Mecodema tibiale Castelnau, 1867

Seldon, David S. & Buckley, Thomas R., 2019, The genus Mecodema Blanchard 1853 (Coleoptera: Carabidae: Broscini) from the North Island, New Zealand, Zootaxa 4598 (1), pp. 1-148 : 123-126

publication ID

https://doi.org/ 10.11646/zootaxa.4598.1.1

publication LSID

lsid:zoobank.org:pub:6BA5F086-1014-46B5-B6D5-77FE0CD07AEC

DOI

https://doi.org/10.5281/zenodo.5678448

persistent identifier

https://treatment.plazi.org/id/03BBD026-027F-B773-90FB-B4CBE9D9FD21

treatment provided by

Plazi

scientific name

Mecodema tibiale Castelnau, 1867
status

comb. nov.

Mecodema tibiale Castelnau, 1867 View in CoL , comb. n.

Figure 60 View FIGURE 60 .

Maoria tibialis Castelnau, 1867 View in CoL (Puerua (Molyneux) River, SL).

Metaglymma tibiale Castelnau, 1868 View in CoL , redescribed.

Metaglymma rufipes Broun, 1886 View in CoL (nr. Outram, DN), synonymised by Britton 1949.

Metaglymma asperum Broun, 1893a View in CoL (Taiera Beach, DN), synonymised by Britton 1949.

Metaglymma junctum Broun, 1893a (Strath-Taiera, CO), synonymised by Britton 1949.

Metaglymma calcaratum Broun, 1903 View in CoL (Maniototo, CO), synonymised by Britton 1949.

Diagnosis: Differs from other Mecodema species by having: 1, ventrites 4 and 5 with multiple setose punctures each side of midline; 2, apical area of elytra with numerous setose punctures; 3, shape of basal lobe and setal distribution along ventral edge of left paramere ( Fig. 60 View FIGURE 60 LP).

Description: Length 14.6–18.4 mm, pronotal width 4.3–5.5 mm, elytral width 5–6.4 mm. Colour of body dorsally matte reddish-brown to glossy black, ventrally matte to glossy reddish-brown on ventrites and legs.

Head: Broad and convex ( Fig. 14B View FIGURE 14 ). Vertex smooth ( Fig. 60 View FIGURE 60 ); vertexal groove slightly impressed laterally, absent medially; large supraorbital puncture bearing 3 setae; supraorbital grooves absent; frons with a small and shallow depression each side of midline (anteriorly); frontoclypeal suture ( Fig. 9 View FIGURE 9 ) a well-defined groove, tentorial pits large and well-defined; anterior area of clypeus without microsculpture, 1 setose puncture each side bearing 1 seta ( Fig. 60 View FIGURE 60 ). Labrum rounded, anterior edge outwardly curved with 3 evenly-spaced setae each side. Mentum lobes rounded ( Fig. 12F View FIGURE 12 ), mentum process long and narrow, slightly angled upward (15°), indentation notched ( Fig. 12D View FIGURE 12 ); mentum setae absent. Submentum sclerite constriction narrow with 6–8 setae, 3–4 clustered laterally. Stipes with 2 basal setae. Gula pits small, suture well-defined, gula ( Fig. 8 View FIGURE 8 ) flat with very fine transverse lines. Gena with very fine wrinkles forming an isodiametric pattern across entire area.

Prothorax: Prothoracic carina broad the entire length, slightly crenulated with 5–8 setae each side ( Fig. 60 View FIGURE 60 ), extended to anterior angle; posterior lateral sinuation indistinctly carinate, angled inward; pronotum broad and moderately deflected, overall shape cordate; midline well-defined, posterior medial diamond-shaped impression present, disc without microsculpture ( Fig. 60 View FIGURE 60 ); pronotal foveae narrow and shallow; anterior edge straight and posterior edge inwardly curved medially. Prosternum flat and smooth; proepisternum without microsculpture. Procoxal setae absent; protibia distally expanded and shovel-like ( Fig. 10B View FIGURE 10 ).

Elytra: Broad and moderately deflected, truncated length due to steep angle to apex; humeral angle anteriorly convergent ( Fig. 6A View FIGURE 6 ); basal margin slightly curved and bevelled to base, interval 1 extended to margin; lateral carina narrow in anterior ¾, broadened posterior ⅓, extended to humeral angle; humerus with 4–5 setose punctures; suture ( Fig. 1 View FIGURE 1 ) impressed; all striae with irregularly sized asetose punctures, stria 5 with asetose punctures 2x size of stria 6 asetose punctures; intervals 1–6 slightly convex, intervals 7–9 moderately convex; interval microsculpture present as lines; 7 th strial setal pattern with 2–3 setose punctures in anterior ½, 3–4 setose punctures in posterior ½, setose punctures large; numerous setose punctures in striae on steep slope to apex ( Fig. 60 View FIGURE 60 ); row of many setae above and along length lateral carina.

