Theronus, Stiller, 2009

Stiller, M., 2009, Revision of Elginus Theron (Hemiptera: Cicadellidae: Deltocephalinae) with the description of two new genera and comments on the grassland leafhopper fauna in South Africa, Zootaxa 2135 (1), pp. 1-56: 33-34

publication ID 10.11646/zootaxa.2135.1.1


persistent identifier

treatment provided by


scientific name



Theronus   gen.n.

Type species: Theronus priapus   sp.n. ( Figs 47–49 View FIGURES 47–50 , 284–290 View FIGURES 284–290 ).

Gender masculine.

Etymology: Named for the collector, J.G. Theron, who published taxonomic papers on leafhoppers between 1970–1988.

External morphology. Vertex with disc smooth, anterior margin shagreened. Clypeus light brown with ochraceous horizontal arcs. Clypellus broad, 1.3 times as long as wide, tapering apically. Pronotum without carina. Submacropter with tegmina with three anteapical cells, four narrow apical cells, reduced appendix; hind wing about half as long as tegmina, jugal lobe present, folded over anal area. Macropters rare, with hind wing as long as tegmina. Profemur with short, stout setae in row AV, about 8 long, fine intercalary setae, one AM1 seta. Protibial setal formula 1+4; mesotibial setal formula 4+4; metafemur setal formula 2+2+1, with median pair consisting of one robust, long seta, one fine, short seta.

Genital capsule. Male. Aedeagus extremely long, narrow, flattened laterally; extending into tergite VIII, sometimes into tergite VII; gonopore subapical, dorsal ( Figs 284, 285 View FIGURES 284–290 ). Connective linear, arms fused ( Fig. 284 View FIGURES 284–290 ). Plate about half length of pygofer; apex denticulate ( Fig. 286 View FIGURES 284–290 ). Anal tube superficial, not invaginated into pygofer. Pygofer lobe setose, posterior margin uniformly rounded; in dorsal view acutely triangular ( Figs 287, 290 View FIGURES 284–290 ).

Genital capsule. Female. Pygofer with numerous macrosetae, ovipositor as long as pygofer.

Relationships. This genus is a typical member of the tribe Deltocephalini   that is defined by having the connective fused to the aedeagus ( Zahniser & Dietrich, 2008; Ossiannilsson, 1983). It is characterized by the male genitalia that are so large that they extend into abdominal segment VIII or even VII, and have the aedeagus blade-like and curved ventroposteriorly. This genus is not closely related to any Afrotropical genus in this tribe such as Recilia Edwards. Occasionally   , however, similar colouration is found in pale specimens of species of genera such as Pravistylus   and Vilargus   that are in the tribe Paralimnini   . No genera from the Palaearctic Region bearing any genitalic resemblance were listed by Ossiannilsson (1983) or Ribaut (1952), although some bear similar external features, such as colour and shape. Linnavuori (1959) described a number of genera from the Nearctic Region in the Deltocephalini   , but none show any relationship with Theronus   .