Viola nevadensis Boissier (1838: 18)

Magrini, Sara, 2019, Typification of Iberian names in Viola Sect. Melanium (Violaceae) with nomenclatural notes, Phytotaxa 399 (2), pp. 147-159 : 157

Viola nevadensis Boissier (1838: 18)

Synonyms:— Viola crassiuscula Bory (1820: 16) ≡ Viola nevadensis Boiss. var. crassiuscula (Bory) Steudel (1841: 772) .

Lectotype (first-step designated by Burdet et al. 1991):—In Sierra Nevada glareosis, Jul. Aug. 1837, E. Boissier 26 [G, ex herbier Boissier (5 feuilles). Le lectotype est le spécimen disposé au bas de la page n° 1; les autres sont des isolectotypes]; (second-step designated here):— SPAIN. Granada: in Sierra Nevada glareosis, E. Boissier 26 (G, Herb. Boissier G 00343986!; Fig. 6 ); syntypes: K000327610, K000327607, K000327623, M0112759, MPU013559, MPU013560.

Notes: —This name was previously typified by Burdet et al. (1991), but the specification of the herbarium and collection were not followed by any available number that permanently and unambiguously identify the lectotype (ICN, Recommendation 9C.1, Turland et al. 2018). In fact, they wrote: “ LECTOTYPUS: 6 étiquettes imprimées: HERB. E. BOISSIER // Viola Nevadensis Boiss. El. n° 26. // in Sierra Nevada glareosis. / Jul. Aug. 1837. / Alt. 9000’–11000’. G, ex herbier Boissier (5 feuilles). Obs. 1. Le lectotype est le spécimen disposé au bas de la page n° 1; les autres sont des isolectotypes.”. Then, according to Art. 9.17 of ICN ( Turland et al. 2018), the type citation of Burdet et al. (1991) must be accepted as the first-step lectotypification, while a lectotype was designated here by a second-step lectotypification ( Fig. 6 ). Additional six syntypes were also traced in K, M, and MPU (see above).

Viola nevadensis is a Spanish endemic, with a distribution limited to the Sierra Nevada (Andalucia).

Viola tricolor L. subsp. olyssiponensis Rouy ex Magnier (1887: 114)

Synonyms:— Viola olyssiponensis Rouy (a sphalm.) ≡ Viola tricolor L. var. olyssiponensis (Rouy) Henriques (1888: 13) ≡ Viola tricolor L. subsp. olyssiponensis (Rouy) Nyman (1889: 47) ≡ Viola arvensis Murray subsp. olyssiponensis (Rouy) W. Becker (1904: 40 , 43).

Lectotype (designated here):— PORTUGAL. Lisbon: coteaux de la Vallée d’Alcantara , près Lisbonne, Mars 1886, J. Daveau s.n. (LY40689!, sub Viola olyssiponensis , Fig. 7 ); syntypes: in LY, MA, MPU, P, and S (LY0040690, LY0040692, LY0040695, MA82504, MPU023604, MPU023603, P04816640, P05114269, P05114270, P05114283).

Notes: —Rouy described in Scrinia Florae Selectae 6 (1887) a new subspecies of Viola tricolor called “ Viola Olyssiponensis Rouy (subsp. nova)”, further specifying its subspecific rank writing after the description “ Sous-espèce du type spécifique V. tricolor ”. After him, this taxon was called mistakenly Viola olyssiponensis , instead of V. tricolor subsp. olyssiponensis . An error probably due to the label of the specimens of “ Flora selecta exsiccata. Publié par Ch. Magnier ” distributed in 1887 and reporting “1380. Viola olyssiponensis Rouy Sp. Nov.”

For the typification of this name, specimens were selected among the exsiccata belonging to such gathering and reporting “ Hab.: Portugal, coteaux argilo-basaltiques de la Vallée d’Alcantara, près Lisbonne, Mars 1886 (J. Daveau) ” as cited by Rouy in the protologue. As the author failed to designate the type, a lectotype ( Fig. 7 ) is chosen among the syntypes belonging to the cited collection found in LY (Art. 9.6, Turland et al. 2018), as the herbarium Rouy is included in the Collection Herbier Roland Bonaparte in LY ( Stafleu & Cowan 1983). Other syntypes were traced in MA, MPU, P, and S.

Viola tricolor subsp. olyssiponensis is considered a heterotypic synonym of V. kitaibeliana ( Muñoz Garmendia et al. 1993) , of V. kitaibeliana subsp. trimestris ( Espeut 1999) and, more recently ( Raab-Straube & Henning 2018), of V. hymettia Boiss. & Heldr. in Boissier (1854: 57). Today, its effective position within the Section Melanium is not clear and needs to be further studied.

Figures

FIGURE 6. Second-step lectotype of Viola nevadensis (G, Herb. Boissier G00343986!; image available from the Catalogue des herbiers de Genève (CHG) - Conservatoire & Jardin botaniques de la Ville de Genève, accessed on 05-02-2019, at http://www.ville-ge.ch/musinfo/ bd/cjb/chg/adetail.php?lang=en&id=252880&base=img).

FIGURE 7. Lectotype of Viola tricolor subsp. olyssiponensis (LY40689!; sub Viola olyssiponensis, image available at https://explore. recolnat.org/picturae/LY0040689).