elytra; S = striae; I1 = interval 1; A = apex; 7S = 7 th strial setae; HSP = humerus setose puncture; HA = humeral angle; BM = basal margin; P = protibia.

FIGURE 5. The entire female reproductive tract (ventral view) including taxonomic structures (for all other abbreviated structure names see Seldon et al. 2012). However, for this revision only the following structures are important: HS = helminthoid sclerite; G1 = gonocoxite 1; G2 = gonocoxite 2; TDC = transverse dorsal carina; R = ramus.

FIGURE 8. Detail of the ventral head with the taxonomic structures used in the species descriptions. SBMS = submentum setae; SBM = submentum sclerite; ML = mentum lobe; MP = mentum process (with indentation); MPW = mentum process width; SBS = stipes basal setae; MPL = mentum process length; MPS = mentum process setae; SBMC = submentum sclerite constriction; GP = gula pits; GS = gula suture.

FIGURE 9. Dorsal view of head with detail of the morphological structures used in the species descriptions. Ƒ = vertex; SPS = supraorbital puncture setae; SP = supraorbital puncture; F = frons area; FCS = frontoclypeal suture; C = clypeus; M = mandible; LS = labrum setae; L = labrum; ACM = anterior clypeal microsculpture; CSP = clypeal setose puncture; TP = tentorial pit; SG = supraorbital grooves; E = eye; ƑG = vertexal groove.

FIGURE 10. The protibia form is a character used mainly to determine between genera, except for one species in Mecodema where the difference between (A) greatly distally expanded (M. antarctica) and (B) expanded distally (all other Mecodema species) is used.