Ventral surface: Mesepisternum with rugose wrinkles; metepisternum ( Fig. 2 View FIGURE 2 ) without microsculpture; setose punctures on mesocoxae absent and present on metacoxae (1). All abdominal ventrites ( Fig. 2 View FIGURE 2 ) finely lineate laterally; ventrite 3 with 2–4 setose punctures each side of midline (not laterally), ventrite 4 with 3–6 setose punctures each side of midline, may be near lateral margin, ventrite 5 with 5–9 setose punctures each side of midline to lateral margin; ventrite 6 setae present: ♂ with 2 setose punctures distantly spaced along straight apical edge, ♀ with 2 evenly spaced setose punctures each side of midline on bluntly rounded apical edge; lateral foveae absent. Anterior metaventrite process ( Fig. 2 View FIGURE 2 ) a long and narrow triangle with a narrow carina the entire length (may be indistinct).

Male genitalia: Apical portion of penis lobe asymmetrically rounded with a distinct deflection to right of vertical axis (VV) ( Fig. 15A View FIGURE 15 ), ventral edge of apex with symmetrical curve that is curved flatly upward to form slightly forward (of vertical axis) dorsal curve (LV) ( Fig. 60 View FIGURE 60 PL); shaft of penis lobe broad and shortened, shaft straight (VV); overall length of penis lobe moderately curved ventrally (LV) ( Fig. 16B View FIGURE 16 ). Structures of the endophallus ( Fig. 4 View FIGURE 4 ): lateral form of the apex of the central spicule rounded; dorsal form of the apex of the central spicule short and narrow; relative extent of setae (scales) of the apical plate sparse (1–25%); size of left setose flange very small; size of right setose flange large. Left paramere basal lobe rounded, narrowly rectangular, short anterior slope to arm; arm long and narrow ( Fig. 60 View FIGURE 60 LP); terminal lobe indistinct from arm, terminus narrowed with apical tuft of sparse medium-lengthed setae ( Fig. 60 View FIGURE 60 LP); sparse short setae along strongly curved ventral edge for ¾ of length. Right paramere long, very narrowly rectangular, narrowed to terminus ( Fig. 60 View FIGURE 60 RP) with a double row of long setae extended along apical ¾ of ventral edge.

Female genitalia: Basal gonocoxite 1 long and narrow, ventral surface with a few rugose wrinkles, internal dorso-lateral carina without setae. Gonocoxite 2 ( Fig. 5 View FIGURE 5 ) long and narrow, bluntly, rounded triangle with egdes reflexed, ventral surface with numerous sensilla pits and short seta posterior to apex on ventral surface (near base of gonocoxite 2). Ramus short and narrow.

Comments: Mecodema tibiale seems to have the narrowest of distributions in comparison to the other former Metaglymma species, Mecodema aberrans and M. monilifer . This species was described by Castelnau who designated four syntypes, all of which were collected from the type locality, here we designate a lectotype and three paralectotypes from this material.

Distribution: New Zealand, South Island only, Central Otago (CO), Dunedin (DN), Southland (SL).

Lectotype: MCNG specimen labelled. Riv. Molyneux, N. Zelanda. Coll. Castelnau [hw, black square border] / Typus [red font, red square border] / Maoria tibiale Cast. [hw] / tibiale Cast. [hw, black square border] / SYNTYPUS Maoria tibalis Castelnau, 1867 [hw, red label] / LECTOTYPE Maoria tibialis Castelnau , Mecodema tibiale design. by DS Seldon, TR Buckley 2019 [red label] / Museo Civico di Genova.

Paralectotypes: 3, label data same as lectotype.

Material examined: 1, NCant. Dunedin N. Zeland Coll. Castelnau [hw, black square border] / Museo Genova Maoria tibialis [hw] Coll. Castelnau (MCNG); 1♂, 1♀, NEW ZEALAND CO, Cromwell-Alexandra terrace 16.NOV.1977, J.C. Watt [hw] / On ground at night [hw] ( NZAC); 1♂ , Kyeburn Ck, Naseby Otago 8 Nov 68 J.I. Townsend ( NZAC); 1♀ , Kyeburn Ck, Naseby Otago 8 Nov 68 J.I. Townsend / NZAC04004586 About NZAC .

NZAC

New Zealand Arthropod Collection

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Carabidae

Genus

Mecodema

Loc

Mecodema tibiale Castelnau, 1867

Seldon, David S. & Buckley, Thomas R. 2019
2019
Loc

Metaglymma calcaratum

Broun 1903
1903
Loc

Metaglymma asperum

Broun 1893
1893
Loc

Metaglymma rufipes

Broun 1886
1886
Loc

Metaglymma tibiale

Castelnau 1868
1868
Loc

Maoria tibialis

Castelnau 1867
1867
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